chr1 : 52,550,519 52,550,830
311 bp 159 TFs 2 linked genes
This 311 bp open chromatin element is linked to TUT4 and ENSG00000272371 and is bound by 159 transcription factors.
Linked Genes
2 genes
Gene Expression Dist. to TSS Distance Link type
TUT4 2.2 kb Proximal Proximity
ENSG00000272371 4.0 kb Proximal Proximity
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr1:52,545,519 – 52,555,830
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
159 transcription factors
Source
Cell type
AHDC1 1 dataset
ChIP HepG2 ENCFF069FSH 311 bp overlap
ARID2 1 dataset
ChIP BIN-67_lentivirus-SMARCA4-K785R GSE117734.ARID2.BIN-67_lentivirus-SMARCA4-K785R 219 bp overlap
ARID3A 4 datasets
ChIP Hep-G2 ENCSR000EDP.ARID3A.Hep-G2 144 bp overlap
ChIP HepG2 ENCFF122GLS 311 bp overlap
ChIP HepG2 ENCFF341DES 311 bp overlap
ChIP HepG2 ENCFF341DES 271 bp overlap
ARID4A 2 datasets
ChIP Hep-G2 ENCSR386UBO.ARID4A.Hep-G2 272 bp overlap
ChIP HepG2 ENCFF142DIE 311 bp overlap
ARID4B 1 dataset
ChIP HepG2 ENCFF519OXJ 201 bp overlap
ARID5B 1 dataset
ChIP HepG2 ENCFF964FWK 204 bp overlap
ASH2L 3 datasets
ChIP Hep-G2 ENCSR508LMH.ASH2L.Hep-G2 311 bp overlap
ChIP HepG2 ENCFF207QHL 311 bp overlap
ChIP HepG2 ENCFF207QHL 311 bp overlap
ATF1 1 dataset
ChIP HepG2 ENCFF239LTQ 311 bp overlap
BRCA1 1 dataset
ChIP HepG2 ENCFF585LUC 311 bp overlap
BRD3 1 dataset
ChIP HEK293T GSE39579.BRD3.HEK293T 146 bp overlap
BRD4 5 datasets
ChIP CLL_patient1 GSE109411.BRD4.CLL_patient1 221 bp overlap
ChIP CLL_patient3 GSE109411.BRD4.CLL_patient3 258 bp overlap
ChIP CLL_patient3_4h_CpG GSE109411.BRD4.CLL_patient3_4h_CpG 208 bp overlap
ChIP K-562_iBET-BD2-IFNG GSE138084.BRD4.K-562_iBET-BD2-IFNG 242 bp overlap
ChIP K-562_iBET-IFNG GSE138084.BRD4.K-562_iBET-IFNG 205 bp overlap
CBFA2T3 2 datasets
ChIP K-562 ENCSR697YLJ.CBFA2T3.K-562 209 bp overlap
ChIP K562 ENCFF673OEZ 311 bp overlap
CBFB 1 dataset
ChIP Hep-G2 ENCSR479RZV.CBFB.Hep-G2 267 bp overlap
CEBPA 2 datasets
ChIP Hep-G2 ERP000209.CEBPA.Hep-G2 171 bp overlap
ChIP HepG2 ENCFF175DFS 167 bp overlap
CEBPB 6 datasets
ChIP Hep-G2 ENCSR000EEX.CEBPB.Hep-G2 158 bp overlap
ChIP Hep-G2 ENCSR000BQI.CEBPB.Hep-G2 138 bp overlap
ChIP HepG2 ENCFF074JWB 155 bp overlap
ChIP HepG2 ENCFF536NTI 173 bp overlap
ChIP K-562 ENCSR000EHE.CEBPB.K-562 137 bp overlap
ChIP K562 ENCFF189VBN 209 bp overlap
CEBPD 2 datasets
ChIP Hep-G2 ENCSR000BQJ.CEBPD.Hep-G2 98 bp overlap
ChIP HepG2 ENCFF345JDB 104 bp overlap
CEBPG 1 dataset
ChIP HepG2 ENCFF503XBC 144 bp overlap
CREB1 1 dataset
ChIP HepG2 ENCFF576ERP 311 bp overlap
CREM 1 dataset
ChIP HepG2 ENCFF049UDY 222 bp overlap
CTCF 1 dataset
ChIP endodermal cell ENCFF471YCZ 274 bp overlap
DDX5 1 dataset
ChIP NTERA2 GSE58641.DDX5.NTERA2 250 bp overlap
DLX6 2 datasets
ChIP Hep-G2 ENCSR272TOJ.DLX6.Hep-G2 131 bp overlap
ChIP HepG2 ENCFF371CVH 311 bp overlap
DMAP1 2 datasets
ChIP Hep-G2 ENCSR670YPQ.DMAP1.Hep-G2 213 bp overlap
ChIP HepG2 ENCFF247MSU 311 bp overlap
DRAP1 1 dataset
ChIP Hep-G2 ENCSR765MKZ.DRAP1.Hep-G2 290 bp overlap
E2F4 2 datasets
ChIP Hep-G2 ENCSR924LSO.E2F4.Hep-G2 224 bp overlap
ChIP HepG2 ENCFF311TOD 269 bp overlap
EGR1 3 datasets
ChIP Hep-G2 ENCSR026GSW.EGR1.Hep-G2 282 bp overlap
ChIP K-562 ENCSR000BNE.EGR1.K-562 147 bp overlap
ChIP K562 ENCFF006PJY 211 bp overlap
EGR2 1 dataset
ChIP HEK293 ENCFF336LFH 235 bp overlap
EP300 2 datasets
ChIP Hep-G2 ENCSR000BLW.EP300.Hep-G2 195 bp overlap
ChIP HepG2 ENCFF354ACD 231 bp overlap
FOSL2 1 dataset
ChIP HepG2 ENCFF548CXY 236 bp overlap
FOXJ3 1 dataset
ChIP Hep-G2 ENCSR413AJG.FOXJ3.Hep-G2 197 bp overlap
FOXK1 1 dataset
ChIP Hep-G2 ENCSR603BJQ.FOXK1.Hep-G2 302 bp overlap
FOXP1 1 dataset
ChIP Hep-G2 ENCSR369YUK.FOXP1.Hep-G2 205 bp overlap
FOXP4 2 datasets
ChIP Hep-G2 ENCSR232LLP.FOXP4.Hep-G2 288 bp overlap
ChIP HepG2 ENCFF462ULY 311 bp overlap
GATA4 1 dataset
ChIP pancreatic-progenitor_PP1 GSE104840.GATA4.pancreatic-progenitor_PP1 173 bp overlap
GATA6 3 datasets
ChIP DE DE-GATA6-1 219 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 251 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 128 bp overlap
GATAD2A 2 datasets
ChIP HepG2 ENCFF252XNH 189 bp overlap
ChIP HepG2 ENCFF252XNH 249 bp overlap
GMEB1 1 dataset
ChIP Hep-G2 ENCSR780OXL.GMEB1.Hep-G2 189 bp overlap
HDAC1 2 datasets
ChIP Hep-G2 ENCSR750OWO.HDAC1.Hep-G2 186 bp overlap
ChIP HepG2 ENCFF304IEJ 311 bp overlap
HDAC2 3 datasets
ChIP Hep-G2 ENCSR000BMC.HDAC2.Hep-G2 199 bp overlap
ChIP HepG2 ENCFF087XCR 239 bp overlap
ChIP HepG2 ENCFF990GUQ 306 bp overlap
HLF 1 dataset
ChIP HepG2 ENCFF854JLR 199 bp overlap
HMG20A 1 dataset
ChIP HepG2 ENCFF599VWU 308 bp overlap
HNF4A 10 datasets
ChIP Hep-G2 ENCSR469FBY.HNF4A.Hep-G2 311 bp overlap
ChIP Hep-G2 ERP000209.HNF4A.Hep-G2 195 bp overlap
ChIP Hep-G2 ENCSR000EEU.HNF4A.Hep-G2 181 bp overlap
ChIP Hep-G2 ENCSR000BLF.HNF4A.Hep-G2 248 bp overlap
ChIP HepG2 ENCFF146SSF 175 bp overlap
ChIP HepG2 ENCFF669NAM 203 bp overlap
ChIP liver ENCFF354NRH 175 bp overlap
ChIP liver ENCFF449HPV 231 bp overlap
ChIP liver ENCFF449HPV 244 bp overlap
ChIP liver ERP002306.HNF4A.liver 130 bp overlap
HNF4G 2 datasets
ChIP Hep-G2 ENCSR000BNJ.HNF4G.Hep-G2 176 bp overlap
ChIP HepG2 ENCFF323ATZ 201 bp overlap
HOXA3 2 datasets
ChIP Hep-G2 ENCSR999JKC.HOXA3.Hep-G2 263 bp overlap
ChIP HepG2 ENCFF374TCI 156 bp overlap
ISL2 2 datasets
ChIP Hep-G2 ENCSR492IHH.ISL2.Hep-G2 299 bp overlap
ChIP HepG2 ENCFF742RIP 311 bp overlap
JUND 1 dataset
ChIP liver ENCSR837GTK.JUND.liver 99 bp overlap
KAT7 1 dataset
ChIP HepG2 ENCFF613PTN 311 bp overlap
KDM1A 5 datasets
ChIP HepG2 ENCFF240UWG 311 bp overlap
ChIP HepG2 ENCFF730KKG 236 bp overlap
ChIP K-562 ENCSR908CMW.KDM1A.K-562 228 bp overlap
ChIP K-562 ENCSR360HRA.KDM1A.K-562 206 bp overlap
ChIP NB4_DMSO GSE128528.KDM1A.NB4_DMSO 301 bp overlap
KDM3A 1 dataset
ChIP Hep-G2 ENCSR387JKT.KDM3A.Hep-G2 136 bp overlap
KLF1 1 dataset
ChIP HEK293 ENCSR859BMR.KLF1.HEK293 217 bp overlap
KLF12 1 dataset
ChIP HepG2 ENCFF395LSO 311 bp overlap
KLF6 1 dataset
ChIP Hep-G2 ENCSR757EKM.KLF6.Hep-G2 264 bp overlap
KMT2A 2 datasets
ChIP Hep-G2 ENCSR692GFR.KMT2A.Hep-G2 311 bp overlap
ChIP HepG2 ENCFF103PKS 154 bp overlap
LCOR 1 dataset
ChIP HepG2 ENCFF499KCU 207 bp overlap
LCORL 1 dataset
ChIP HepG2 ENCFF017FTI 311 bp overlap
LIN54 2 datasets
ChIP Hep-G2 ENCSR597MCV.LIN54.Hep-G2 251 bp overlap
ChIP HepG2 ENCFF662XDE 311 bp overlap
MAX 2 datasets
ChIP Hep-G2 ENCSR168DYA.MAX.Hep-G2 188 bp overlap
ChIP HepG2 ENCFF507HCX 311 bp overlap
MED1 2 datasets
ChIP Hep-G2 ENCSR959XNY.MED1.Hep-G2 224 bp overlap
ChIP HepG2 ENCFF495TSS 311 bp overlap
MEF2A 1 dataset
ChIP Hep-G2 ENCSR291MJH.MEF2A.Hep-G2 142 bp overlap
MIER3 1 dataset
ChIP HepG2 ENCFF032KTL 297 bp overlap
MLX 1 dataset
ChIP Hep-G2 ENCSR125DAD.MLX.Hep-G2 296 bp overlap
MNX1 1 dataset
ChIP Hep-G2 ENCSR108MKR.MNX1.Hep-G2 203 bp overlap
MTA1 1 dataset
ChIP Hep-G2 ENCSR983KRB.MTA1.Hep-G2 211 bp overlap
MXD4 1 dataset
ChIP Hep-G2 ENCSR441KFW.MXD4.Hep-G2 270 bp overlap
MYBL2 1 dataset
ChIP Hep-G2 ENCSR581KCO.MYBL2.Hep-G2 291 bp overlap
MYC 1 dataset
ChIP HepG2 ENCFF575FXK 311 bp overlap
MYNN 1 dataset
ChIP Hep-G2 ENCSR398TMP.MYNN.Hep-G2 226 bp overlap
NCOR1 1 dataset
ChIP HepG2 ENCFF685NAH 311 bp overlap
NFIL3 1 dataset
ChIP HepG2 ENCFF686VLI 230 bp overlap
NFYC 1 dataset
ChIP Hep-G2 ENCSR569ARC.NFYC.Hep-G2 131 bp overlap
NR2C2 2 datasets
ChIP Hep-G2 ENCSR559ZKI.NR2C2.Hep-G2 226 bp overlap
ChIP HepG2 ENCFF944PRH 311 bp overlap
NR2F6 3 datasets
ChIP Hep-G2 ENCSR518WPL.NR2F6.Hep-G2 126 bp overlap
ChIP HepG2 ENCFF429VKC 267 bp overlap
ChIP HepG2 ENCFF514UJI 252 bp overlap
NR5A1 2 datasets
ChIP Hep-G2 ENCSR310OZS.NR5A1.Hep-G2 303 bp overlap
ChIP HepG2 ENCFF970YZO 181 bp overlap
OSR2 1 dataset
ChIP HEK293 ENCSR324LTM.OSR2.HEK293 218 bp overlap
PAXIP1 2 datasets
ChIP Hep-G2 ENCSR530ARJ.PAXIP1.Hep-G2 296 bp overlap
ChIP HepG2 ENCFF526NOJ 154 bp overlap
PHF5A 1 dataset
ChIP Hep-G2 ENCSR191TLD.PHF5A.Hep-G2 123 bp overlap
PHF8 1 dataset
ChIP Hep-G2 ENCSR604VAE.PHF8.Hep-G2 264 bp overlap
PITX1 1 dataset
ChIP HepG2 ENCFF468QTQ 311 bp overlap
POLR2A 1 dataset
ChIP adrenal gland ENCFF843OBJ 208 bp overlap
PPARG 1 dataset
ChIP Hep-G2 ENCSR130VQL.PPARG.Hep-G2 281 bp overlap
PROX1 1 dataset
ChIP HepG2 ENCFF016ZJS 311 bp overlap
RAD21 3 datasets
ChIP HepG2 ENCFF360ZSW 151 bp overlap
ChIP HepG2 ENCFF906QIS 176 bp overlap
ChIP liver ENCFF522JHE 294 bp overlap
RARA 1 dataset
ChIP HepG2 ENCFF582XUA 168 bp overlap
RCOR1 1 dataset
ChIP K562 ENCFF216EEJ 248 bp overlap
RFXAP 1 dataset
ChIP Hep-G2 ENCSR524RLU.RFXAP.Hep-G2 143 bp overlap
RXRA 3 datasets
ChIP HepG2 ENCFF763IEA 164 bp overlap
ChIP liver ENCFF077DAP 311 bp overlap
ChIP liver ENCFF807CIA 281 bp overlap
RXRB 2 datasets
ChIP HepG2 ENCFF539ZAY 267 bp overlap
ChIP HepG2 ENCFF539ZAY 231 bp overlap
SALL1 1 dataset
ChIP HepG2 ENCFF426MCK 311 bp overlap
SAP130 1 dataset
ChIP HepG2 ENCFF892EHZ 222 bp overlap
SCRT1 1 dataset
ChIP HEK293 ENCSR605MGM.SCRT1.HEK293 212 bp overlap
SCRT2 1 dataset
ChIP HEK293 ENCFF711QQB 303 bp overlap
SIX1 1 dataset
ChIP Hep-G2 ENCSR561BQM.SIX1.Hep-G2 263 bp overlap
SKI 2 datasets
ChIP Hep-G2 ENCSR754MUD.SKI.Hep-G2 140 bp overlap
ChIP HepG2 ENCFF631IPX 265 bp overlap
SMAD3 1 dataset
ChIP Hep-G2 ENCSR005GZH.SMAD3.Hep-G2 311 bp overlap
SMAD4 1 dataset
ChIP HepG2 ENCFF615GTE 157 bp overlap
SMARCA4 1 dataset
ChIP endometrial-epithelial-cells_KO GSE106660.SMARCA4.endometrial-epithelial-cells_KO 281 bp overlap
SMARCB1 1 dataset
ChIP Hep-G2 GSE69566.SMARCB1.Hep-G2 311 bp overlap
SOX13 1 dataset
ChIP HepG2 ENCFF062VSQ 93 bp overlap
SOX5 1 dataset
ChIP HepG2 ENCFF470KZD 270 bp overlap
SOX6 2 datasets
ChIP Hep-G2 ENCSR766TSU.SOX6.Hep-G2 196 bp overlap
ChIP HepG2 ENCFF767OCK 311 bp overlap
SP1 5 datasets
ChIP Hep-G2 ENCSR000BJX.SP1.Hep-G2 173 bp overlap
ChIP Hep-G2 ENCSR460YAM.SP1.Hep-G2 153 bp overlap
ChIP HepG2 ENCFF123KAM 229 bp overlap
ChIP liver ENCFF597LFJ 311 bp overlap
ChIP liver ENCFF769YSM 311 bp overlap
SP5 2 datasets
ChIP Hep-G2 ENCSR019NPF.SP5.Hep-G2 308 bp overlap
ChIP HepG2 ENCFF931FHV 262 bp overlap
STAG1 1 dataset
ChIP Hep-G2 ENCSR167MTG.STAG1.Hep-G2 160 bp overlap
TAF1 1 dataset
ChIP Hep-G2 ENCSR918OKK.TAF1.Hep-G2 275 bp overlap
TARDBP 1 dataset
ChIP HepG2 ENCFF356JNC 311 bp overlap
TBX2 2 datasets
ChIP Hep-G2 ENCSR410FEH.TBX2.Hep-G2 194 bp overlap
ChIP HepG2 ENCFF811TLA 311 bp overlap
TCF12 1 dataset
ChIP WA01 ENCSR000BIT.TCF12.WA01 109 bp overlap
TFAP4 1 dataset
ChIP HepG2 ENCFF932XOY 278 bp overlap
TFDP2 2 datasets
ChIP Hep-G2 ENCSR069JKP.TFDP2.Hep-G2 219 bp overlap
ChIP HepG2 ENCFF794WDW 300 bp overlap
TFE3 1 dataset
ChIP Hep-G2 ENCSR589SNT.TFE3.Hep-G2 271 bp overlap
THAP11 1 dataset
ChIP HepG2 ENCFF272SWH 311 bp overlap
THRA 1 dataset
ChIP HepG2 ENCFF025KMX 115 bp overlap
THRB 2 datasets
ChIP Hep-G2 ENCSR430JGJ.THRB.Hep-G2 311 bp overlap
ChIP HepG2 ENCFF476INC 291 bp overlap
ZBED4 1 dataset
ChIP Hep-G2 ENCSR409PMR.ZBED4.Hep-G2 267 bp overlap
ZBTB33 2 datasets
ChIP K-562 ENCSR876GXA.ZBTB33.K-562 146 bp overlap
ChIP K562 ENCFF875HLX 282 bp overlap
ZBTB40 1 dataset
ChIP HepG2 ENCFF130IRD 311 bp overlap
ZEB2 2 datasets
ChIP HEK293 ENCSR417VWF.ZEB2.HEK293 275 bp overlap
ChIP K562 ENCFF795CMH 311 bp overlap
ZFHX2 1 dataset
ChIP HEK293 ENCFF167TUA 275 bp overlap
ZFP90 1 dataset
ChIP HepG2 ENCFF409XXV 311 bp overlap
ZFY 1 dataset
ChIP Hep-G2 ENCSR949OEV.ZFY.Hep-G2 152 bp overlap
ZGPAT 2 datasets
ChIP Hep-G2 ENCSR704IGU.ZGPAT.Hep-G2 311 bp overlap
ChIP HepG2 ENCFF055YSO 311 bp overlap
ZMYM3 1 dataset
ChIP HepG2 ENCFF408KTI 311 bp overlap
ZMYM4 1 dataset
ChIP HepG2 ENCFF567SQY 311 bp overlap
ZNF12 1 dataset
ChIP Hep-G2 ENCSR711KBM.ZNF12.Hep-G2 117 bp overlap
ZNF121 6 datasets
ChIP HEK293 ENCFF839FUF 279 bp overlap
ChIP HEK293 ENCSR224QDY.ZNF121.HEK293 255 bp overlap
ChIP HEK293 GSE76494.ZNF121.HEK293 129 bp overlap
ChIP Hep-G2 ENCSR945QEW.ZNF121.Hep-G2 246 bp overlap
ChIP HepG2 ENCFF343YSL 269 bp overlap
ChIP WTC11 ENCFF291API 218 bp overlap
ZNF143 2 datasets
ChIP Hep-G2 ENCSR080UEM.ZNF143.Hep-G2 300 bp overlap
ChIP HepG2 ENCFF658YIR 311 bp overlap
ZNF217 2 datasets
ChIP Hep-G2 ENCSR055FQB.ZNF217.Hep-G2 290 bp overlap
ChIP HepG2 ENCFF455XGO 311 bp overlap
ZNF219 1 dataset
ChIP HepG2 ENCFF266JIR 280 bp overlap
ZNF292 1 dataset
ChIP HepG2 ENCFF975MAJ 311 bp overlap
ZNF335 2 datasets
ChIP HEK293 ENCFF784SLD 286 bp overlap
ChIP HEK293 ENCSR328SUD.ZNF335.HEK293 311 bp overlap
ZNF362 1 dataset
ChIP HepG2 ENCFF256AZN 105 bp overlap
ZNF384 2 datasets
ChIP Hep-G2 ENCSR101FJU.ZNF384.Hep-G2 162 bp overlap
ChIP HepG2 ENCFF129PLC 103 bp overlap
ZNF44 1 dataset
ChIP Hep-G2 ENCSR914HPP.ZNF44.Hep-G2 253 bp overlap
ZNF501 2 datasets
ChIP Hep-G2 ENCSR227PHM.ZNF501.Hep-G2 286 bp overlap
ChIP HepG2 ENCFF879XZR 311 bp overlap
ZNF503 2 datasets
ChIP Hep-G2 ENCSR998YJI.ZNF503.Hep-G2 282 bp overlap
ChIP HepG2 ENCFF923HZL 311 bp overlap
ZNF574 1 dataset
ChIP HepG2 ENCFF206MMY 311 bp overlap
ZNF614 1 dataset
ChIP HepG2 ENCFF677IUD 311 bp overlap
ZNF629 2 datasets
ChIP Hep-G2 ENCSR363ASY.ZNF629.Hep-G2 244 bp overlap
ChIP HepG2 ENCFF490FFQ 283 bp overlap
ZNF792 1 dataset
ChIP HepG2 ENCFF825WPU 311 bp overlap