chr9 : 33,381,203 33,381,714
511 bp 134 TFs 6 linked genes
This 511 bp open chromatin element is linked to 6 target genes and is bound by 134 transcription factors.
Linked Genes
6 genes
Gene Expression Dist. to TSS Distance Link type
NFX1 91.0 kb Distal Multiome
NOL6 92.4 kb Distal Multiome
CHMP5 116.5 kb Distal Multiome
BAG1 116.8 kb Distal Multiome
ANKRD18B 142.7 kb Distal Multiome
B4GALT1 214.2 kb Distal Multiome
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr9:33,376,203 – 33,386,714
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
134 transcription factors
Source
Cell type
AR 2 datasets
ChIP MDA-MB-453 ERP003503.AR.MDA-MB-453 145 bp overlap
ChIP myofibroblast GSE90772.AR.myofibroblast 267 bp overlap
ARID4A 1 dataset
ChIP HepG2 ENCFF142DIE 145 bp overlap
ARNTL 1 dataset
ChIP Hep-G2 ENCSR794LVK.ARNTL.Hep-G2 164 bp overlap
ARRB1 1 dataset
ChIP prostate GSE55615.ARRB1.prostate 135 bp overlap
ASH2L 3 datasets
ChIP H1 ENCFF399KAM 430 bp overlap
ChIP Hep-G2 ENCSR508LMH.ASH2L.Hep-G2 321 bp overlap
ChIP WA01 ENCSR850KIP.ASH2L.WA01 511 bp overlap
ATF2 2 datasets
ChIP H1 ENCFF295GZO 463 bp overlap
ChIP WA01 ENCSR000BQU.ATF2.WA01 303 bp overlap
ATF3 1 dataset
ChIP HCT-116_CAMP GSE74355.ATF3.HCT-116_CAMP 136 bp overlap
BCOR 3 datasets
ChIP WA01_KDM2B_KO GSE104690.BCOR.WA01_KDM2B_KO 196 bp overlap
ChIP WA01_RNF2-R GSE104690.BCOR.WA01_RNF2-R 239 bp overlap
ChIP WA01_RNF2-R_Doxycyclin GSE104690.BCOR.WA01_RNF2-R_Doxycyclin 337 bp overlap
BRD1 1 dataset
ChIP HUES-64 GSE104059.BRD1.HUES-64 455 bp overlap
BRD2 1 dataset
ChIP MV4-11_IBET151_500nM GSE120715.BRD2.MV4-11_IBET151_500nM 151 bp overlap
BRD4 1 dataset
ChIP SW480_16h_TNF-a GSE110473.BRD4.SW480_16h_TNF-a 132 bp overlap
CHD2 3 datasets
ChIP H1 ENCFF991MKH 331 bp overlap
ChIP HeLa-S3 ENCSR000ECP.CHD2.HeLa-S3 112 bp overlap
ChIP WA01 ENCSR000EBT.CHD2.WA01 151 bp overlap
CHD7 2 datasets
ChIP hiPSC_AF22_abD3F5 GSE108506.CHD7.hiPSC_AF22_abD3F5 187 bp overlap
ChIP hiPSC_derived_lt-NES GSE108506.CHD7.hiPSC_derived_lt-NES 332 bp overlap
CREB1 5 datasets
ChIP GM23338 ENCFF432ZEW 258 bp overlap
ChIP GM23338 ENCSR214ZAV.CREB1.GM23338 282 bp overlap
ChIP H1 ENCFF955PMP 125 bp overlap
ChIP Hep-G2 ENCSR000BVL.CREB1.Hep-G2 124 bp overlap
ChIP WA01 ENCSR000BSN.CREB1.WA01 432 bp overlap
CTBP2 2 datasets
ChIP H1 ENCFF329MAX 511 bp overlap
ChIP WA01 ENCSR000EUO.CTBP2.WA01 445 bp overlap
CTCF 8 datasets
Motif DE_12h DE_12h-CTCF_MA1929.2 31 bp overlap
ChIP GSC23 GSE139416.CTCF.GSC23 230 bp overlap
ChIP HepG2 ENCFF757EKU 86 bp overlap
ChIP MM1-S GSE43743.CTCF.MM1-S 62 bp overlap
ChIP anterior-temporal-cortex_fetal GSE116825.CTCF.anterior-temporal-cortex_fetal 154 bp overlap
ChIP endodermal cell ENCFF471YCZ 133 bp overlap
ChIP hESC_NAIVE GSE69646.CTCF.hESC_NAIVE 110 bp overlap
ChIP hiPSC_failed-genome-editing GSE106870.CTCF.hiPSC_failed-genome-editing 125 bp overlap
CTNNB1 1 dataset
ChIP hiPSC_D2 GSE132532.CTNNB1.hiPSC_D2 425 bp overlap
CUX1 1 dataset
ChIP Hep-G2 ENCSR049KIZ.CUX1.Hep-G2 179 bp overlap
Cebpa 1 dataset
ChIP BLaER1 ENCFF274GAT 287 bp overlap
E2F6 1 dataset
ChIP WA01 ENCSR000BSI.E2F6.WA01 68 bp overlap
EED 1 dataset
ChIP ProEs GSE59087.EED.ProEs 91 bp overlap
EGR1 1 dataset
ChIP Hep-G2 ENCSR026GSW.EGR1.Hep-G2 191 bp overlap
ELF1 1 dataset
ChIP HepG2 ENCFF367ZWV 107 bp overlap
ELL2 2 datasets
ChIP HeLa_DOX GSE40632.ELL2.HeLa_DOX 254 bp overlap
ChIP HeLa_EGF GSE40632.ELL2.HeLa_EGF 180 bp overlap
EP300 3 datasets
ChIP H1 ENCFF927IYK 297 bp overlap
ChIP WA01 ENCSR000AUQ.EP300.WA01 239 bp overlap
ChIP WA01 ENCSR000BKK.EP300.WA01 237 bp overlap
ESR1 1 dataset
ChIP Ishikawa_siNTC-72h GSE129803.ESR1.Ishikawa_siNTC-72h 318 bp overlap
ETS1 1 dataset
ChIP Hep-G2 ENCSR681WHQ.ETS1.Hep-G2 206 bp overlap
FOSL2 1 dataset
ChIP HepG2 ENCFF796NIA 255 bp overlap
FOXA1 1 dataset
ChIP breast-cancer_heregulin GSE101407.FOXA1.breast-cancer_heregulin 251 bp overlap
FOXO3 1 dataset
ChIP Hep-G2 GSE97661.FOXO3.Hep-G2 131 bp overlap
FOXP1 1 dataset
ChIP WTC11 ENCFF338WGC 384 bp overlap
GLIS1 1 dataset
Motif DE_12h DE_12h-GLIS1_MA0735.2 15 bp overlap
GLIS3 1 dataset
ChIP H9_plus GSE109562.GLIS3.H9_plus 382 bp overlap
HDAC2 1 dataset
ChIP WA01 ENCSR000AVB.HDAC2.WA01 324 bp overlap
HNF4A 1 dataset
ChIP IM95 GSE114018.HNF4A.IM95 183 bp overlap
JUN 7 datasets
ChIP DE_D1 S08-DE-d1-JUN-exp1 501 bp overlap
ChIP DE_D1 S40-DE-d1-JUN-exp2 395 bp overlap
ChIP DE_D2 S02-DE-d2-JUN-exp1 338 bp overlap
ChIP ESC S24-ESC-d0-JUN-exp1 314 bp overlap
ChIP ESC S34-ESC-d0-JUN-exp2 458 bp overlap
ChIP HUES-8 GSE109524.JUN.HUES-8 491 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.JUN.HUES-8_DE_ctrl 508 bp overlap
JUND 3 datasets
ChIP H1 ENCFF010YXS 301 bp overlap
ChIP WA01 ENCSR000BKP.JUND.WA01 214 bp overlap
ChIP WA01 ENCSR000EBZ.JUND.WA01 136 bp overlap
KDM4A 2 datasets
ChIP WA01 ENCSR000AVC.KDM4A.WA01 436 bp overlap
ChIP hiPSC_IID12 GSE106870.KDM4A.hiPSC_IID12 222 bp overlap
KLF17 1 dataset
ChIP HEK293 ENCSR065WUF.KLF17.HEK293 389 bp overlap
KLF5 1 dataset
ChIP HEK293_E419Q GSE88976.KLF5.HEK293_E419Q 490 bp overlap
KMT2C 1 dataset
ChIP BIN-67_lentivirus-SMARCA4 GSE117734.KMT2C.BIN-67_lentivirus-SMARCA4 251 bp overlap
LCORL 1 dataset
ChIP HepG2 ENCFF017FTI 473 bp overlap
MAX 2 datasets
ChIP H1 ENCFF914VQY 92 bp overlap
ChIP Hep-G2 ENCSR168DYA.MAX.Hep-G2 246 bp overlap
MAZ 1 dataset
ChIP IMR-90 ENCSR000EFF.MAZ.IMR-90 84 bp overlap
MED1 5 datasets
ChIP HCT-116 GSE121798.MED1.HCT-116 487 bp overlap
ChIP HCT-116_shLuc GSE121798.MED1.HCT-116_shLuc 494 bp overlap
ChIP HCT-116_shLuc_thaps GSE121798.MED1.HCT-116_shLuc_thaps 486 bp overlap
ChIP HCT-116_thaps GSE121798.MED1.HCT-116_thaps 407 bp overlap
ChIP HUVEC-C_siCTL GSE124892.MED1.HUVEC-C_siCTL 318 bp overlap
MED26 1 dataset
ChIP HEK293T GSE121024.MED26.HEK293T 179 bp overlap
MTERF4 1 dataset
ChIP HepG2 ENCFF831NAM 420 bp overlap
MXD4 1 dataset
ChIP Hep-G2 ENCSR441KFW.MXD4.Hep-G2 204 bp overlap
MYC 1 dataset
ChIP H1 ENCFF794ZJT 265 bp overlap
MYF5 1 dataset
Motif DE_12h DE_12h-MYF5_MA1641.2 8 bp overlap
MYOD1 2 datasets
ChIP IMR-90_GM GSE128527.MYOD1.IMR-90_GM 183 bp overlap
ChIP myoblast GSE50413.MYOD1.myoblast 157 bp overlap
NANOG 2 datasets
ChIP HUES-8 GSE109524.NANOG.HUES-8 391 bp overlap
ChIP WA01 ERP004238.NANOG.WA01 254 bp overlap
NELFE 1 dataset
ChIP HeLa GSE125534.NELFE.HeLa 231 bp overlap
NFKB2 1 dataset
ChIP HepG2 ENCFF165NTY 306 bp overlap
NIPBL 1 dataset
ChIP HCT-116_RAD21-mAC GSE104888.NIPBL.HCT-116_RAD21-mAC 202 bp overlap
Neurod2 2 datasets
Motif DE_12h DE_12h-Neurod2_MA0668.3 8 bp overlap
Motif DE_12h DE_12h-Neurod2_MA1993.2 6 bp overlap
Olig2 1 dataset
Motif DE_12h DE_12h-Olig2_MA1997.2 6 bp overlap
PARP1 1 dataset
ChIP MCF-10A GSE93038.PARP1.MCF-10A 385 bp overlap
PATZ1 2 datasets
ChIP HEK293 ENCSR966ULI.PATZ1.HEK293 254 bp overlap
ChIP HepG2 ENCFF723PFC 308 bp overlap
PHF8 1 dataset
ChIP Hep-G2 ENCSR604VAE.PHF8.Hep-G2 146 bp overlap
POLR2A 1 dataset
ChIP H1 ENCFF833NJP 411 bp overlap
POU5F1 3 datasets
ChIP BG03 GSE21614.POU5F1.BG03 435 bp overlap
ChIP DE_D1 DED1-OCT4_Batch_II 388 bp overlap
ChIP HUES-8 GSE109524.POU5F1.HUES-8 415 bp overlap
POU5F1_M 1 dataset
ChIP DE_D1 DED1-OCT4-M_Batch_II 374 bp overlap
PRDM14 1 dataset
ChIP hESC GSE138674.PRDM14.hESC 218 bp overlap
Ptf1A 1 dataset
Motif DE_12h DE_12h-Ptf1A_MA1619.2 8 bp overlap
RAD21 5 datasets
ChIP HeLa-S3_CTCF-s-OE GSE108869.RAD21.HeLa-S3_CTCF-s-OE 163 bp overlap
ChIP HepG2 ENCFF360ZSW 217 bp overlap
ChIP hiPSC_HUES9 GSE106870.RAD21.hiPSC_HUES9 132 bp overlap
ChIP hiPSC_parental-line GSE106870.RAD21.hiPSC_parental-line 113 bp overlap
ChIP hiPSC_parental-line GSE106870.RAD21.hiPSC_parental-line 200 bp overlap
RBBP5 2 datasets
ChIP H1 ENCFF905HFL 420 bp overlap
ChIP WA01 ENCSR000AQC.RBBP5.WA01 511 bp overlap
REL 1 dataset
Motif DE_12h DE_12h-REL_MA0101.1 10 bp overlap
RELA 3 datasets
ChIP HAEC_TNFa_4h GSE89970.RELA.HAEC_TNFa_4h 153 bp overlap
ChIP HeLa-B2_TNFA GSE24518.RELA.HeLa-B2_TNFA 198 bp overlap
ChIP SGBS_TNF GSE64233.RELA.SGBS_TNF 301 bp overlap
RXRA 1 dataset
ChIP H1 ENCFF570NHK 201 bp overlap
SAP130 1 dataset
ChIP HepG2 ENCFF892EHZ 416 bp overlap
SAP30 2 datasets
ChIP H1 ENCFF149IOE 391 bp overlap
ChIP WA01 ENCSR000ATR.SAP30.WA01 399 bp overlap
SFPQ 1 dataset
ChIP HepG2 ENCFF145CDF 456 bp overlap
SIN3A 4 datasets
ChIP H1 ENCFF042ZSL 479 bp overlap
ChIP WA01 ENCSR000EBO.SIN3A.WA01 276 bp overlap
ChIP hiPSC_IB12 GSE106870.SIN3A.hiPSC_IB12 273 bp overlap
ChIP hiPSC_IID12 GSE106870.SIN3A.hiPSC_IID12 395 bp overlap
SMAD1 1 dataset
ChIP Hep-G2 ENCSR213QOZ.SMAD1.Hep-G2 225 bp overlap
SMAD2-3 1 dataset
ChIP HUES-8_DE_ctrl GSE109524.SMAD2-3.HUES-8_DE_ctrl 351 bp overlap
SMARCA4 5 datasets
ChIP BIN-67_lentivirus-SMARCA4 GSE117734.SMARCA4.BIN-67_lentivirus-SMARCA4 236 bp overlap
ChIP TTC-1240_delC GSE124903.SMARCA4.TTC-1240_delC 313 bp overlap
ChIP TTC-549_Dox GSE71504.SMARCA4.TTC-549_Dox 197 bp overlap
ChIP hiPSC GSE124903.SMARCA4.hiPSC 326 bp overlap
ChIP hiPSC_WT GSE124903.SMARCA4.hiPSC_WT 301 bp overlap
SMARCB1 2 datasets
ChIP hiPSC GSE124903.SMARCB1.hiPSC 184 bp overlap
ChIP proliferating-human-fibroblast_ASO_Ctrl GSE131711.SMARCB1.proliferating-human-fibroblast_ASO_Ctrl 161 bp overlap
SMARCC1 3 datasets
ChIP BIN-67_lentivirus-SMARCA4-K785R GSE117734.SMARCC1.BIN-67_lentivirus-SMARCA4-K785R 260 bp overlap
ChIP hiPSC GSE124903.SMARCC1.hiPSC 163 bp overlap
ChIP hiPSC_WT GSE124903.SMARCC1.hiPSC_WT 315 bp overlap
SMC3 3 datasets
ChIP HeLa GSE126990.SMC3.HeLa 210 bp overlap
ChIP HeLa-Kyoto GSE102884.SMC3.HeLa-Kyoto 210 bp overlap
ChIP HeLa-Kyoto GSE138405.SMC3.HeLa-Kyoto 210 bp overlap
SOX2 1 dataset
ChIP HNSC GSE69479.SOX2.HNSC 224 bp overlap
SP1 5 datasets
ChIP H1 ENCFF263FUH 145 bp overlap
ChIP HEK293 GSE76494.SP1.HEK293 153 bp overlap
ChIP Hep-G2 ENCSR460YAM.SP1.Hep-G2 189 bp overlap
ChIP WA01 ENCSR000BIR.SP1.WA01 342 bp overlap
ChIP WTC11 ENCFF688PEU 413 bp overlap
SP4 3 datasets
Motif DE_12h DE_12h-SP4_MA0685.2 9 bp overlap
ChIP HEK293 GSE76494.SP4.HEK293 149 bp overlap
ChIP WA01 ENCSR000BQV.SP4.WA01 308 bp overlap
SP5 1 dataset
Motif DE_12h DE_12h-SP5_MA1965.2 6 bp overlap
STAG1 2 datasets
ChIP HeLa GSE126990.STAG1.HeLa 64 bp overlap
ChIP HeLa-Kyoto GSE138405.STAG1.HeLa-Kyoto 64 bp overlap
SUPT5H 1 dataset
ChIP HeLa GSE125534.SUPT5H.HeLa 223 bp overlap
T 1 dataset
ChIP H9_MESODERM GSE60606.T.H9_MESODERM 183 bp overlap
TAF1 1 dataset
ChIP WA01 ENCSR000BHO.TAF1.WA01 153 bp overlap
TBP 2 datasets
ChIP H1 ENCFF859IIO 377 bp overlap
ChIP WA01 ENCSR000ECB.TBP.WA01 141 bp overlap
TCF12 2 datasets
ChIP H1 ENCFF203EBH 251 bp overlap
ChIP WA01 ENCSR000BIT.TCF12.WA01 293 bp overlap
TCF3 1 dataset
ChIP NPC GSE154479.TCF3.NPC 269 bp overlap
TEAD1 3 datasets
ChIP HepG2 ENCFF661PNM 377 bp overlap
ChIP WTC11 ENCFF502QUV 405 bp overlap
ChIP pancreas_12D ERP008682.TEAD1.pancreas_12D 168 bp overlap
TEAD3 1 dataset
ChIP HepG2 ENCFF054UUL 371 bp overlap
TEAD4 6 datasets
ChIP ESC S26-ESC-d0-TEAD4-exp1 468 bp overlap
ChIP H1 ENCFF778PAX 211 bp overlap
ChIP HepG2 ENCFF250NXO 311 bp overlap
ChIP Ishikawa ENCSR000BSW.TEAD4.Ishikawa 168 bp overlap
ChIP WTC11 ENCFF114TZS 341 bp overlap
ChIP hESC GSE99202.TEAD4.hESC 247 bp overlap
TFAP2C 1 dataset
ChIP UCLA1-hESCs_TFAP2C--_1d_TFAP2C_Induc GSE101074.TFAP2C.UCLA1-hESCs_TFAP2C--_1d_TFAP2C_Induc 442 bp overlap
TFAP4 3 datasets
ChIP Hep-G2 ENCSR103SZL.TFAP4.Hep-G2 233 bp overlap
ChIP HepG2 ENCFF030SRU 319 bp overlap
ChIP HepG2 ENCFF932XOY 397 bp overlap
THRB 1 dataset
ChIP HepG2 ENCFF476INC 60 bp overlap
TP53 1 dataset
ChIP WTC11 ENCFF359JCU 401 bp overlap
TRIM25 1 dataset
ChIP MDA-MB-231 GSE79588.TRIM25.MDA-MB-231 240 bp overlap
TRIM28 1 dataset
ChIP hESC GSE115387.TRIM28.hESC 184 bp overlap
Tcf12 1 dataset
Motif DE_12h DE_12h-Tcf12_MA0521.3 6 bp overlap
Twist2 1 dataset
Motif DE_12h DE_12h-Twist2_MA0633.3 6 bp overlap
U2AF1 1 dataset
ChIP Hep-G2 GSE120104.U2AF1.Hep-G2 167 bp overlap
YAP1 1 dataset
ChIP hiPSC GSE111930.YAP1.hiPSC 124 bp overlap
YY1 3 datasets
ChIP Hep-G2 ENCSR000BNT.YY1.Hep-G2 217 bp overlap
ChIP Huh-7 GSE97411.YY1.Huh-7 309 bp overlap
ChIP WA01 ENCSR000BKD.YY1.WA01 376 bp overlap
ZBED4 1 dataset
ChIP Hep-G2 ENCSR409PMR.ZBED4.Hep-G2 146 bp overlap
ZBTB18 1 dataset
Motif DE_12h DE_12h-ZBTB18_MA0698.2 11 bp overlap
ZBTB21 1 dataset
ChIP WTC11 ENCFF677ZYY 380 bp overlap
ZBTB7A 4 datasets
ChIP Hep-G2 ENCSR000BQA.ZBTB7A.Hep-G2 134 bp overlap
ChIP HepG2 ENCFF492YYQ 388 bp overlap
ChIP K-562 ENCSR000BME.ZBTB7A.K-562 126 bp overlap
ChIP VCaP_FBS GSE123089.ZBTB7A.VCaP_FBS 215 bp overlap
ZFHX2 1 dataset
ChIP HEK293 ENCFF167TUA 271 bp overlap
ZFX 2 datasets
ChIP HEK293T_FLAG GSE145160.ZFX.HEK293T_FLAG 230 bp overlap
ChIP HepG2 ENCFF016NZF 511 bp overlap
ZNF143 2 datasets
ChIP FLP143HA_T4 GSE39263.ZNF143.FLP143HA_T4 163 bp overlap
ChIP HeLa GSE39263.ZNF143.HeLa 78 bp overlap
ZNF263 1 dataset
Motif DE_12h DE_12h-ZNF263_MA0528.3 7 bp overlap
ZNF281 1 dataset
ChIP HepG2 ENCFF585QNU 325 bp overlap
ZNF398 1 dataset
ChIP BG01V GSE133630.ZNF398.BG01V 186 bp overlap
ZNF506 1 dataset
ChIP HEK293T GSE78099.ZNF506.HEK293T 62 bp overlap
ZNF687 1 dataset
ChIP HepG2 ENCFF653WIX 511 bp overlap
ZNF692 1 dataset
Motif DE_12h DE_12h-ZNF692_MA1986.2 8 bp overlap
ZNF707 1 dataset
Motif DE_12h DE_12h-ZNF707_MA1715.1 15 bp overlap
Zic2 1 dataset
Motif DE_12h DE_12h-Zic2_MA1629.2 9 bp overlap