chr15 : 84,979,692 84,980,043
351 bp 156 TFs 1 linked gene
This 351 bp open chromatin element is linked to PDE8A and is bound by 156 transcription factors.
Linked Genes
1 gene
Gene Expression Dist. to TSS Distance Link type
PDE8A 468 bp At TSS Proximity
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr15:84,974,692 – 84,985,043
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
156 transcription factors
Source
Cell type
ASH2L 1 dataset
ChIP WA01 ENCSR850KIP.ASH2L.WA01 246 bp overlap
ATF3 1 dataset
ChIP Hep-G2 ENCSR402ZCY.ATF3.Hep-G2 254 bp overlap
BCOR 3 datasets
ChIP K-562 ENCSR808AKZ.BCOR.K-562 253 bp overlap
ChIP K562 ENCFF343XWA 247 bp overlap
ChIP WA01_RNF2-R GSE104690.BCOR.WA01_RNF2-R 331 bp overlap
BRD1 1 dataset
ChIP HUES-64 GSE104059.BRD1.HUES-64 351 bp overlap
BRD4 1 dataset
ChIP K-562_iBET-IFNG GSE138084.BRD4.K-562_iBET-IFNG 203 bp overlap
CBFB 1 dataset
ChIP Hep-G2 ENCSR479RZV.CBFB.Hep-G2 242 bp overlap
CDKN1B 3 datasets
ChIP MDA-BoM-1833_shp27 GSE112444.CDKN1B.MDA-BoM-1833_shp27 204 bp overlap
ChIP MDA-MB-231 GSE112444.CDKN1B.MDA-MB-231 171 bp overlap
ChIP MDA-MB-231_p27CK-DD GSE112444.CDKN1B.MDA-MB-231_p27CK-DD 292 bp overlap
CTCF 1 dataset
ChIP MCF-7 ENCFF424NQR 191 bp overlap
DRAP1 1 dataset
ChIP Hep-G2 ENCSR765MKZ.DRAP1.Hep-G2 133 bp overlap
E2F6 3 datasets
ChIP K-562 ENCSR000BLI.E2F6.K-562 102 bp overlap
ChIP K562 ENCFF136LTS 307 bp overlap
ChIP WA01 ENCSR000BSI.E2F6.WA01 116 bp overlap
EGR1 1 dataset
ChIP Hep-G2 ENCSR026GSW.EGR1.Hep-G2 297 bp overlap
ELK1::HOXB13 2 datasets
Motif DE_12h DE_12h-ELK1HOXB13_MA1932.2 15 bp overlap
Motif DE_24h DE_24h-ELK1HOXB13_MA1932.2 15 bp overlap
ESRRA 4 datasets
Motif DE_12h DE_12h-ESRRA_MA0592.4 9 bp overlap
Motif DE_24h DE_24h-ESRRA_MA0592.4 9 bp overlap
ChIP K562 ENCFF968PEP 351 bp overlap
ChIP MCF-7 ENCSR954WVZ.ESRRA.MCF-7 209 bp overlap
ESRRB 2 datasets
Motif DE_12h DE_12h-ESRRB_MA0141.4 10 bp overlap
Motif DE_24h DE_24h-ESRRB_MA0141.4 10 bp overlap
ETV4 2 datasets
ChIP HepG2 ENCFF534CDD 266 bp overlap
ChIP HepG2 ENCFF534CDD 66 bp overlap
ETV6 1 dataset
ChIP WTC11 ENCFF812SCD 320 bp overlap
EZH2 8 datasets
ChIP GM23338 ENCSR045YHA.EZH2.GM23338 279 bp overlap
ChIP H1 ENCFF232NZA 351 bp overlap
ChIP H1 ENCFF232NZA 211 bp overlap
ChIP WA01 ENCSR000ASY.EZH2.WA01 256 bp overlap
ChIP neural progenitor cell ENCFF018MKA 204 bp overlap
ChIP neural progenitor cell ENCFF018MKA 204 bp overlap
ChIP neural progenitor cell ENCFF472NFV 351 bp overlap
ChIP neural progenitor cell ENCFF472NFV 351 bp overlap
Esrrg 2 datasets
Motif DE_12h DE_12h-Esrrg_MA0643.2 9 bp overlap
Motif DE_24h DE_24h-Esrrg_MA0643.2 9 bp overlap
FOXA1 2 datasets
ChIP T-47D_shRNF2 GSE137579.FOXA1.T-47D_shRNF2 78 bp overlap
ChIP T-47D_shRNF2_8h GSE137579.FOXA1.T-47D_shRNF2_8h 72 bp overlap
FOXA2 3 datasets
ChIP BJ1-hTERT_FoxHnf1aCoExp GSE90454.FOXA2.BJ1-hTERT_FoxHnf1aCoExp 266 bp overlap
ChIP HepG2 ENCFF533COJ 156 bp overlap
ChIP PANC-1_HNF1B-OE GSE119930.FOXA2.PANC-1_HNF1B-OE 72 bp overlap
GABPA 1 dataset
ChIP K-562 ENCSR000BLO.GABPA.K-562 197 bp overlap
GMEB1 1 dataset
ChIP Hep-G2 ENCSR780OXL.GMEB1.Hep-G2 216 bp overlap
HCFC1 1 dataset
ChIP Hep-G2 ENCSR529JYA.HCFC1.Hep-G2 128 bp overlap
HDAC1 2 datasets
ChIP Hep-G2 ENCSR750OWO.HDAC1.Hep-G2 158 bp overlap
ChIP K-562 ENCSR711VWL.HDAC1.K-562 139 bp overlap
HDAC2 3 datasets
ChIP K-562 ENCSR893WSB.HDAC2.K-562 204 bp overlap
ChIP K-562 ENCSR075HTM.HDAC2.K-562 160 bp overlap
ChIP K562 ENCFF744ALD 294 bp overlap
HDGF 2 datasets
ChIP K-562 ENCSR197ALX.HDGF.K-562 183 bp overlap
ChIP K-562 ENCSR563YDA.HDGF.K-562 194 bp overlap
HNF1B 1 dataset
ChIP Hep-G2 ENCSR127XTZ.HNF1B.Hep-G2 274 bp overlap
HOXA3 1 dataset
ChIP Hep-G2 ENCSR999JKC.HOXA3.Hep-G2 214 bp overlap
HOXB2::ELK1 2 datasets
Motif DE_12h DE_12h-HOXB2ELK1_MA1957.1 14 bp overlap
Motif DE_24h DE_24h-HOXB2ELK1_MA1957.1 14 bp overlap
HOXD12::ELK1 2 datasets
Motif DE_12h DE_12h-HOXD12ELK1_MA1958.2 13 bp overlap
Motif DE_24h DE_24h-HOXD12ELK1_MA1958.2 13 bp overlap
INO80 1 dataset
ChIP Hep-G2 GSE107730.INO80.Hep-G2 347 bp overlap
JARID2 2 datasets
ChIP MRC-5_IPS25_CTR GSE48515.JARID2.MRC-5_IPS25_CTR 351 bp overlap
ChIP MRC-5_IPS25_MEG3 GSE48515.JARID2.MRC-5_IPS25_MEG3 351 bp overlap
JUN 3 datasets
ChIP MDA-BoM-1833 GSE112444.JUN.MDA-BoM-1833 190 bp overlap
ChIP MDA-BoM-1833_shp27 GSE112444.JUN.MDA-BoM-1833_shp27 308 bp overlap
ChIP MDA-MB-231 GSE112444.JUN.MDA-MB-231 239 bp overlap
JUND 1 dataset
ChIP Hep-G2 ENCSR000BGK.JUND.Hep-G2 110 bp overlap
KDM1A 9 datasets
ChIP A-549 ENCSR639GWS.KDM1A.A-549 190 bp overlap
ChIP K-562 GSE117944.KDM1A.K-562 351 bp overlap
ChIP K-562_Wnt GSE117944.KDM1A.K-562_Wnt 200 bp overlap
ChIP K562 ENCFF128TYE 281 bp overlap
ChIP K562 ENCFF133OLU 286 bp overlap
ChIP K562 ENCFF934ZRG 320 bp overlap
ChIP pancreatic-progenitor_GT GSE104840.KDM1A.pancreatic-progenitor_GT 220 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.KDM1A.pancreatic-progenitor_PP1 200 bp overlap
ChIP pancreatic-progenitor_PP2-KDM1A-inh GSE104840.KDM1A.pancreatic-progenitor_PP2-KDM1A-inh 184 bp overlap
KDM2B 1 dataset
ChIP K562 ENCFF392YVR 256 bp overlap
KDM3A 1 dataset
ChIP Hep-G2 ENCSR387JKT.KDM3A.Hep-G2 186 bp overlap
KDM4A 1 dataset
ChIP WA01 ENCSR000AVC.KDM4A.WA01 206 bp overlap
KDM5B 2 datasets
ChIP Hep-G2 ENCSR227MRE.KDM5B.Hep-G2 251 bp overlap
ChIP SUM185_SHCTCF GSE46055.KDM5B.SUM185_SHCTCF 214 bp overlap
KLF1 1 dataset
Motif DE_12h DE_12h-KLF1_MA0493.3 8 bp overlap
KLF10 1 dataset
Motif DE_12h DE_12h-KLF10_MA1511.2 9 bp overlap
KLF12 1 dataset
Motif DE_12h DE_12h-KLF12_MA0742.2 9 bp overlap
KLF14 1 dataset
Motif DE_12h DE_12h-KLF14_MA0740.2 9 bp overlap
KLF15 1 dataset
Motif DE_12h DE_12h-KLF15_MA1513.2 8 bp overlap
KLF2 1 dataset
Motif DE_12h DE_12h-KLF2_MA1515.2 8 bp overlap
KLF4 1 dataset
Motif DE_12h DE_12h-KLF4_MA0039.5 8 bp overlap
KLF5 1 dataset
Motif DE_12h DE_12h-KLF5_MA0599.1 10 bp overlap
KLF7 1 dataset
Motif DE_12h DE_12h-KLF7_MA1959.2 8 bp overlap
KMT2A 2 datasets
ChIP HEK293T_N-term_shCtrl_TASP1-KO GSE90762.KMT2A.HEK293T_N-term_shCtrl_TASP1-KO 190 bp overlap
ChIP Hep-G2 ENCSR692GFR.KMT2A.Hep-G2 171 bp overlap
MAX 1 dataset
ChIP K-562 ENCSR000EFV.MAX.K-562 167 bp overlap
MAZ 2 datasets
Motif ES_0h ES_0h-MAZ_MA1522.2 8 bp overlap
ChIP K-562 ENCSR000EFX.MAZ.K-562 111 bp overlap
MED1 1 dataset
ChIP HUVEC-C_siCTL GSE124892.MED1.HUVEC-C_siCTL 246 bp overlap
MNX1 1 dataset
ChIP Hep-G2 ENCSR108MKR.MNX1.Hep-G2 215 bp overlap
MTA1 1 dataset
ChIP Hep-G2 ENCSR983KRB.MTA1.Hep-G2 276 bp overlap
MXD4 1 dataset
ChIP Hep-G2 ENCSR441KFW.MXD4.Hep-G2 167 bp overlap
MYBL2 1 dataset
ChIP Hep-G2 ENCSR581KCO.MYBL2.Hep-G2 123 bp overlap
MYC 1 dataset
ChIP P493-6_scramble-shRNA GSE101853.MYC.P493-6_scramble-shRNA 174 bp overlap
MYCN 2 datasets
ChIP COG-N-415 GSE138295.MYCN.COG-N-415 297 bp overlap
ChIP LA-N-5 GSE138295.MYCN.LA-N-5 263 bp overlap
NANOG 2 datasets
ChIP HUES-8 GSE109524.NANOG.HUES-8 276 bp overlap
ChIP WA01 ENCSR000BMT.NANOG.WA01 131 bp overlap
NELFE 2 datasets
ChIP K-562_HS GSE112379.NELFE.K-562_HS 314 bp overlap
ChIP K-562_NHS GSE112379.NELFE.K-562_NHS 202 bp overlap
NFE2L2 1 dataset
ChIP A-549 GSE113497.NFE2L2.A-549 234 bp overlap
NFKB1 1 dataset
ChIP HEK293T GSE129618.NFKB1.HEK293T 216 bp overlap
NR1I3 1 dataset
Motif DE_12h DE_12h-NR1I3_MA1534.2 8 bp overlap
NR2C1 2 datasets
Motif DE_12h DE_12h-NR2C1_MA1535.2 6 bp overlap
Motif DE_24h DE_24h-NR2C1_MA1535.2 6 bp overlap
NR2C2 3 datasets
Motif DE_12h DE_12h-NR2C2_MA0504.2 14 bp overlap
Motif DE_12h DE_12h-NR2C2_MA1536.2 6 bp overlap
Motif DE_24h DE_24h-NR2C2_MA1536.2 6 bp overlap
NR2F2 3 datasets
Motif DE_12h DE_12h-NR2F2_MA1111.2 7 bp overlap
Motif DE_24h DE_24h-NR2F2_MA1111.2 7 bp overlap
ChIP liver ENCSR168SMX.NR2F2.liver 163 bp overlap
NR2F6 1 dataset
ChIP Hep-G2 ENCSR518WPL.NR2F6.Hep-G2 242 bp overlap
NR5A1 2 datasets
ChIP Hep-G2 ENCSR310OZS.NR5A1.Hep-G2 168 bp overlap
ChIP HepG2 ENCFF970YZO 309 bp overlap
NR5A2 1 dataset
ChIP A549 ENCFF834RVE 351 bp overlap
Nr1H2 2 datasets
Motif DE_12h DE_12h-Nr1H2_MA1996.2 6 bp overlap
Motif DE_24h DE_24h-Nr1H2_MA1996.2 6 bp overlap
Nr1H4 2 datasets
Motif DE_12h DE_12h-Nr1H4_MA1110.3 6 bp overlap
Motif DE_24h DE_24h-Nr1H4_MA1110.3 6 bp overlap
Nr1h3 2 datasets
Motif DE_12h DE_12h-Nr1h3_MA2337.1 6 bp overlap
Motif DE_24h DE_24h-Nr1h3_MA2337.1 6 bp overlap
Nr5A2 4 datasets
Motif DE_12h DE_12h-Nr5A2_MA0505.3 9 bp overlap
Motif DE_12h DE_12h-Nr5A2_MA0505.3 9 bp overlap
Motif DE_24h DE_24h-Nr5A2_MA0505.3 9 bp overlap
Motif DE_24h DE_24h-Nr5A2_MA0505.3 9 bp overlap
OSR2 2 datasets
Motif DE_12h DE_12h-OSR2_MA1646.2 8 bp overlap
Motif DE_24h DE_24h-OSR2_MA1646.2 8 bp overlap
PARP1 1 dataset
ChIP MCF-10A GSE93038.PARP1.MCF-10A 280 bp overlap
PAXIP1 1 dataset
ChIP HepG2 ENCFF526NOJ 240 bp overlap
PCBP1 2 datasets
ChIP HepG2 ENCFF447SRJ 318 bp overlap
ChIP HepG2 ENCFF447SRJ 274 bp overlap
PGR 1 dataset
ChIP MCF-7_PROG GSE68355.PGR.MCF-7_PROG 183 bp overlap
PHF8 1 dataset
ChIP K-562 ENCSR000AQH.PHF8.K-562 139 bp overlap
PHIP 2 datasets
ChIP HCT-116_MLL1-KO__ab833 GSE101646.PHIP.HCT-116_MLL1-KO__ab833 351 bp overlap
ChIP HCT-116_ab833 GSE101646.PHIP.HCT-116_ab833 300 bp overlap
PPARG 1 dataset
ChIP Hep-G2 ENCSR130VQL.PPARG.Hep-G2 140 bp overlap
PRDM15 1 dataset
ChIP WTC11 ENCFF108TMF 189 bp overlap
PTBP1 2 datasets
ChIP Hep-G2 GSE120104.PTBP1.Hep-G2 198 bp overlap
ChIP Hep-G2 ENCSR156APP.PTBP1.Hep-G2 210 bp overlap
Ppara 2 datasets
Motif DE_12h DE_12h-Ppara_MA2338.1 7 bp overlap
Motif DE_24h DE_24h-Ppara_MA2338.1 7 bp overlap
RBBP5 1 dataset
ChIP K-562 ENCSR000AQI.RBBP5.K-562 171 bp overlap
RBM39 2 datasets
ChIP Hep-G2 ENCSR339JTP.RBM39.Hep-G2 210 bp overlap
ChIP Hep-G2 GSE120104.RBM39.Hep-G2 206 bp overlap
RCOR1 1 dataset
ChIP K562 ENCFF216EEJ 297 bp overlap
REST 1 dataset
ChIP A-549 ENCSR000BQP.REST.A-549 61 bp overlap
RYBP 1 dataset
ChIP WA01 GSE104690.RYBP.WA01 351 bp overlap
Rarg 1 dataset
Motif DE_12h DE_12h-Rarg_MA0860.1 17 bp overlap
SIRT6 1 dataset
ChIP SK-MEL-239_L-C-B GSE102813.SIRT6.SK-MEL-239_L-C-B 275 bp overlap
SKIL 1 dataset
ChIP HepG2 ENCFF823HPQ 96 bp overlap
SMARCA4 1 dataset
ChIP A-549_AG15689 GSE132290.SMARCA4.A-549_AG15689 256 bp overlap
SOX13 1 dataset
ChIP HepG2 ENCFF062VSQ 293 bp overlap
SP1 2 datasets
Motif DE_12h DE_12h-SP1_MA0079.5 9 bp overlap
ChIP WA01 ENCSR000BIR.SP1.WA01 92 bp overlap
SP140L 1 dataset
ChIP HepG2 ENCFF203CWF 181 bp overlap
SP2 4 datasets
Motif DE_12h DE_12h-SP2_MA0516.3 9 bp overlap
ChIP HEK293 ENCFF181QXT 263 bp overlap
ChIP HEK293 ENCSR807LQP.SP2.HEK293 263 bp overlap
ChIP HEK293 GSE76494.SP2.HEK293 185 bp overlap
SP4 1 dataset
Motif DE_12h DE_12h-SP4_MA0685.2 9 bp overlap
SP5 2 datasets
Motif DE_12h DE_12h-SP5_MA1965.2 6 bp overlap
Motif ES_0h ES_0h-SP5_MA1965.2 6 bp overlap
SRF 1 dataset
ChIP K-562 ENCSR582IAO.SRF.K-562 143 bp overlap
STAT3 1 dataset
Motif DE_12h DE_12h-STAT3_MA0144.3 9 bp overlap
SUZ12 2 datasets
ChIP H1 ENCFF881NFR 351 bp overlap
ChIP H1 ENCFF881NFR 351 bp overlap
Stat5a 1 dataset
Motif DE_12h DE_12h-Stat5a_MA1624.2 9 bp overlap
TAF1 1 dataset
ChIP Hep-G2 ENCSR918OKK.TAF1.Hep-G2 341 bp overlap
TBP 1 dataset
ChIP K-562 GSE55306.TBP.K-562 244 bp overlap
TFAP2C 1 dataset
ChIP UCLA1-hESCs_TFAP2C--_1d_TFAP2C_Induc GSE101074.TFAP2C.UCLA1-hESCs_TFAP2C--_1d_TFAP2C_Induc 194 bp overlap
TFDP2 1 dataset
ChIP Hep-G2 ENCSR069JKP.TFDP2.Hep-G2 171 bp overlap
TP53 2 datasets
ChIP H9_ectoderm GSE142050.TP53.H9_ectoderm 348 bp overlap
ChIP MDA-MB-231 GSE95303.TP53.MDA-MB-231 278 bp overlap
TP63 1 dataset
ChIP breast-organoid GSE113909.TP63.breast-organoid 123 bp overlap
U2AF1 1 dataset
ChIP Hep-G2 GSE120104.U2AF1.Hep-G2 167 bp overlap
USF2 1 dataset
ChIP HepG2 ENCFF433IUE 59 bp overlap
Vdr 1 dataset
Motif DE_12h DE_12h-Vdr_MA0693.4 7 bp overlap
WT1 1 dataset
ChIP HEK293 ENCSR966PJJ.WT1.HEK293 252 bp overlap
YY1 1 dataset
ChIP Huh-7 GSE97411.YY1.Huh-7 291 bp overlap
ZBED4 2 datasets
Motif DE_12h DE_12h-ZBED4_MA2328.1 10 bp overlap
ChIP Hep-G2 ENCSR409PMR.ZBED4.Hep-G2 176 bp overlap
ZEB1 1 dataset
ChIP HepG2 ENCFF808RQT 273 bp overlap
ZFAT 1 dataset
ChIP HepG2 ENCFF236QRV 104 bp overlap
ZFP14 3 datasets
Motif DE_12h DE_12h-ZFP14_MA1972.1 15 bp overlap
Motif DE_24h DE_24h-ZFP14_MA1972.1 15 bp overlap
Motif ES_0h ES_0h-ZFP14_MA1972.1 15 bp overlap
ZFP36 1 dataset
ChIP K-562 ENCSR776CYN.ZFP36.K-562 319 bp overlap
ZFP64 1 dataset
ChIP Hep-G2 ENCSR487CPI.ZFP64.Hep-G2 131 bp overlap
ZKSCAN5 2 datasets
Motif DE_12h DE_12h-ZKSCAN5_MA1652.2 9 bp overlap
Motif ES_0h ES_0h-ZKSCAN5_MA1652.2 9 bp overlap
ZMYM3 7 datasets
ChIP Hep-G2 ENCSR848YWD.ZMYM3.Hep-G2 264 bp overlap
ChIP Hep-G2_AC_JH39-2-2B9 GSE97661.ZMYM3.Hep-G2_AC_JH39-2-2B9 164 bp overlap
ChIP Hep-G2_Ab_JH39-2-2F10 GSE97661.ZMYM3.Hep-G2_Ab_JH39-2-2F10 285 bp overlap
ChIP HepG2 ENCFF408KTI 151 bp overlap
ChIP K-562 ENCSR102KIN.ZMYM3.K-562 193 bp overlap
ChIP K-562_Ab_JH39-2-2F10 GSE97661.ZMYM3.K-562_Ab_JH39-2-2F10 183 bp overlap
ChIP K562 ENCFF361LXT 219 bp overlap
ZNF121 1 dataset
ChIP WTC11 ENCFF291API 214 bp overlap
ZNF140 1 dataset
Motif ES_0h ES_0h-ZNF140_MA1589.2 19 bp overlap
ZNF165 2 datasets
ChIP K-562 ENCSR172XJS.ZNF165.K-562 214 bp overlap
ChIP K562 ENCFF039BMN 337 bp overlap
ZNF24 1 dataset
ChIP K-562 ENCSR099NCH.ZNF24.K-562 287 bp overlap
ZNF257 2 datasets
Motif DE_12h DE_12h-ZNF257_MA1710.2 10 bp overlap
ChIP HEK293T GSE78099.ZNF257.HEK293T 239 bp overlap
ZNF263 1 dataset
Motif DE_12h DE_12h-ZNF263_MA0528.3 7 bp overlap
ZNF3 1 dataset
ChIP Hep-G2 ENCSR182QWU.ZNF3.Hep-G2 160 bp overlap
ZNF335 1 dataset
ChIP HEK293 ENCFF784SLD 152 bp overlap
ZNF418 3 datasets
Motif DE_12h DE_12h-ZNF418_MA1980.1 15 bp overlap
Motif DE_24h DE_24h-ZNF418_MA1980.1 15 bp overlap
Motif ES_0h ES_0h-ZNF418_MA1980.1 15 bp overlap
ZNF44 1 dataset
ChIP Hep-G2 ENCSR914HPP.ZNF44.Hep-G2 306 bp overlap
ZNF530 2 datasets
Motif DE_12h DE_12h-ZNF530_MA1981.2 14 bp overlap
Motif DE_24h DE_24h-ZNF530_MA1981.2 14 bp overlap
ZNF592 1 dataset
ChIP K562 ENCFF547OSS 50 bp overlap
ZNF607 1 dataset
ChIP HepG2 ENCFF118ANP 351 bp overlap
ZNF768 3 datasets
Motif DE_12h DE_12h-ZNF768_MA1731.2 9 bp overlap
Motif DE_24h DE_24h-ZNF768_MA1731.2 9 bp overlap
ChIP HepG2 ENCFF388QCK 335 bp overlap
ZNF800 1 dataset
ChIP HepG2 ENCFF840FYM 351 bp overlap