chr3 : 71,247,266 71,247,567
301 bp 150 TFs 0 linked genes
This 301 bp open chromatin element has no linked target genes and is bound by 150 transcription factors.
Linked Genes
No linked genes
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr3:71,242,266 – 71,252,567
This element Other elements ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
150 transcription factors
Source
Cell type
AR 2 datasets
ChIP MCF-7 GSE48930.AR.MCF-7 118 bp overlap
ChIP prostate_fetal-fibroblast GSE90772.AR.prostate_fetal-fibroblast 258 bp overlap
ARID5B 1 dataset
ChIP HepG2 ENCFF964FWK 181 bp overlap
ARNTL 1 dataset
ChIP Hep-G2 ENCSR794LVK.ARNTL.Hep-G2 211 bp overlap
ASH2L 2 datasets
ChIP Hep-G2 ENCSR508LMH.ASH2L.Hep-G2 239 bp overlap
ChIP WA01 ENCSR850KIP.ASH2L.WA01 109 bp overlap
ATF7 1 dataset
ChIP K-562 ENCSR972ZBV.ATF7.K-562 84 bp overlap
BNC2 2 datasets
ChIP SK-N-SH ENCFF174EMC 301 bp overlap
ChIP SK-N-SH_all-trans-retinoic-acid_6-uM_48-hour ENCSR754GYI.BNC2.SK-N-SH_all-trans-retinoic-acid_6-uM_48-hour 280 bp overlap
BRD3 1 dataset
ChIP HEK293T GSE39579.BRD3.HEK293T 111 bp overlap
BRD4 4 datasets
ChIP HEK293_sgNT GSE129407.BRD4.HEK293_sgNT 226 bp overlap
ChIP Kelly_sensitive GSE103030.BRD4.Kelly_sensitive 206 bp overlap
ChIP RH4_shCHD4 GSE140115.BRD4.RH4_shCHD4 104 bp overlap
ChIP SK-N-BE2-C_Resistant-veh GSE107706.BRD4.SK-N-BE2-C_Resistant-veh 256 bp overlap
CASZ1 1 dataset
ChIP rhabdomyosarcoma_DOX GSE126142.CASZ1.rhabdomyosarcoma_DOX 148 bp overlap
CBFB 1 dataset
ChIP Hep-G2 ENCSR479RZV.CBFB.Hep-G2 150 bp overlap
CDX2 1 dataset
ChIP LS180_125 GSE31939.CDX2.LS180_125 207 bp overlap
CEBPA 1 dataset
ChIP HepG2 ENCFF175DFS 284 bp overlap
CHD1 1 dataset
ChIP WA01 ENCSR000AQK.CHD1.WA01 163 bp overlap
CHD7 1 dataset
ChIP hiPSC_derived_lt-NES GSE108506.CHD7.hiPSC_derived_lt-NES 289 bp overlap
CSDC2 2 datasets
ChIP SK-N-SH ENCFF868MXA 273 bp overlap
ChIP SK-N-SH_all-trans-retinoic-acid_6-uM_48-hour ENCSR474KBG.CSDC2.SK-N-SH_all-trans-retinoic-acid_6-uM_48-hour 274 bp overlap
CTBP2 2 datasets
ChIP H1 ENCFF329MAX 301 bp overlap
ChIP H1 ENCFF329MAX 204 bp overlap
CTCF 10 datasets
ChIP 22Rv1 ENCFF466OXN 74 bp overlap
ChIP H9 ENCFF152GTF 301 bp overlap
ChIP HeLa-S3 ENCSR000AOA.CTCF.HeLa-S3 174 bp overlap
ChIP RWPE2 ENCFF911IEE 190 bp overlap
ChIP SU-DHL-6 ENCFF116KKR 67 bp overlap
ChIP cardiac muscle cell ENCFF777TNC 95 bp overlap
ChIP dorsolateral prefrontal cortex ENCFF483ZLP 80 bp overlap
ChIP dorsolateral prefrontal cortex ENCFF835ZSJ 62 bp overlap
ChIP epithelial cell of prostate ENCFF086GTI 154 bp overlap
ChIP hESC_D5 GSE116862.CTCF.hESC_D5 196 bp overlap
CTNNB1 1 dataset
ChIP LS180_125 GSE31939.CTNNB1.LS180_125 141 bp overlap
Cebpa 1 dataset
ChIP BLaER1 ENCFF364PUR 242 bp overlap
E2F8 2 datasets
ChIP HepG2 ENCFF117UYU 301 bp overlap
ChIP HepG2 ENCFF117UYU 142 bp overlap
EGR1 1 dataset
ChIP Hep-G2 ENCSR026GSW.EGR1.Hep-G2 137 bp overlap
ELL2 1 dataset
ChIP HeLa GSE40632.ELL2.HeLa 140 bp overlap
EOMES 1 dataset
ChIP hESC GSE26097.EOMES.hESC 198 bp overlap
EP300 1 dataset
ChIP Hep-G2 ENCSR000BLW.EP300.Hep-G2 209 bp overlap
ESR1 1 dataset
ChIP Ishikawa_siETV4-72h GSE129803.ESR1.Ishikawa_siETV4-72h 152 bp overlap
ESRRG 1 dataset
ChIP cardiomyocyte GSE113760.ESRRG.cardiomyocyte 301 bp overlap
EZH2 1 dataset
ChIP RH30_DMSO GSE85169.EZH2.RH30_DMSO 262 bp overlap
FEZF1 1 dataset
ChIP HEK293 GSE76494.FEZF1.HEK293 156 bp overlap
FOXA1 1 dataset
ChIP breast-cancer_Veh-2856 GSE128018.FOXA1.breast-cancer_Veh-2856 218 bp overlap
FOXA2 2 datasets
ChIP HepG2 ENCFF894AYY 286 bp overlap
ChIP HepG2 ENCFF894AYY 184 bp overlap
FOXK1 1 dataset
ChIP Hep-G2 ENCSR603BJQ.FOXK1.Hep-G2 146 bp overlap
FOXO1 1 dataset
ChIP HepG2 ENCFF088FIR 301 bp overlap
FOXP1 2 datasets
ChIP Hep-G2 ENCSR369YUK.FOXP1.Hep-G2 252 bp overlap
ChIP HepG2 ENCFF823ERM 301 bp overlap
FOXP4 3 datasets
ChIP Hep-G2 ENCSR232LLP.FOXP4.Hep-G2 231 bp overlap
ChIP HepG2 ENCFF462ULY 301 bp overlap
ChIP HepG2 ENCFF462ULY 129 bp overlap
GATA2 1 dataset
ChIP hiPSC_SLC6ebs GSE107639.GATA2.hiPSC_SLC6ebs 183 bp overlap
GATA4 2 datasets
ChIP DE DE-GATA4-2 296 bp overlap
ChIP cardiomyocyte_7 GSE85628.GATA4.cardiomyocyte_7 235 bp overlap
GATA6 2 datasets
ChIP DE_D2 S55-DE-d2-GATA6-exp2 201 bp overlap
ChIP PATU8988 GSE47535.GATA6.PATU8988 203 bp overlap
GATAD2A 1 dataset
ChIP HepG2 ENCFF252XNH 301 bp overlap
GLIS1 2 datasets
ChIP HEK293 ENCFF299RSE 301 bp overlap
ChIP HEK293 ENCSR482BBZ.GLIS1.HEK293 268 bp overlap
HAND2 2 datasets
ChIP CLB-Ga GSE90683.HAND2.CLB-Ga 301 bp overlap
ChIP SK-N-BE2-C GSE94822.HAND2.SK-N-BE2-C 284 bp overlap
HBP1 1 dataset
ChIP Hep-G2 ENCSR616WEG.HBP1.Hep-G2 102 bp overlap
HDAC1 1 dataset
ChIP Hep-G2 ENCSR750OWO.HDAC1.Hep-G2 179 bp overlap
HDAC2 3 datasets
ChIP Hep-G2 ENCSR000BMC.HDAC2.Hep-G2 197 bp overlap
ChIP RH4_DMSO-6H GSE116344.HDAC2.RH4_DMSO-6H 248 bp overlap
ChIP RH4_Entinostat-6H GSE116344.HDAC2.RH4_Entinostat-6H 301 bp overlap
HNF4A 2 datasets
ChIP Hep-G2 ENCSR469FBY.HNF4A.Hep-G2 200 bp overlap
ChIP HepG2 ENCFF146SSF 299 bp overlap
HNRNPH1 2 datasets
ChIP Hep-G2 ENCSR066FXN.HNRNPH1.Hep-G2 206 bp overlap
ChIP Hep-G2 GSE120104.HNRNPH1.Hep-G2 206 bp overlap
HOXA3 2 datasets
ChIP Hep-G2 ENCSR999JKC.HOXA3.Hep-G2 254 bp overlap
ChIP HepG2 ENCFF374TCI 301 bp overlap
IKZF3 1 dataset
ChIP HEK293 ENCSR304AMN.IKZF3.HEK293 301 bp overlap
KDM1A 1 dataset
ChIP K-562 GSE117944.KDM1A.K-562 52 bp overlap
KDM3A 1 dataset
ChIP Hep-G2 ENCSR387JKT.KDM3A.Hep-G2 202 bp overlap
KLF1 2 datasets
ChIP HEK293 ENCFF159QSW 154 bp overlap
ChIP HEK293 ENCSR859BMR.KLF1.HEK293 78 bp overlap
KLF6 1 dataset
ChIP Hep-G2 ENCSR757EKM.KLF6.Hep-G2 195 bp overlap
KLF8 2 datasets
ChIP HEK293 ENCFF929IAJ 301 bp overlap
ChIP HEK293 ENCFF929IAJ 139 bp overlap
KMT2A 3 datasets
ChIP HEK293T_N-term_C49 GSE90762.KMT2A.HEK293T_N-term_C49 221 bp overlap
ChIP HEK293T_N-term_shCtrl_TASP1-KO GSE90762.KMT2A.HEK293T_N-term_shCtrl_TASP1-KO 301 bp overlap
ChIP Hep-G2 ENCSR692GFR.KMT2A.Hep-G2 301 bp overlap
KMT2C 1 dataset
ChIP BIN-67_lentivirus-SMARCA4 GSE117734.KMT2C.BIN-67_lentivirus-SMARCA4 193 bp overlap
LCOR 1 dataset
ChIP HepG2 ENCFF499KCU 301 bp overlap
MAX 2 datasets
ChIP Hep-G2 ENCSR168DYA.MAX.Hep-G2 244 bp overlap
ChIP NCI-H2171 GSE36354.MAX.NCI-H2171 207 bp overlap
MAZ 1 dataset
ChIP HepG2 ENCFF068NYH 208 bp overlap
MED1 5 datasets
ChIP G296S GSE85628.MED1.G296S 301 bp overlap
ChIP G296S_2 GSE85628.MED1.G296S_2 301 bp overlap
ChIP RH4 GSE83726.MED1.RH4 235 bp overlap
ChIP cardiomyocyte GSE85628.MED1.cardiomyocyte 161 bp overlap
ChIP cardiomyocyte_1 GSE85628.MED1.cardiomyocyte_1 162 bp overlap
MTA1 1 dataset
ChIP Hep-G2 ENCSR983KRB.MTA1.Hep-G2 236 bp overlap
MTERF4 1 dataset
ChIP HepG2 ENCFF831NAM 301 bp overlap
MXD4 1 dataset
ChIP Hep-G2 ENCSR441KFW.MXD4.Hep-G2 174 bp overlap
MYBL2 1 dataset
ChIP Hep-G2 ENCSR581KCO.MYBL2.Hep-G2 189 bp overlap
MYCN 2 datasets
ChIP CHP-134 GSE129588.MYCN.CHP-134 173 bp overlap
ChIP RH4 GSE83726.MYCN.RH4 263 bp overlap
MYNN 1 dataset
ChIP Hep-G2 ENCSR398TMP.MYNN.Hep-G2 166 bp overlap
MYOD1 2 datasets
ChIP RH30_DMSO GSE85169.MYOD1.RH30_DMSO 195 bp overlap
ChIP RH4 GSE83726.MYOD1.RH4 301 bp overlap
MYPOP 2 datasets
ChIP HepG2 ENCFF176TQL 301 bp overlap
ChIP HepG2 ENCFF176TQL 301 bp overlap
NANOG 1 dataset
ChIP WA09 GSE105028.NANOG.WA09 215 bp overlap
NCOA2 1 dataset
ChIP HepG2 ENCFF853BJJ 301 bp overlap
NEUROG2 1 dataset
ChIP MRC-5_NFD_1DPT GSE75910.NEUROG2.MRC-5_NFD_1DPT 168 bp overlap
NFIA 2 datasets
ChIP Hep-G2 GSE97661.NFIA.Hep-G2 177 bp overlap
ChIP HepG2 ENCFF815HWK 249 bp overlap
NFIC 1 dataset
ChIP Hep-G2 GSE108514.NFIC.Hep-G2 275 bp overlap
NFYC 1 dataset
ChIP Hep-G2 ENCSR569ARC.NFYC.Hep-G2 209 bp overlap
NKX2-5 1 dataset
ChIP hESC_sc-14033 GSE89457.NKX2-5.hESC_sc-14033 135 bp overlap
NR2F6 1 dataset
ChIP HepG2 ENCFF514UJI 301 bp overlap
ONECUT1 4 datasets
ChIP Hep-G2 ENCSR956OSX.ONECUT1.Hep-G2 293 bp overlap
ChIP HepG2 ENCFF243FIR 301 bp overlap
ChIP liver ERP002306.ONECUT1.liver 189 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.ONECUT1.pancreatic-progenitor_PP1 301 bp overlap
PATZ1 1 dataset
ChIP HepG2 ENCFF723PFC 301 bp overlap
PAXIP1 3 datasets
ChIP Hep-G2 ENCSR530ARJ.PAXIP1.Hep-G2 268 bp overlap
ChIP HepG2 ENCFF526NOJ 301 bp overlap
ChIP HepG2 ENCFF526NOJ 213 bp overlap
PDX1 1 dataset
ChIP hiPSC_derived_PPs GSE106949.PDX1.hiPSC_derived_PPs 172 bp overlap
PHF5A 1 dataset
ChIP Hep-G2 ENCSR191TLD.PHF5A.Hep-G2 132 bp overlap
PITX1 1 dataset
ChIP HepG2 ENCFF468QTQ 301 bp overlap
PPARG 3 datasets
ChIP HT29_ROSIG_2H GSE77039.PPARG.HT29_ROSIG_2H 185 bp overlap
ChIP HT29_ROSIG_48H GSE77039.PPARG.HT29_ROSIG_48H 213 bp overlap
ChIP Hep-G2 ENCSR130VQL.PPARG.Hep-G2 237 bp overlap
PRDM1 1 dataset
ChIP HEK293 ENCFF302TBP 210 bp overlap
PRDM10 2 datasets
ChIP HEK293 ENCFF145WQQ 301 bp overlap
ChIP Hep-G2 ENCSR289PSX.PRDM10.Hep-G2 301 bp overlap
RAD21 3 datasets
ChIP HEK293_siE1A GSE130135.RAD21.HEK293_siE1A 76 bp overlap
ChIP Hep-G2 ENCSR054FKH.RAD21.Hep-G2 100 bp overlap
ChIP RH4 GSE83726.RAD21.RH4 273 bp overlap
RARA 1 dataset
ChIP HepG2 ENCFF582XUA 301 bp overlap
RBBP4 2 datasets
ChIP RH5 GSE155861.RBBP4.RH5 145 bp overlap
ChIP SCMC GSE155861.RBBP4.SCMC 146 bp overlap
RBBP5 1 dataset
ChIP WA01 ENCSR000AQC.RBBP5.WA01 250 bp overlap
RREB1 1 dataset
ChIP HepG2 ENCFF986CSN 271 bp overlap
SALL1 2 datasets
ChIP HepG2 ENCFF426MCK 301 bp overlap
ChIP HepG2 ENCFF426MCK 102 bp overlap
SAP130 2 datasets
ChIP HepG2 ENCFF892EHZ 301 bp overlap
ChIP HepG2 ENCFF892EHZ 245 bp overlap
SMAD2-3 1 dataset
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 257 bp overlap
SMAD3 1 dataset
ChIP Hep-G2 ENCSR005GZH.SMAD3.Hep-G2 172 bp overlap
SMARCA4 4 datasets
ChIP BIN-67_lentivirus-SMARCA4-T910M GSE117734.SMARCA4.BIN-67_lentivirus-SMARCA4-T910M 223 bp overlap
ChIP NGP_ARID1A-het GSE134626.SMARCA4.NGP_ARID1A-het 85 bp overlap
ChIP NPC_siCT GSE122631.SMARCA4.NPC_siCT 257 bp overlap
ChIP NSC GSE125033.SMARCA4.NSC 301 bp overlap
SMARCC1 2 datasets
ChIP BIN-67_lentivirus-SMARCA4-T910M GSE117734.SMARCC1.BIN-67_lentivirus-SMARCA4-T910M 175 bp overlap
ChIP HS-SY-2 GSE108025.SMARCC1.HS-SY-2 301 bp overlap
SMC3 1 dataset
ChIP neural ENCSR404BPV.SMC3.neural 224 bp overlap
SNAI2 1 dataset
ChIP SK-N-SH_all-trans-retinoic-acid_6-uM_48-hour ENCSR106GVM.SNAI2.SK-N-SH_all-trans-retinoic-acid_6-uM_48-hour 207 bp overlap
SOX13 1 dataset
ChIP HepG2 ENCFF062VSQ 248 bp overlap
SOX17 1 dataset
ChIP DE_D2 DED2-SOX17_Batch_II 301 bp overlap
SOX17_M 1 dataset
ChIP DE_D2 DED2-SOX17-M_Batch_II 273 bp overlap
SOX2 1 dataset
ChIP HNSC GSE69479.SOX2.HNSC 278 bp overlap
SOX5 1 dataset
ChIP HepG2 ENCFF470KZD 214 bp overlap
SOX6 2 datasets
ChIP Hep-G2 ENCSR766TSU.SOX6.Hep-G2 301 bp overlap
ChIP HepG2 ENCFF767OCK 301 bp overlap
SP1 1 dataset
ChIP WA01 ENCSR000BIR.SP1.WA01 161 bp overlap
SP3 1 dataset
ChIP HEK293 ENCSR141PZA.SP3.HEK293 155 bp overlap
SP5 1 dataset
ChIP Hep-G2 ENCSR019NPF.SP5.Hep-G2 119 bp overlap
SS18 1 dataset
ChIP BIN-67_lentivirus-SMARCA4-T910M GSE117734.SS18.BIN-67_lentivirus-SMARCA4-T910M 250 bp overlap
T 1 dataset
ChIP H9_MESODERM GSE60606.T.H9_MESODERM 199 bp overlap
TAF1 2 datasets
ChIP H1 ENCFF478SZO 260 bp overlap
ChIP WA01 ENCSR000BHO.TAF1.WA01 149 bp overlap
TBX2 2 datasets
ChIP Hep-G2 ENCSR410FEH.TBX2.Hep-G2 301 bp overlap
ChIP HepG2 ENCFF811TLA 301 bp overlap
TBX3 1 dataset
ChIP HepG2 ENCFF178RIL 301 bp overlap
TBX5 6 datasets
ChIP G296S GSE85628.TBX5.G296S 285 bp overlap
ChIP G296S_2 GSE85628.TBX5.G296S_2 285 bp overlap
ChIP G296S_4 GSE85628.TBX5.G296S_4 301 bp overlap
ChIP cardiomyocyte_5 GSE85628.TBX5.cardiomyocyte_5 282 bp overlap
ChIP cardiomyocyte_7 GSE85628.TBX5.cardiomyocyte_7 301 bp overlap
ChIP hiPSC GSE81585.TBX5.hiPSC 290 bp overlap
TCF12 2 datasets
ChIP H1 ENCFF203EBH 251 bp overlap
ChIP WA01 ENCSR000BIT.TCF12.WA01 194 bp overlap
TCF7L2 2 datasets
ChIP Hep-G2 ENCSR346BOD.TCF7L2.Hep-G2 167 bp overlap
ChIP HepG2 ENCFF510OLG 301 bp overlap
TEAD1 3 datasets
ChIP HepG2 ENCFF661PNM 167 bp overlap
ChIP MSTO GSE68170.TEAD1.MSTO 213 bp overlap
ChIP pancreas_12D ERP008682.TEAD1.pancreas_12D 200 bp overlap
TEAD3 1 dataset
ChIP HepG2 ENCFF054UUL 291 bp overlap
TEAD4 9 datasets
ChIP BJ_fibroblast_CD13-neg GSE114367.TEAD4.BJ_fibroblast_CD13-neg 219 bp overlap
ChIP H1 ENCFF778PAX 101 bp overlap
ChIP HEK293_siCtrl GSE130135.TEAD4.HEK293_siCtrl 243 bp overlap
ChIP HEK293_siE1A GSE130135.TEAD4.HEK293_siE1A 223 bp overlap
ChIP Hep-G2 ENCSR000BRP.TEAD4.Hep-G2 262 bp overlap
ChIP HepG2 ENCFF006QNB 301 bp overlap
ChIP HepG2 ENCFF250NXO 172 bp overlap
ChIP Ishikawa ENCSR000BSW.TEAD4.Ishikawa 110 bp overlap
ChIP hMSC-TERT4 GSE140782.TEAD4.hMSC-TERT4 212 bp overlap
TFAP4 2 datasets
ChIP Hep-G2 ENCSR103SZL.TFAP4.Hep-G2 163 bp overlap
ChIP HepG2 ENCFF932XOY 298 bp overlap
TFE3 1 dataset
ChIP Hep-G2 ENCSR589SNT.TFE3.Hep-G2 163 bp overlap
THRB 1 dataset
ChIP Hep-G2 ENCSR430JGJ.THRB.Hep-G2 144 bp overlap
TWIST1 2 datasets
ChIP SHEP-21N GSE80151.TWIST1.SHEP-21N 181 bp overlap
ChIP SHEP-21N_DOX_0H GSE80151.TWIST1.SHEP-21N_DOX_0H 181 bp overlap
YAP1 1 dataset
ChIP hiPSC GSE111930.YAP1.hiPSC 127 bp overlap
YY1 5 datasets
ChIP H1 ENCFF524BTL 262 bp overlap
ChIP Hep-G2_NC GSE120104.YY1.Hep-G2_NC 301 bp overlap
ChIP Hep-G2_RBM25-KD GSE120104.YY1.Hep-G2_RBM25-KD 269 bp overlap
ChIP WA01 ENCSR000BKD.YY1.WA01 131 bp overlap
ChIP liver ENCSR994YLZ.YY1.liver 212 bp overlap
ZBTB7A 1 dataset
ChIP VCaP_FBS GSE123089.ZBTB7A.VCaP_FBS 108 bp overlap
ZEB1 1 dataset
ChIP neuron_bipolar_doxy_4d ENCSR418KUS.ZEB1.neuron_bipolar_doxy_4d 218 bp overlap
ZFHX3 1 dataset
ChIP HepG2 ENCFF082SJV 301 bp overlap
ZFX 1 dataset
ChIP HEK293T ENCFF402JZW 208 bp overlap
ZGPAT 1 dataset
ChIP Hep-G2 ENCSR704IGU.ZGPAT.Hep-G2 301 bp overlap
ZMYM4 1 dataset
ChIP HepG2 ENCFF567SQY 128 bp overlap
ZNF217 2 datasets
ChIP Hep-G2 ENCSR055FQB.ZNF217.Hep-G2 301 bp overlap
ChIP HepG2 ENCFF455XGO 301 bp overlap
ZNF292 1 dataset
ChIP HepG2 ENCFF975MAJ 301 bp overlap
ZNF331 1 dataset
ChIP HepG2 ENCFF842SZN 291 bp overlap
ZNF44 1 dataset
ChIP HepG2 ENCFF984YCN 55 bp overlap
ZNF600 1 dataset
ChIP HEK293 ENCFF785JSX 301 bp overlap
ZNF644 1 dataset
ChIP HepG2 ENCFF352VGJ 297 bp overlap
ZNF711 1 dataset
ChIP HEK293T GSE145160.ZNF711.HEK293T 267 bp overlap
ZNF770 1 dataset
ChIP HEK293 GSE76494.ZNF770.HEK293 204 bp overlap
ZSCAN22 1 dataset
ChIP HEK293 GSE76494.ZSCAN22.HEK293 222 bp overlap