chr6 : 28,357,204 28,357,431
227 bp 129 TFs 2 linked genes
This 227 bp open chromatin element is linked to ZSCAN31 and ZKSCAN3 and is bound by 129 transcription factors.
Linked Genes
2 genes
Distance
Gene Expression Dist. to TSS Distance Link type
ZSCAN31 933 bp At TSS Proximity
ZKSCAN3 7.3 kb Proximal Proximity
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr6:28,352,204 – 28,362,431
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
129 transcription factors
Source
Cell type
ARID1B 2 datasets
ChIP K-562 ENCSR822CCM.ARID1B.K-562 206 bp overlap
ChIP K562 ENCFF938UXQ 227 bp overlap
ARNT 1 dataset
ChIP K-562 ENCSR155KHM.ARNT.K-562 227 bp overlap
ARNTL 3 datasets
ChIP U2OS GSE130602.ARNTL.U2OS 147 bp overlap
ChIP U2OS_DMSO GSE130506.ARNTL.U2OS_DMSO 147 bp overlap
ChIP U2OS_cordycepin GSE130506.ARNTL.U2OS_cordycepin 123 bp overlap
ATF1 1 dataset
ChIP K562 ENCFF469GPI 227 bp overlap
ATF3 2 datasets
ChIP K-562 ENCSR028UIU.ATF3.K-562 190 bp overlap
ChIP K562 ENCFF604FPV 227 bp overlap
ATF7 1 dataset
ChIP K562 ENCFF308SKS 227 bp overlap
BACH1 6 datasets
ChIP AsPC-1 GSE124406.BACH1.AsPC-1 183 bp overlap
ChIP K-562 ENCSR740NPG.BACH1.K-562 227 bp overlap
ChIP K562 ENCFF419VIM 227 bp overlap
ChIP K562 ENCFF419VIM 227 bp overlap
ChIP K562 ENCFF990JHI 100 bp overlap
ChIP SW1990 GSE124406.BACH1.SW1990 185 bp overlap
BAF155 1 dataset
ChIP VCaP_shARID1A GSE110655.BAF155.VCaP_shARID1A 193 bp overlap
BCOR 2 datasets
ChIP K-562 ENCSR808AKZ.BCOR.K-562 69 bp overlap
ChIP K562 ENCFF343XWA 190 bp overlap
BHLHE40 2 datasets
ChIP IMR-90 ENCFF312JYK 107 bp overlap
ChIP IMR-90 ENCSR957KYB.BHLHE40.IMR-90 59 bp overlap
BRD2 6 datasets
ChIP MDA-MB-231_BAZ2AsiRNA GSE116879.BRD2.MDA-MB-231_BAZ2AsiRNA 227 bp overlap
ChIP MDA-MB-231_BAZ2BsiRNA GSE116879.BRD2.MDA-MB-231_BAZ2BsiRNA 227 bp overlap
ChIP MDA-MB-231_MGSK2801 GSE116879.BRD2.MDA-MB-231_MGSK2801 227 bp overlap
ChIP MDA-MB-231_MGSK2801_rDNA GSE116879.BRD2.MDA-MB-231_MGSK2801_rDNA 227 bp overlap
ChIP MDA-MB-231_NTsiRNA GSE116879.BRD2.MDA-MB-231_NTsiRNA 227 bp overlap
ChIP SK-MEL-147 GSE94488.BRD2.SK-MEL-147 227 bp overlap
BRD4 10 datasets
ChIP IMR-90_QUIES GSE74238.BRD4.IMR-90_QUIES 119 bp overlap
ChIP MDA-MB-231_DMSO GSE116879.BRD4.MDA-MB-231_DMSO 86 bp overlap
ChIP MDA-MB-231_DMSO_rDNA GSE116879.BRD4.MDA-MB-231_DMSO_rDNA 86 bp overlap
ChIP MDA-MB-231_JQ1-neg_L GSE136151.BRD4.MDA-MB-231_JQ1-neg_L 196 bp overlap
ChIP MDA-MB-231_JQ1-neg_S_3f GSE136151.BRD4.MDA-MB-231_JQ1-neg_S_3f 227 bp overlap
ChIP MDA-MB-231_MGSK2801 GSE116879.BRD4.MDA-MB-231_MGSK2801 206 bp overlap
ChIP MDA-MB-231_MGSK2801_rDNA GSE116879.BRD4.MDA-MB-231_MGSK2801_rDNA 206 bp overlap
ChIP SUM159PT_30nMbortezomib_8h GSE87418.BRD4.SUM159PT_30nMbortezomib_8h 142 bp overlap
ChIP SUM159PT_shMYC_DMSO_48h GSE87418.BRD4.SUM159PT_shMYC_DMSO_48h 226 bp overlap
ChIP SUM159PT_shMYC_dox_48h GSE87418.BRD4.SUM159PT_shMYC_dox_48h 196 bp overlap
BRD9 2 datasets
ChIP K562 ENCFF480JXZ 142 bp overlap
ChIP K562 ENCFF480JXZ 227 bp overlap
CAVIN1 1 dataset
ChIP K562 ENCFF731YBD 223 bp overlap
CBX3 1 dataset
ChIP K562 ENCFF410AQU 91 bp overlap
CBX5 2 datasets
ChIP K-562 ENCSR272JAT.CBX5.K-562 55 bp overlap
ChIP K562 ENCFF188CYP 115 bp overlap
CEBPZ 2 datasets
ChIP K-562 ENCSR618GDK.CEBPZ.K-562 77 bp overlap
ChIP K562 ENCFF909BYC 136 bp overlap
CREB3 3 datasets
ChIP K-562 ENCSR093FKD.CREB3.K-562 227 bp overlap
ChIP K562 ENCFF985QJI 227 bp overlap
ChIP K562 ENCFF985QJI 196 bp overlap
CREBBP 1 dataset
ChIP fibroblast_senescent GSE106146.CREBBP.fibroblast_senescent 101 bp overlap
CREM 1 dataset
ChIP K562 ENCFF180STA 96 bp overlap
CTCF 1 dataset
ChIP HCT-116 GSE131606.CTCF.HCT-116 178 bp overlap
CUX1 1 dataset
ChIP K562 ENCFF902MYN 227 bp overlap
CXXC5 1 dataset
ChIP K562 ENCFF497CZN 227 bp overlap
DMBX1 1 dataset
ChIP K562 ENCFF972HXB 200 bp overlap
DPF2 1 dataset
ChIP K562 ENCFF775HUO 69 bp overlap
E2F1 1 dataset
ChIP MDA-MB-231 GSE95303.E2F1.MDA-MB-231 227 bp overlap
E4F1 1 dataset
ChIP K-562 ENCSR731LHZ.E4F1.K-562 212 bp overlap
EHMT2 1 dataset
ChIP K-562 ENCSR175EOM.EHMT2.K-562 186 bp overlap
ELF1 1 dataset
ChIP K-562 ENCSR000BMD.ELF1.K-562 57 bp overlap
EMSY 2 datasets
ChIP K562 ENCFF511ZZZ 164 bp overlap
ChIP K562 ENCFF511ZZZ 227 bp overlap
EP300 3 datasets
ChIP fibroblast_senescent GSE106146.EP300.fibroblast_senescent 218 bp overlap
ChIP pulmonary-artery_endothelial-cell_siCtrl GSE89786.EP300.pulmonary-artery_endothelial-cell_siCtrl 173 bp overlap
ChIP pulmonary-artery_endothelial-cell_siPFKFB3 GSE89786.EP300.pulmonary-artery_endothelial-cell_siPFKFB3 144 bp overlap
ERG 2 datasets
ChIP HUVEC-C GSE124891.ERG.HUVEC-C 127 bp overlap
ChIP VCaP_shARID1A GSE110655.ERG.VCaP_shARID1A 203 bp overlap
ETS1 7 datasets
ChIP HUVEC-C GSE41166.ETS1.HUVEC-C 120 bp overlap
ChIP HUVEC-C_12h GSE93030.ETS1.HUVEC-C_12h 151 bp overlap
ChIP HUVEC-C_4h GSE93030.ETS1.HUVEC-C_4h 82 bp overlap
ChIP HUVEC-C_VEGF_12H GSE41166.ETS1.HUVEC-C_VEGF_12H 185 bp overlap
ChIP HUVEC-C_VEGF_12h GSE109625.ETS1.HUVEC-C_VEGF_12h 151 bp overlap
ChIP HUVEC-C_VEGF_4H GSE41166.ETS1.HUVEC-C_VEGF_4H 102 bp overlap
ChIP HUVEC-C_VEGF_4h GSE109625.ETS1.HUVEC-C_VEGF_4h 82 bp overlap
ETV1 2 datasets
ChIP K-562 ENCSR277DMR.ETV1.K-562 138 bp overlap
ChIP K562 ENCFF389WTI 176 bp overlap
FLI1 1 dataset
ChIP HUVEC-C GSE109695.FLI1.HUVEC-C 147 bp overlap
FOS 2 datasets
ChIP IMR-90 ENCFF179EDA 137 bp overlap
ChIP IMR-90 ENCSR124AIG.FOS.IMR-90 130 bp overlap
FOSL1 1 dataset
ChIP MDA-MB-231 GSE95303.FOSL1.MDA-MB-231 137 bp overlap
FOXA2 2 datasets
ChIP CFPAC-1 GSE119930.FOXA2.CFPAC-1 160 bp overlap
ChIP DE DE-FOXA2-1 210 bp overlap
FOXA3 2 datasets
ChIP K562 ENCFF348SOM 227 bp overlap
ChIP K562 ENCFF781VSC 182 bp overlap
FOXK2 4 datasets
ChIP K-562 ENCSR508DQA.FOXK2.K-562 109 bp overlap
ChIP K-562 ENCSR302AWT.FOXK2.K-562 227 bp overlap
ChIP K562 ENCFF245WKP 219 bp overlap
ChIP K562 ENCFF851PFH 219 bp overlap
GABPA 1 dataset
ChIP K562 ENCFF139LXS 227 bp overlap
GABPB1 2 datasets
ChIP K-562 ENCSR138YYY.GABPB1.K-562 227 bp overlap
ChIP K562 ENCFF015GDS 227 bp overlap
GATA4 2 datasets
ChIP DE DE-GATA4-1 227 bp overlap
ChIP DE DE-GATA4-2 227 bp overlap
GATA6 4 datasets
ChIP DE DE-GATA6-1 227 bp overlap
ChIP DE DE-GATA6-2 227 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 227 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 227 bp overlap
HDAC2 2 datasets
ChIP K-562 ENCSR893WSB.HDAC2.K-562 227 bp overlap
ChIP K-562 ENCSR075HTM.HDAC2.K-562 226 bp overlap
HDGF 1 dataset
ChIP K-562 ENCSR197ALX.HDGF.K-562 227 bp overlap
HIF1A 2 datasets
ChIP HUVEC-C_HYPOX GSE39089.HIF1A.HUVEC-C_HYPOX 227 bp overlap
ChIP U2OS_DMOG GSE85096.HIF1A.U2OS_DMOG 84 bp overlap
HINFP 1 dataset
ChIP K562 ENCFF361QXJ 202 bp overlap
HLF 1 dataset
ChIP HepG2 ENCFF854JLR 60 bp overlap
IRF1 1 dataset
ChIP K-562 ENCSR854MCV.IRF1.K-562 227 bp overlap
JUN 3 datasets
ChIP K562 ENCFF372VWH 151 bp overlap
ChIP MDA-BoM-1833 GSE112444.JUN.MDA-BoM-1833 94 bp overlap
ChIP MDA-MB-231 GSE112444.JUN.MDA-MB-231 79 bp overlap
JUND 3 datasets
ChIP K-562 ENCSR000EGN.JUND.K-562 61 bp overlap
ChIP K562 ENCFF336RCR 209 bp overlap
ChIP K562 ENCFF830LVJ 112 bp overlap
KDM1A 1 dataset
ChIP K562 ENCFF128TYE 227 bp overlap
KLF13 1 dataset
ChIP K562 ENCFF738YZC 209 bp overlap
KLF6 2 datasets
ChIP K-562 ENCSR297CGF.KLF6.K-562 195 bp overlap
ChIP K562 ENCFF948QSP 227 bp overlap
L3MBTL2 2 datasets
ChIP K-562 ENCSR530XQI.L3MBTL2.K-562 227 bp overlap
ChIP K562 ENCFF320EQC 227 bp overlap
LEF1 1 dataset
ChIP K562 ENCFF198WCP 227 bp overlap
MAFB 1 dataset
ChIP islet ERP004003.MAFB.islet 64 bp overlap
MAFF 2 datasets
ChIP K-562 ENCSR000EGI.MAFF.K-562 98 bp overlap
ChIP K562 ENCFF071YKK 125 bp overlap
MAFG 2 datasets
ChIP K-562 ENCSR818DQV.MAFG.K-562 227 bp overlap
ChIP K562 ENCFF455EEO 206 bp overlap
MAFK 4 datasets
ChIP H1 ENCFF854XWE 102 bp overlap
ChIP IMR-90 ENCFF336DHZ 138 bp overlap
ChIP K-562 ENCSR000EGX.MAFK.K-562 88 bp overlap
ChIP K562 ENCFF380WHM 137 bp overlap
MAX 1 dataset
ChIP K562 ENCFF110LJS 141 bp overlap
MED1 1 dataset
ChIP HUVEC-C_siERG GSE124892.MED1.HUVEC-C_siERG 184 bp overlap
MEF2D 1 dataset
ChIP K562 ENCFF392LDT 227 bp overlap
MGA 1 dataset
ChIP K562 ENCFF140CEX 227 bp overlap
MNT 1 dataset
ChIP K562 ENCFF820IGH 227 bp overlap
MTA3 1 dataset
ChIP K-562 ENCSR914NEI.MTA3.K-562 227 bp overlap
MYC 1 dataset
ChIP endothelial_umbilical-vein ENCSR000DLU.MYC.endothelial_umbilical-vein 53 bp overlap
NCOR1 1 dataset
ChIP K-562 ENCSR798ILC.NCOR1.K-562 200 bp overlap
NFE2 5 datasets
ChIP K-562 ENCSR552YGL.NFE2.K-562 185 bp overlap
ChIP K-562 ENCSR000FCC.NFE2.K-562 145 bp overlap
ChIP K562 ENCFF047YKA 180 bp overlap
ChIP K562 ENCFF163BSI 76 bp overlap
ChIP erythroid GSE125753.NFE2.erythroid 64 bp overlap
NFE2L1 2 datasets
ChIP K-562 ENCSR632SHZ.NFE2L1.K-562 185 bp overlap
ChIP K562 ENCFF138UHK 130 bp overlap
NFE2L2 7 datasets
ChIP A-549 ENCSR584GHV.NFE2L2.A-549 85 bp overlap
ChIP A549 ENCFF474YMB 118 bp overlap
ChIP Hep-G2 ENCSR488EES.NFE2L2.Hep-G2 68 bp overlap
ChIP HepG2 ENCFF178DRC 113 bp overlap
ChIP IMR-90 ENCFF059WEE 86 bp overlap
ChIP IMR-90 ENCSR197WGI.NFE2L2.IMR-90 204 bp overlap
ChIP K-562 ENCSR485GKE.NFE2L2.K-562 200 bp overlap
NFRKB 1 dataset
ChIP K-562 ENCSR657EOF.NFRKB.K-562 227 bp overlap
NR1H2 1 dataset
ChIP K562 ENCFF386VZB 172 bp overlap
NR2C1 2 datasets
ChIP K-562 ENCSR178DEG.NR2C1.K-562 168 bp overlap
ChIP K562 ENCFF568JLK 219 bp overlap
NR4A1 2 datasets
ChIP K-562 ENCSR130PDE.NR4A1.K-562 227 bp overlap
ChIP K562 ENCFF998LHF 227 bp overlap
POLR2A 3 datasets
ChIP Panc1 ENCFF290KAB 227 bp overlap
ChIP endothelial cell of umbilical vein ENCFF467WJF 227 bp overlap
ChIP stomach ENCFF820WZN 174 bp overlap
PTTG1 1 dataset
ChIP K562 ENCFF017KRM 158 bp overlap
PYGO2 1 dataset
ChIP K562 ENCFF458IIA 149 bp overlap
RELA 1 dataset
ChIP aortic-endothelial-cell_IL1B_D8 GSE139377.RELA.aortic-endothelial-cell_IL1B_D8 70 bp overlap
RFX5 2 datasets
ChIP IMR-90 ENCFF886KPO 96 bp overlap
ChIP IMR-90 ENCSR000EFD.RFX5.IMR-90 58 bp overlap
RNF2 1 dataset
ChIP K562 ENCFF653BQJ 227 bp overlap
RUVBL2 1 dataset
ChIP U2OS GSE130602.RUVBL2.U2OS 227 bp overlap
SETDB1 2 datasets
ChIP K-562 ENCSR000EWI.SETDB1.K-562 117 bp overlap
ChIP K562 ENCFF745PAW 174 bp overlap
SMAD2-3 1 dataset
ChIP HUVEC-C_TGFB1 GSE134556.SMAD2-3.HUVEC-C_TGFB1 207 bp overlap
SMAD3 1 dataset
ChIP K-562 ENCSR376XAV.SMAD3.K-562 125 bp overlap
SMAD4 1 dataset
ChIP K562 ENCFF628RBP 227 bp overlap
SMARCE1 1 dataset
ChIP K562 ENCFF690CFF 227 bp overlap
SOX17 1 dataset
ChIP DE_D2 DED2-SOX17_Batch_II 227 bp overlap
SOX17_M 1 dataset
ChIP DE_D2 DED2-SOX17-M_Batch_II 227 bp overlap
SREBF1 1 dataset
ChIP K562 ENCFF441TTT 227 bp overlap
SRF 2 datasets
ChIP K562 ENCFF766EOO 170 bp overlap
ChIP K562 ENCFF766EOO 227 bp overlap
STAT3 1 dataset
ChIP MDA-MB-231_EtOH GSE85579.STAT3.MDA-MB-231_EtOH 69 bp overlap
TEAD1 1 dataset
ChIP K-562 ENCSR591ASD.TEAD1.K-562 134 bp overlap
TEAD4 1 dataset
ChIP K562 ENCFF843TII 227 bp overlap
TFDP1 1 dataset
ChIP K562 ENCFF794ZXJ 227 bp overlap
TP53 2 datasets
ChIP IMR-90_NUT3A GSE58740.TP53.IMR-90_NUT3A 169 bp overlap
ChIP MDA-MB-231 GSE95303.TP53.MDA-MB-231 224 bp overlap
TRIM24 2 datasets
ChIP K-562 ENCSR907MZR.TRIM24.K-562 167 bp overlap
ChIP K562 ENCFF616RIL 198 bp overlap
TRIM28 6 datasets
ChIP K-562 ENCSR000EVY.TRIM28.K-562 227 bp overlap
ChIP K-562 ENCSR474CVP.TRIM28.K-562 198 bp overlap
ChIP K562 ENCFF172UPN 159 bp overlap
ChIP K562 ENCFF671NCI 227 bp overlap
ChIP K562 ENCFF671NCI 227 bp overlap
ChIP hESC GSE115387.TRIM28.hESC 147 bp overlap
YY1 2 datasets
ChIP K-562 ENCSR000BMH.YY1.K-562 104 bp overlap
ChIP K562 ENCFF660QRE 149 bp overlap
ZBTB12 1 dataset
ChIP K562 ENCFF933CVM 227 bp overlap
ZBTB40 1 dataset
ChIP K562 ENCFF337GJB 227 bp overlap
ZBTB9 1 dataset
ChIP K562 ENCFF233EFX 207 bp overlap
ZEB2 2 datasets
ChIP K562 ENCFF795CMH 227 bp overlap
ChIP K562 ENCFF975RXS 196 bp overlap
ZFP36 1 dataset
ChIP K562 ENCFF255RZG 105 bp overlap
ZFX 3 datasets
ChIP K562 ENCFF169LZT 227 bp overlap
ChIP K562 ENCFF169LZT 227 bp overlap
ChIP K562 ENCFF169LZT 158 bp overlap
ZMYND8 1 dataset
ChIP MDA-MB-231 GSE108833.ZMYND8.MDA-MB-231 68 bp overlap
ZNF134 1 dataset
ChIP K562 ENCFF502NWS 227 bp overlap
ZNF143 1 dataset
ChIP K562 ENCFF554TVF 227 bp overlap
ZNF215 1 dataset
ChIP K562 ENCFF317MOH 183 bp overlap
ZNF24 2 datasets
ChIP K-562 ENCSR695EQB.ZNF24.K-562 227 bp overlap
ChIP K562 ENCFF877JCX 227 bp overlap
ZNF280A 1 dataset
ChIP K562 ENCFF706EWX 104 bp overlap
ZNF281 1 dataset
ChIP K562 ENCFF594VNM 227 bp overlap
ZNF282 2 datasets
ChIP K562 ENCFF657WOV 94 bp overlap
ChIP K562 ENCFF657WOV 227 bp overlap
ZNF311 1 dataset
ChIP K562 ENCFF986QSP 183 bp overlap
ZNF316 1 dataset
ChIP K562 ENCFF838QCD 177 bp overlap
ZNF317 1 dataset
ChIP K562 ENCFF896LCF 171 bp overlap
ZNF395 1 dataset
ChIP K562 ENCFF464EIT 227 bp overlap
ZNF449 1 dataset
ChIP K562 ENCFF038VEZ 227 bp overlap
ZNF592 2 datasets
ChIP K-562 ENCSR249BHQ.ZNF592.K-562 227 bp overlap
ChIP K562 ENCFF547OSS 227 bp overlap
ZNF639 1 dataset
ChIP K562 ENCFF271FQR 227 bp overlap
ZNF740 1 dataset
ChIP K562 ENCFF447IXE 227 bp overlap
ZNF83 2 datasets
ChIP K-562 ENCSR257XVY.ZNF83.K-562 164 bp overlap
ChIP K562 ENCFF340RTV 227 bp overlap