chr10 : 26,524,443 26,524,835
392 bp 139 TFs 0 linked genes
This 392 bp open chromatin element has no linked target genes and is bound by 139 transcription factors.
Linked Genes
No linked genes
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr10:26,519,443 – 26,529,835
This element Other elements ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
139 transcription factors
Source
Cell type
ATF2 2 datasets
ChIP HEK293 ENCFF194VKZ 63 bp overlap
ChIP HEK293 ENCSR217HTK.ATF2.HEK293 372 bp overlap
BRCA1 2 datasets
ChIP HeLa-S3 ENCFF218GPC 141 bp overlap
ChIP HeLa-S3 ENCSR000EDB.BRCA1.HeLa-S3 78 bp overlap
CHD2 2 datasets
ChIP HeLa-S3 ENCFF078QRQ 188 bp overlap
ChIP HeLa-S3 ENCSR000ECP.CHD2.HeLa-S3 242 bp overlap
CHD4 1 dataset
ChIP HaCaT GSE139685.CHD4.HaCaT 177 bp overlap
EP300 2 datasets
ChIP HeLa-S3 ENCSR680OFU.EP300.HeLa-S3 392 bp overlap
ChIP HeLa-S3 ENCSR000ECV.EP300.HeLa-S3 392 bp overlap
FEZF1 3 datasets
ChIP HEK293 ENCFF528YED 379 bp overlap
ChIP HEK293 ENCSR827NWO.FEZF1.HEK293 392 bp overlap
ChIP HEK293 GSE76494.FEZF1.HEK293 240 bp overlap
FOS 1 dataset
ChIP HeLa-S3 ENCSR000EZE.FOS.HeLa-S3 113 bp overlap
FOXA1 2 datasets
ChIP 22Rv1_CST_Crispr-70 GSE123618.FOXA1.22Rv1_CST_Crispr-70 63 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.FOXA1.pancreatic-progenitor_PP1 241 bp overlap
FOXA2 2 datasets
ChIP DE DE-FOXA2-1 392 bp overlap
ChIP DE DE-FOXA2-2 392 bp overlap
FOXL2 4 datasets
ChIP HGrC1_C134W-TGF GSE138496.FOXL2.HGrC1_C134W-TGF 64 bp overlap
ChIP HGrC1_C134W-TGF GSE138496.FOXL2.HGrC1_C134W-TGF 128 bp overlap
ChIP HGrC1_C134W-TGF_parental GSE138496.FOXL2.HGrC1_C134W-TGF_parental 94 bp overlap
ChIP HGrC1_C134W-TGF_parental GSE138496.FOXL2.HGrC1_C134W-TGF_parental 202 bp overlap
GATA1 4 datasets
ChIP CD34_Day7_30min GSE104676.GATA1.CD34_Day7_30min 181 bp overlap
ChIP CD34_ERYTH_BMP GSE29194.GATA1.CD34_ERYTH_BMP 142 bp overlap
ChIP HUDEP-2_30min GSE104676.GATA1.HUDEP-2_30min 111 bp overlap
ChIP K-562_enCRISPRi-LK GSE132212.GATA1.K-562_enCRISPRi-LK 168 bp overlap
GATA2 3 datasets
ChIP dermal-fibroblast_alone GSE51025.GATA2.dermal-fibroblast_alone 130 bp overlap
ChIP primary-endometrial-stromal-cell_Veh_P1 GSE145673.GATA2.primary-endometrial-stromal-cell_Veh_P1 245 bp overlap
ChIP primary-endometrial-stromal-cell_Veh_P2 GSE145673.GATA2.primary-endometrial-stromal-cell_Veh_P2 211 bp overlap
GATA4 9 datasets
ChIP BJ1-hTERT_FOXA2_GATA4_Coexp GSE92491.GATA4.BJ1-hTERT_FOXA2_GATA4_Coexp 136 bp overlap
ChIP DE DE-GATA4-1 392 bp overlap
ChIP DE DE-GATA4-2 329 bp overlap
ChIP Hep-G2 GSE135714.GATA4.Hep-G2 65 bp overlap
ChIP Hep-G2 GSE135714.GATA4.Hep-G2 231 bp overlap
ChIP YCC-3 GSE51705.GATA4.YCC-3 226 bp overlap
ChIP foregut GSE117136.GATA4.foregut 329 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA4.pancreatic-progenitor 392 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA4.pancreatic-progenitor_PP1 390 bp overlap
GATA6 18 datasets
ChIP AGS GSE51705.GATA6.AGS 237 bp overlap
ChIP Caco-2_PROLIF GSE23436.GATA6.Caco-2_PROLIF 172 bp overlap
ChIP DE DE-GATA6-1 354 bp overlap
ChIP DE DE-GATA6-2 392 bp overlap
ChIP DE_D1 S09-DE-d1-GATA6-exp1 391 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 249 bp overlap
ChIP DE_D1 S41-DE-d1-GATA6-exp2 265 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 392 bp overlap
ChIP H9 ERP004206.GATA6.H9 241 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 256 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 392 bp overlap
ChIP YCC-3 GSE51705.GATA6.YCC-3 246 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 286 bp overlap
ChIP foregut GSE117136.GATA6.foregut 307 bp overlap
ChIP foregut_KO GSE117136.GATA6.foregut_KO 294 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA6.pancreatic-progenitor 373 bp overlap
ChIP pancreatic-progenitor_KO GSE117136.GATA6.pancreatic-progenitor_KO 392 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA6.pancreatic-progenitor_PP1 314 bp overlap
GFI1B 1 dataset
ChIP HEK293 ENCSR445PDR.GFI1B.HEK293 207 bp overlap
GLI2 1 dataset
ChIP HEK293 ENCSR978EQY.GLI2.HEK293 81 bp overlap
GLIS1 2 datasets
ChIP HEK293 ENCFF299RSE 244 bp overlap
ChIP HEK293 ENCSR482BBZ.GLIS1.HEK293 392 bp overlap
GLIS2 2 datasets
ChIP HEK293 ENCFF446EIF 135 bp overlap
ChIP HEK293 ENCSR535DIA.GLIS2.HEK293 139 bp overlap
HIC1 2 datasets
ChIP HEK293 ENCFF252CFL 269 bp overlap
ChIP HEK293 ENCSR803GYT.HIC1.HEK293 392 bp overlap
IKZF3 1 dataset
ChIP HEK293 ENCSR304AMN.IKZF3.HEK293 392 bp overlap
INSM2 1 dataset
ChIP HEK293 ENCSR382GSF.INSM2.HEK293 389 bp overlap
JUN 8 datasets
ChIP DE_D1 S08-DE-d1-JUN-exp1 392 bp overlap
ChIP DE_D1 S13-DE-d1-JUN-exp1 281 bp overlap
ChIP DE_D1 S40-DE-d1-JUN-exp2 392 bp overlap
ChIP DE_D2 S02-DE-d2-JUN-exp1 392 bp overlap
ChIP DE_D2 S54-DE-d2-JUN-exp2 232 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.JUN.HUES-8_DE_ctrl 392 bp overlap
ChIP HeLa-S3 ENCFF668QVP 275 bp overlap
ChIP HeLa-S3 ENCSR000EDG.JUN.HeLa-S3 165 bp overlap
JUND 1 dataset
ChIP HeLa-S3 ENCSR000EDH.JUND.HeLa-S3 317 bp overlap
KLF1 1 dataset
ChIP HEK293 ENCFF159QSW 117 bp overlap
KLF10 3 datasets
ChIP HEK293 ENCFF326EGX 279 bp overlap
ChIP HEK293 ENCFF326EGX 392 bp overlap
ChIP HEK293 ENCSR006GAQ.KLF10.HEK293 392 bp overlap
KLF13 1 dataset
ChIP HEK293 ENCSR946ZLI.KLF13.HEK293 153 bp overlap
KLF17 1 dataset
ChIP HEK293 ENCFF658MHR 143 bp overlap
KLF8 2 datasets
ChIP HEK293 ENCFF929IAJ 50 bp overlap
ChIP HEK293 ENCSR635NOQ.KLF8.HEK293 392 bp overlap
LDB1 1 dataset
ChIP HEP GSE52637.LDB1.HEP 188 bp overlap
MAZ 2 datasets
ChIP HEK293 ENCSR290SSQ.MAZ.HEK293 60 bp overlap
ChIP HEK293 ENCSR290SSQ.MAZ.HEK293 136 bp overlap
MZF1 3 datasets
ChIP HEK293 ENCFF683ZWN 122 bp overlap
ChIP HEK293 ENCFF683ZWN 383 bp overlap
ChIP HEK293 ENCSR298QUH.MZF1.HEK293 376 bp overlap
NFYB 1 dataset
ChIP HeLa-S3 ENCSR000DNR.NFYB.HeLa-S3 182 bp overlap
NKX2-1 1 dataset
ChIP NCI-H1819 GSE39998.NKX2-1.NCI-H1819 159 bp overlap
ONECUT1 2 datasets
ChIP H9 ERP004206.ONECUT1.H9 219 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.ONECUT1.pancreatic-progenitor_PP1 345 bp overlap
ONECUT2 2 datasets
ChIP AGS GSE113045.ONECUT2.AGS 169 bp overlap
ChIP AGS_Overexpression GSE113045.ONECUT2.AGS_Overexpression 364 bp overlap
OSR2 2 datasets
ChIP HEK293 ENCFF875BDB 77 bp overlap
ChIP HEK293 ENCSR324LTM.OSR2.HEK293 382 bp overlap
OVOL3 2 datasets
ChIP HEK293 ENCFF898STB 97 bp overlap
ChIP HEK293 ENCSR768LIO.OVOL3.HEK293 58 bp overlap
PATZ1 1 dataset
ChIP HEK293 ENCSR966ULI.PATZ1.HEK293 392 bp overlap
PDX1 1 dataset
ChIP hiPSC GSE125768.PDX1.hiPSC 234 bp overlap
PHIP 1 dataset
ChIP HEK293_ab833 GSE101646.PHIP.HEK293_ab833 392 bp overlap
PKNOX1 2 datasets
ChIP HEK293T ENCFF174WDB 240 bp overlap
ChIP HEK293T ENCSR233FAG.PKNOX1.HEK293T 345 bp overlap
PRDM1 1 dataset
ChIP HEK293 ENCFF302TBP 108 bp overlap
PRDM6 3 datasets
ChIP HEK293 ENCFF283AJL 129 bp overlap
ChIP HEK293 ENCSR892QHR.PRDM6.HEK293 384 bp overlap
ChIP HEK293 GSE76494.PRDM6.HEK293 63 bp overlap
REST 1 dataset
ChIP HEK293 ENCSR896UBV.REST.HEK293 313 bp overlap
RFX5 1 dataset
ChIP HeLa-S3 ENCSR000ECX.RFX5.HeLa-S3 84 bp overlap
RUNX1 1 dataset
ChIP Jurkat GSE68976.RUNX1.Jurkat 173 bp overlap
SCRT1 1 dataset
ChIP HEK293 ENCSR605MGM.SCRT1.HEK293 223 bp overlap
SIX2 1 dataset
ChIP kidney_fetal_17w GSE75948.SIX2.kidney_fetal_17w 96 bp overlap
SMAD2-3 1 dataset
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 266 bp overlap
SMAD2_3 2 datasets
ChIP DE_D2 S01-DE-d2-SMAD2_3-exp1 313 bp overlap
ChIP DE_D2 S53-DE-d2-SMAD2_3-exp2 159 bp overlap
SMAD4 1 dataset
ChIP Caco-2 GSE112946.SMAD4.Caco-2 197 bp overlap
SP2 3 datasets
ChIP HEK293 ENCFF181QXT 378 bp overlap
ChIP HEK293 ENCSR807LQP.SP2.HEK293 392 bp overlap
ChIP HEK293 GSE76494.SP2.HEK293 336 bp overlap
SP3 1 dataset
ChIP HEK293 ENCSR141PZA.SP3.HEK293 309 bp overlap
SP7 2 datasets
ChIP HEK293 ENCFF733RBE 144 bp overlap
ChIP HEK293 ENCSR468IJT.SP7.HEK293 392 bp overlap
STAG1 4 datasets
ChIP HeLa GSE126990.STAG1.HeLa 84 bp overlap
ChIP HeLa GSE126990.STAG1.HeLa 104 bp overlap
ChIP HeLa-Kyoto GSE138405.STAG1.HeLa-Kyoto 84 bp overlap
ChIP HeLa-Kyoto GSE138405.STAG1.HeLa-Kyoto 104 bp overlap
STAT3 1 dataset
ChIP HeLa-S3 ENCSR000EDC.STAT3.HeLa-S3 51 bp overlap
TAL1 4 datasets
ChIP CD34 GSE52924.TAL1.CD34 174 bp overlap
ChIP HSPC-CD34pos GSE93372.TAL1.HSPC-CD34pos 92 bp overlap
ChIP K-562_enCRISPRi-LK GSE132212.TAL1.K-562_enCRISPRi-LK 161 bp overlap
ChIP ProEs GSE59087.TAL1.ProEs 192 bp overlap
TBP 2 datasets
ChIP HeLa-S3 ENCFF715NNJ 258 bp overlap
ChIP HeLa-S3 ENCSR000EDD.TBP.HeLa-S3 392 bp overlap
TCF7L2 4 datasets
ChIP HEK293 ENCFF513JQN 214 bp overlap
ChIP HEK293 ENCSR000EUY.TCF7L2.HEK293 224 bp overlap
ChIP HeLa-S3 ENCFF673QAB 146 bp overlap
ChIP HeLa-S3 ENCSR000EVE.TCF7L2.HeLa-S3 64 bp overlap
TEAD1 3 datasets
ChIP HUCCT1 GSE68296.TEAD1.HUCCT1 319 bp overlap
ChIP MSTO GSE68170.TEAD1.MSTO 392 bp overlap
ChIP pancreas_12D ERP008682.TEAD1.pancreas_12D 214 bp overlap
TEAD4 3 datasets
ChIP HEK293_siCtrl GSE130135.TEAD4.HEK293_siCtrl 202 bp overlap
ChIP PC-9_1DF GSE131687.TEAD4.PC-9_1DF 267 bp overlap
ChIP PC-9_2DF GSE131687.TEAD4.PC-9_2DF 284 bp overlap
TSHZ1 3 datasets
ChIP HEK293 ENCFF893BGV 58 bp overlap
ChIP HEK293 ENCFF893BGV 281 bp overlap
ChIP HEK293 ENCSR217WRC.TSHZ1.HEK293 310 bp overlap
WT1 2 datasets
ChIP HEK293 ENCFF906HIR 84 bp overlap
ChIP HEK293 ENCSR966PJJ.WT1.HEK293 392 bp overlap
YY1 2 datasets
ChIP Hep-G2_NC GSE120104.YY1.Hep-G2_NC 102 bp overlap
ChIP Hep-G2_RBM25-KD GSE120104.YY1.Hep-G2_RBM25-KD 392 bp overlap
YY1AP1 4 datasets
ChIP HEK293_siCtrl GSE130135.YY1AP1.HEK293_siCtrl 51 bp overlap
ChIP HEK293_siCtrl GSE130135.YY1AP1.HEK293_siCtrl 222 bp overlap
ChIP PC-9_1DF GSE131687.YY1AP1.PC-9_1DF 254 bp overlap
ChIP PC-9_2DF GSE131687.YY1AP1.PC-9_2DF 236 bp overlap
ZBTB1 1 dataset
ChIP HEK293 ENCFF916DEM 82 bp overlap
ZBTB12 2 datasets
ChIP HEK293 ENCFF963HPT 220 bp overlap
ChIP HEK293 ENCSR543KOA.ZBTB12.HEK293 217 bp overlap
ZBTB17 1 dataset
ChIP HEK293 ENCFF865LIO 131 bp overlap
ZBTB21 2 datasets
ChIP HEK293 ENCFF509WYZ 107 bp overlap
ChIP HEK293 ENCSR321MSF.ZBTB21.HEK293 358 bp overlap
ZBTB44 2 datasets
ChIP HEK293 ENCFF560VPN 138 bp overlap
ChIP HEK293 ENCSR076STQ.ZBTB44.HEK293 392 bp overlap
ZBTB8A 2 datasets
ChIP HEK293 ENCFF303WRD 240 bp overlap
ChIP HEK293 ENCSR481FEC.ZBTB8A.HEK293 374 bp overlap
ZEB1 2 datasets
ChIP HEK293 ENCFF007TAP 284 bp overlap
ChIP HEK293 ENCSR909HMT.ZEB1.HEK293 392 bp overlap
ZEB2 2 datasets
ChIP HEK293 ENCFF847JIE 392 bp overlap
ChIP HEK293 ENCSR417VWF.ZEB2.HEK293 392 bp overlap
ZFP37 1 dataset
ChIP HEK293 ENCSR365GRX.ZFP37.HEK293 353 bp overlap
ZFP64 1 dataset
ChIP HEK293 GSE76494.ZFP64.HEK293 122 bp overlap
ZFP69B 2 datasets
ChIP HEK293 ENCFF942LFP 186 bp overlap
ChIP HEK293 ENCSR381VYR.ZFP69B.HEK293 357 bp overlap
ZHX1 2 datasets
ChIP HeLa-S3 ENCSR855WKQ.ZHX1.HeLa-S3 85 bp overlap
ChIP HeLa-S3 ENCSR855WKQ.ZHX1.HeLa-S3 181 bp overlap
ZIC2 1 dataset
ChIP HEK293 ENCFF033NQQ 67 bp overlap
ZNF146 2 datasets
ChIP HEK293 ENCFF602LWH 108 bp overlap
ChIP HEK293 ENCFF602LWH 307 bp overlap
ZNF18 2 datasets
ChIP HEK293 ENCFF066NGR 166 bp overlap
ChIP HEK293 ENCSR977HTH.ZNF18.HEK293 50 bp overlap
ZNF189 2 datasets
ChIP HEK293 ENCFF638TIB 54 bp overlap
ChIP HEK293 ENCSR163RYW.ZNF189.HEK293 188 bp overlap
ZNF2 1 dataset
ChIP HEK293 ENCSR011CKE.ZNF2.HEK293 386 bp overlap
ZNF24 1 dataset
ChIP HEK293 ENCSR984MDV.ZNF24.HEK293 392 bp overlap
ZNF263 1 dataset
ChIP HEK293 ENCFF336CWQ 57 bp overlap
ZNF280A 1 dataset
ChIP HEK293 GSE76494.ZNF280A.HEK293 249 bp overlap
ZNF283 1 dataset
ChIP HEK293T GSE78099.ZNF283.HEK293T 80 bp overlap
ZNF30 1 dataset
ChIP HEK293 GSE76494.ZNF30.HEK293 220 bp overlap
ZNF302 2 datasets
ChIP HEK293 ENCFF832SDW 211 bp overlap
ChIP HEK293 ENCSR513MGG.ZNF302.HEK293 193 bp overlap
ZNF335 2 datasets
ChIP HEK293 ENCFF784SLD 367 bp overlap
ChIP HEK293 ENCSR328SUD.ZNF335.HEK293 392 bp overlap
ZNF34 1 dataset
ChIP HEK293 ENCSR727PIC.ZNF34.HEK293 371 bp overlap
ZNF35 2 datasets
ChIP HEK293 GSE76494.ZNF35.HEK293 89 bp overlap
ChIP HEK293 GSE76494.ZNF35.HEK293 195 bp overlap
ZNF350 2 datasets
ChIP HEK293 ENCSR854ORP.ZNF350.HEK293 273 bp overlap
ChIP HEK293 GSE76494.ZNF350.HEK293 141 bp overlap
ZNF366 3 datasets
ChIP HEK293 ENCFF799ATK 130 bp overlap
ChIP HEK293 ENCFF799ATK 134 bp overlap
ChIP HEK293 ENCSR106EBH.ZNF366.HEK293 392 bp overlap
ZNF384 2 datasets
ChIP GM12878 ENCFF229VSP 134 bp overlap
ChIP HEK293T ENCFF019DZX 135 bp overlap
ZNF391 2 datasets
ChIP HEK293 ENCFF835SNY 112 bp overlap
ChIP HEK293 ENCSR210MET.ZNF391.HEK293 55 bp overlap
ZNF394 2 datasets
ChIP HEK293 ENCFF236OPX 172 bp overlap
ChIP HEK293 ENCSR125DNC.ZNF394.HEK293 194 bp overlap
ZNF449 2 datasets
ChIP HEK293 ENCFF764ZIC 139 bp overlap
ChIP HEK293 ENCSR738SLS.ZNF449.HEK293 97 bp overlap
ZNF488 2 datasets
ChIP HEK293 ENCFF780TIG 50 bp overlap
ChIP HEK293 ENCSR363XBR.ZNF488.HEK293 58 bp overlap
ZNF518A 3 datasets
ChIP HEK293 ENCFF892ULS 97 bp overlap
ChIP HEK293 ENCFF892ULS 302 bp overlap
ChIP HEK293 ENCSR159GFL.ZNF518A.HEK293 392 bp overlap
ZNF561 2 datasets
ChIP HEK293 ENCFF399XKF 84 bp overlap
ChIP HEK293 ENCSR125ULS.ZNF561.HEK293 216 bp overlap
ZNF580 2 datasets
ChIP HEK293 ENCFF906MQV 195 bp overlap
ChIP HEK293 ENCSR668HOP.ZNF580.HEK293 392 bp overlap
ZNF629 2 datasets
ChIP HEK293 ENCFF096ELQ 392 bp overlap
ChIP HEK293 ENCSR351NON.ZNF629.HEK293 392 bp overlap
ZNF639 2 datasets
ChIP HEK293 ENCFF971ZNH 338 bp overlap
ChIP HEK293 ENCSR080CST.ZNF639.HEK293 392 bp overlap
ZNF654 1 dataset
ChIP HEK293 ENCFF636WIC 128 bp overlap
ZNF660 2 datasets
ChIP HEK293 ENCFF282RUS 141 bp overlap
ChIP HEK293 ENCSR283DOU.ZNF660.HEK293 333 bp overlap
ZNF692 2 datasets
ChIP HEK293 ENCFF040AZE 74 bp overlap
ChIP HEK293 ENCSR418MKG.ZNF692.HEK293 392 bp overlap
ZNF76 1 dataset
ChIP HEK293 ENCSR072LQF.ZNF76.HEK293 205 bp overlap
ZNF843 3 datasets
ChIP HEK293 ENCFF241QRH 72 bp overlap
ChIP HEK293 ENCFF241QRH 372 bp overlap
ChIP HEK293 ENCSR502KPJ.ZNF843.HEK293 392 bp overlap
ZSCAN18 1 dataset
ChIP HEK293 ENCFF537OVZ 72 bp overlap
ZSCAN22 1 dataset
ChIP HEK293 GSE76494.ZSCAN22.HEK293 161 bp overlap
ZSCAN23 3 datasets
ChIP HEK293 ENCFF127TFV 116 bp overlap
ChIP HEK293 ENCFF127TFV 329 bp overlap
ChIP HEK293 ENCSR705ASR.ZSCAN23.HEK293 392 bp overlap
ZSCAN30 1 dataset
ChIP HEK293 ENCFF082YBI 63 bp overlap
ZSCAN4 2 datasets
ChIP HEK293 ENCFF381BKT 166 bp overlap
ChIP HEK293 ENCSR211GNP.ZSCAN4.HEK293 376 bp overlap
ZXDB 2 datasets
ChIP HEK293 ENCFF835SGA 198 bp overlap
ChIP HEK293 ENCSR559IOZ.ZXDB.HEK293 392 bp overlap