chr9 : 14,908,720 14,909,125
405 bp 143 TFs 1 linked gene
This 405 bp open chromatin element is linked to FREM1 and is bound by 143 transcription factors.
Linked Genes
1 gene
Gene Expression Dist. to TSS Distance Link type
FREM1 1.3 kb Proximal Proximity
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr9:14,903,720 – 14,914,125
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
143 transcription factors
Source
Cell type
AR 1 dataset
ChIP epididymis_HEE_R1881 GSE109061.AR.epididymis_HEE_R1881 91 bp overlap
ARID3A 3 datasets
ChIP Hep-G2 ENCSR000EDP.ARID3A.Hep-G2 79 bp overlap
ChIP HepG2 ENCFF341DES 288 bp overlap
ChIP HepG2 ENCFF341DES 405 bp overlap
ARID4B 1 dataset
ChIP HepG2 ENCFF519OXJ 328 bp overlap
ARID5B 1 dataset
ChIP HepG2 ENCFF964FWK 130 bp overlap
ARNTL 1 dataset
ChIP Hep-G2 ENCSR794LVK.ARNTL.Hep-G2 175 bp overlap
ASCL1 1 dataset
ChIP SCLC_ASCLP GSE61197.ASCL1.SCLC_ASCLP 113 bp overlap
ASH2L 2 datasets
ChIP Hep-G2 ENCSR508LMH.ASH2L.Hep-G2 405 bp overlap
ChIP HepG2 ENCFF207QHL 261 bp overlap
BRD3 1 dataset
ChIP HEK293T GSE39579.BRD3.HEK293T 311 bp overlap
BRD4 2 datasets
ChIP NCI-H1963 GSE145028.BRD4.NCI-H1963 204 bp overlap
ChIP SK-N-BE2-C_Resistant-veh GSE107706.BRD4.SK-N-BE2-C_Resistant-veh 277 bp overlap
CDK8 1 dataset
ChIP leiomyoma_PT886 GSE128230.CDK8.leiomyoma_PT886 84 bp overlap
CDX1 1 dataset
Motif DE_12h DE_12h-CDX1_MA0878.3 10 bp overlap
CDX2 1 dataset
Motif DE_12h DE_12h-CDX2_MA0465.3 8 bp overlap
CEBPA 1 dataset
ChIP HepG2 ENCFF175DFS 140 bp overlap
CHD1 2 datasets
ChIP H1 ENCFF998XEK 356 bp overlap
ChIP H1 ENCFF998XEK 151 bp overlap
CREB1 1 dataset
ChIP HepG2 ENCFF576ERP 317 bp overlap
DRAP1 1 dataset
ChIP Hep-G2 ENCSR765MKZ.DRAP1.Hep-G2 159 bp overlap
EGR1 1 dataset
ChIP Hep-G2 ENCSR026GSW.EGR1.Hep-G2 270 bp overlap
EP300 2 datasets
ChIP Hep-G2 ENCSR000BLW.EP300.Hep-G2 199 bp overlap
ChIP HepG2 ENCFF354ACD 197 bp overlap
ESR1 3 datasets
ChIP Ishikawa_ETV4-KO2_Mili GSE129803.ESR1.Ishikawa_ETV4-KO2_Mili 172 bp overlap
ChIP endometrioid-adenocarcinoma_tumor_3 GSE94031.ESR1.endometrioid-adenocarcinoma_tumor_3 119 bp overlap
ChIP primary-endometrium-cancer_E1_DSG GSE114737.ESR1.primary-endometrium-cancer_E1_DSG 199 bp overlap
FEZF1 3 datasets
ChIP HEK293 ENCFF528YED 158 bp overlap
ChIP HEK293 ENCSR827NWO.FEZF1.HEK293 206 bp overlap
ChIP HEK293 GSE76494.FEZF1.HEK293 114 bp overlap
FEZF2 1 dataset
Motif DE_12h DE_12h-FEZF2_MA2341.1 8 bp overlap
FOXA1 11 datasets
ChIP 22Rv1_CST GSE129951.FOXA1.22Rv1_CST 164 bp overlap
ChIP 22Rv1_R1881 GSE80742.FOXA1.22Rv1_R1881 120 bp overlap
ChIP CFPAC-1 GSE119930.FOXA1.CFPAC-1 207 bp overlap
ChIP CFPAC-1_FOXA2-KO GSE119930.FOXA1.CFPAC-1_FOXA2-KO 147 bp overlap
ChIP HepG2 ENCFF207NVJ 126 bp overlap
ChIP HepG2 ENCFF361KNY 170 bp overlap
ChIP HepG2 ENCFF740VZW 110 bp overlap
ChIP MCF-7_estrogen_ab2 GSE112969.FOXA1.MCF-7_estrogen_ab2 200 bp overlap
ChIP PDAC GSE64557.FOXA1.PDAC 273 bp overlap
ChIP PDAC_KOKLF5 GSE64557.FOXA1.PDAC_KOKLF5 167 bp overlap
ChIP PDAC_SHCTR GSE64557.FOXA1.PDAC_SHCTR 321 bp overlap
FOXA2 10 datasets
ChIP CFPAC-1 GSE119930.FOXA2.CFPAC-1 240 bp overlap
ChIP CFPAC-1_EV GSE119930.FOXA2.CFPAC-1_EV 284 bp overlap
ChIP CFPAC-1_HOXB8-OE GSE119930.FOXA2.CFPAC-1_HOXB8-OE 158 bp overlap
ChIP DE DE-FOXA2-1 324 bp overlap
ChIP DE DE-FOXA2-2 279 bp overlap
Motif DE_12h DE_12h-FOXA2_MA0047.4 8 bp overlap
ChIP HepG2 ENCFF533COJ 123 bp overlap
ChIP HepG2 ENCFF570ABM 151 bp overlap
ChIP HepG2 ENCFF894AYY 158 bp overlap
ChIP pancreas_CARN1618 ERP008682.FOXA2.pancreas_CARN1618 177 bp overlap
FOXA3 1 dataset
ChIP HepG2 ENCFF005KGL 103 bp overlap
FOXG1 1 dataset
Motif DE_12h DE_12h-FOXG1_MA0613.1 8 bp overlap
FOXJ3 1 dataset
ChIP HepG2 ENCFF430OSX 127 bp overlap
FOXK1 3 datasets
Motif DE_12h DE_12h-FOXK1_MA0852.3 7 bp overlap
ChIP Hep-G2 ENCSR603BJQ.FOXK1.Hep-G2 176 bp overlap
ChIP HepG2 ENCFF635XWY 163 bp overlap
FOXK2 1 dataset
Motif DE_12h DE_12h-FOXK2_MA1103.3 7 bp overlap
FOXL1 1 dataset
Motif DE_12h DE_12h-FOXL1_MA0033.2 7 bp overlap
FOXO3 1 dataset
ChIP Hep-G2 GSE97661.FOXO3.Hep-G2 117 bp overlap
FOXO4 1 dataset
Motif DE_12h DE_12h-FOXO4_MA0848.1 7 bp overlap
FOXO6 1 dataset
Motif DE_12h DE_12h-FOXO6_MA0849.1 7 bp overlap
FOXP1 1 dataset
ChIP Hep-G2 ENCSR369YUK.FOXP1.Hep-G2 225 bp overlap
FOXP2 1 dataset
Motif DE_12h DE_12h-FOXP2_MA0593.2 9 bp overlap
FOXP3 1 dataset
Motif DE_12h DE_12h-FOXP3_MA0850.1 7 bp overlap
FOXP4 2 datasets
ChIP Hep-G2 ENCSR232LLP.FOXP4.Hep-G2 184 bp overlap
ChIP HepG2 ENCFF462ULY 213 bp overlap
FOXS1 1 dataset
Motif DE_12h DE_12h-FOXS1_MA2118.1 8 bp overlap
Foxf1 1 dataset
Motif DE_12h DE_12h-Foxf1_MA1606.2 7 bp overlap
Foxo1 1 dataset
Motif DE_12h DE_12h-Foxo1_MA0480.3 7 bp overlap
Foxo3 1 dataset
Motif DE_12h DE_12h-Foxo3_MA0157.4 7 bp overlap
GATA4 2 datasets
ChIP DE DE-GATA4-1 172 bp overlap
ChIP DE DE-GATA4-2 330 bp overlap
GATA6 1 dataset
ChIP DE DE-GATA6-2 328 bp overlap
GATAD2B 2 datasets
ChIP HepG2 ENCFF829IBY 233 bp overlap
ChIP HepG2 ENCFF829IBY 405 bp overlap
GMEB1 1 dataset
ChIP Hep-G2 ENCSR780OXL.GMEB1.Hep-G2 241 bp overlap
HDAC1 4 datasets
ChIP Hep-G2 ENCSR750OWO.HDAC1.Hep-G2 187 bp overlap
ChIP HepG2 ENCFF304IEJ 360 bp overlap
ChIP HepG2 ENCFF750ZWM 405 bp overlap
ChIP HepG2 ENCFF750ZWM 225 bp overlap
HDAC2 3 datasets
ChIP Hep-G2 ENCSR000BMC.HDAC2.Hep-G2 254 bp overlap
ChIP HepG2 ENCFF087XCR 274 bp overlap
ChIP HepG2 ENCFF990GUQ 263 bp overlap
HDAC3 1 dataset
ChIP RH4_DMSO-6H GSE116344.HDAC3.RH4_DMSO-6H 214 bp overlap
HNF1A 5 datasets
ChIP HEE_1 GSE76376.HNF1A.HEE_1 117 bp overlap
ChIP HEE_5 GSE76376.HNF1A.HEE_5 85 bp overlap
ChIP Hep-G2 ENCSR800QIT.HNF1A.Hep-G2 133 bp overlap
ChIP HepG2 ENCFF352VYI 197 bp overlap
ChIP HepG2 ENCFF540TRC 277 bp overlap
HNF1B 2 datasets
ChIP Hep-G2 ENCSR127XTZ.HNF1B.Hep-G2 316 bp overlap
ChIP PDAC GSE64557.HNF1B.PDAC 148 bp overlap
HNF4A 3 datasets
ChIP Hep-G2 ENCSR000BLF.HNF4A.Hep-G2 106 bp overlap
ChIP Hep-G2 ENCSR469FBY.HNF4A.Hep-G2 170 bp overlap
ChIP HepG2 ENCFF146SSF 224 bp overlap
HNF4G 1 dataset
ChIP Hep-G2 ENCSR000BNJ.HNF4G.Hep-G2 114 bp overlap
HOXA3 1 dataset
ChIP Hep-G2 ENCSR999JKC.HOXA3.Hep-G2 170 bp overlap
ISL2 1 dataset
ChIP Hep-G2 ENCSR492IHH.ISL2.Hep-G2 206 bp overlap
KMT2A 1 dataset
ChIP Hep-G2 ENCSR692GFR.KMT2A.Hep-G2 258 bp overlap
LIN54 2 datasets
ChIP Hep-G2 ENCSR597MCV.LIN54.Hep-G2 133 bp overlap
ChIP HepG2 ENCFF662XDE 123 bp overlap
MAX 3 datasets
ChIP Hep-G2 ENCSR168DYA.MAX.Hep-G2 297 bp overlap
ChIP Hep-G2 ENCSR000BTM.MAX.Hep-G2 118 bp overlap
ChIP MCF-7 ENCSR000BUL.MAX.MCF-7 109 bp overlap
MED1 2 datasets
ChIP Hep-G2 ENCSR959XNY.MED1.Hep-G2 290 bp overlap
ChIP HepG2 ENCFF495TSS 322 bp overlap
MEF2A 1 dataset
ChIP Hep-G2 ENCSR291MJH.MEF2A.Hep-G2 150 bp overlap
MSC 1 dataset
Motif DE_12h DE_12h-MSC_MA0665.1 10 bp overlap
MXD4 1 dataset
ChIP Hep-G2 ENCSR441KFW.MXD4.Hep-G2 189 bp overlap
MYBL2 1 dataset
ChIP Hep-G2 ENCSR581KCO.MYBL2.Hep-G2 138 bp overlap
MYPOP 2 datasets
ChIP HepG2 ENCFF176TQL 405 bp overlap
ChIP HepG2 ENCFF176TQL 165 bp overlap
NCOA2 1 dataset
ChIP HepG2 ENCFF853BJJ 245 bp overlap
NFIA 2 datasets
ChIP Hep-G2 GSE97661.NFIA.Hep-G2 299 bp overlap
ChIP HepG2 ENCFF815HWK 192 bp overlap
NFIC 6 datasets
ChIP Hep-G2 GSE108514.NFIC.Hep-G2 186 bp overlap
ChIP Hep-G2 ENCSR000BQX.NFIC.Hep-G2 169 bp overlap
ChIP HepG2 ENCFF169TKU 332 bp overlap
ChIP Ishikawa ENCSR000BUT.NFIC.Ishikawa 151 bp overlap
ChIP K-562 ENCSR796ITY.NFIC.K-562 221 bp overlap
ChIP K562 ENCFF167YID 289 bp overlap
NFIL3 1 dataset
ChIP HepG2 ENCFF686VLI 177 bp overlap
NIPBL 1 dataset
ChIP Hep-G2 GSE76893.NIPBL.Hep-G2 79 bp overlap
NR1H2 1 dataset
ChIP HT29_GW3965_48H GSE77039.NR1H2.HT29_GW3965_48H 196 bp overlap
NR2F2 1 dataset
ChIP Hep-G2 ENCSR000BVM.NR2F2.Hep-G2 174 bp overlap
NR2F6 3 datasets
ChIP Hep-G2 ENCSR518WPL.NR2F6.Hep-G2 320 bp overlap
ChIP HepG2 ENCFF429VKC 237 bp overlap
ChIP HepG2 ENCFF514UJI 207 bp overlap
NR5A1 2 datasets
ChIP Hep-G2 ENCSR310OZS.NR5A1.Hep-G2 187 bp overlap
ChIP HepG2 ENCFF970YZO 228 bp overlap
Nr2e3 1 dataset
Motif DE_12h DE_12h-Nr2e3_MA0164.2 6 bp overlap
PAX5 3 datasets
ChIP GM12878 ENCSR000BHJ.PAX5.GM12878 107 bp overlap
ChIP GM12878 ENCSR000BHD.PAX5.GM12878 95 bp overlap
ChIP NALM-6 GSE126300.PAX5.NALM-6 250 bp overlap
PAX6 1 dataset
Motif DE_12h DE_12h-PAX6_MA0069.1 14 bp overlap
PAXIP1 2 datasets
ChIP Hep-G2 ENCSR530ARJ.PAXIP1.Hep-G2 405 bp overlap
ChIP HepG2 ENCFF526NOJ 310 bp overlap
PHF8 2 datasets
ChIP Hep-G2 ENCSR604VAE.PHF8.Hep-G2 209 bp overlap
ChIP HepG2 ENCFF065NWR 405 bp overlap
PITX1 1 dataset
ChIP HepG2 ENCFF468QTQ 295 bp overlap
POGK 1 dataset
ChIP HepG2 ENCFF029WNT 176 bp overlap
POLR2A 1 dataset
ChIP body of pancreas ENCFF727UBE 405 bp overlap
POLR2G 4 datasets
ChIP HepG2 ENCFF241AEG 82 bp overlap
ChIP HepG2 ENCFF241AEG 405 bp overlap
ChIP HepG2 ENCFF508UTS 82 bp overlap
ChIP HepG2 ENCFF508UTS 405 bp overlap
PSIP1 1 dataset
ChIP T-REx-293 GSE117155.PSIP1.T-REx-293 141 bp overlap
RAD21 1 dataset
ChIP HepG2 ENCFF360ZSW 202 bp overlap
RARA 1 dataset
ChIP HepG2 ENCFF582XUA 202 bp overlap
RBPJ 1 dataset
Motif DE_12h DE_12h-RBPJ_MA1116.2 6 bp overlap
SALL1 1 dataset
ChIP HepG2 ENCFF426MCK 297 bp overlap
SAP130 1 dataset
ChIP HepG2 ENCFF892EHZ 187 bp overlap
SIX1 1 dataset
ChIP Hep-G2 ENCSR561BQM.SIX1.Hep-G2 302 bp overlap
SIX2 3 datasets
ChIP HEK GSE73865.SIX2.HEK 227 bp overlap
ChIP kidney_fetal_16w GSE75948.SIX2.kidney_fetal_16w 187 bp overlap
ChIP kidney_fetal_17w GSE75948.SIX2.kidney_fetal_17w 214 bp overlap
SMAD3 1 dataset
ChIP Hep-G2 ENCSR005GZH.SMAD3.Hep-G2 275 bp overlap
SMAD7 1 dataset
ChIP HepG2 ENCFF850FXR 81 bp overlap
SMARCA4 1 dataset
ChIP HS-SY-2_HA-tagged GSE108926.SMARCA4.HS-SY-2_HA-tagged 333 bp overlap
SNAI1 1 dataset
ChIP HepG2 ENCFF017SIW 91 bp overlap
SNAI2 1 dataset
ChIP SMS-CTR_shSNAI2 GSE137168.SNAI2.SMS-CTR_shSNAI2 190 bp overlap
SOX13 1 dataset
ChIP HepG2 ENCFF062VSQ 216 bp overlap
SOX6 2 datasets
ChIP Hep-G2 ENCSR766TSU.SOX6.Hep-G2 364 bp overlap
ChIP HepG2 ENCFF767OCK 354 bp overlap
TBX2 2 datasets
ChIP Hep-G2 ENCSR410FEH.TBX2.Hep-G2 247 bp overlap
ChIP HepG2 ENCFF811TLA 379 bp overlap
TCF7L2 2 datasets
ChIP Hep-G2 ENCSR346BOD.TCF7L2.Hep-G2 142 bp overlap
ChIP HepG2 ENCFF510OLG 254 bp overlap
TEAD1 1 dataset
ChIP HepG2 ENCFF661PNM 157 bp overlap
TEAD4 1 dataset
ChIP Hep-G2 ENCSR000BRP.TEAD4.Hep-G2 138 bp overlap
TFAP4 3 datasets
ChIP Hep-G2 ENCSR103SZL.TFAP4.Hep-G2 238 bp overlap
ChIP HepG2 ENCFF030SRU 128 bp overlap
ChIP HepG2 ENCFF932XOY 180 bp overlap
TFE3 1 dataset
ChIP Hep-G2 ENCSR589SNT.TFE3.Hep-G2 212 bp overlap
TGIF2 2 datasets
ChIP Hep-G2 ENCSR993LMB.TGIF2.Hep-G2 101 bp overlap
ChIP HepG2 ENCFF421ZJN 244 bp overlap
THRA 1 dataset
ChIP HepG2 ENCFF025KMX 122 bp overlap
THRB 2 datasets
ChIP Hep-G2 ENCSR430JGJ.THRB.Hep-G2 211 bp overlap
ChIP HepG2 ENCFF476INC 112 bp overlap
TSC22D4 1 dataset
ChIP Hep-G2 GSE97661.TSC22D4.Hep-G2 156 bp overlap
Tcf21 1 dataset
Motif DE_12h DE_12h-Tcf21_MA0832.2 10 bp overlap
VDR 1 dataset
ChIP kidney-cortex GSE129585.VDR.kidney-cortex 193 bp overlap
YY1 2 datasets
ChIP Hep-G2_NC GSE120104.YY1.Hep-G2_NC 321 bp overlap
ChIP Hep-G2_RBM25-KD GSE120104.YY1.Hep-G2_RBM25-KD 177 bp overlap
ZBED4 1 dataset
ChIP Hep-G2 ENCSR409PMR.ZBED4.Hep-G2 242 bp overlap
ZBTB6 1 dataset
ChIP HEK293 GSE76494.ZBTB6.HEK293 153 bp overlap
ZBTB7A 1 dataset
ChIP VCaP_FBS GSE123089.ZBTB7A.VCaP_FBS 215 bp overlap
ZBTB7B 2 datasets
ChIP Hep-G2 ENCSR322ULL.ZBTB7B.Hep-G2 405 bp overlap
ChIP HepG2 ENCFF763OCV 292 bp overlap
ZEB1 2 datasets
ChIP Hep-G2 ENCSR000BVN.ZEB1.Hep-G2 189 bp overlap
ChIP HepG2 ENCFF808RQT 334 bp overlap
ZFP64 1 dataset
ChIP HEK293 GSE76494.ZFP64.HEK293 155 bp overlap
ZFX 1 dataset
ChIP HepG2 ENCFF016NZF 405 bp overlap
ZFY 3 datasets
ChIP Hep-G2 ENCSR949OEV.ZFY.Hep-G2 155 bp overlap
ChIP HepG2 ENCFF106ELT 405 bp overlap
ChIP HepG2 ENCFF106ELT 405 bp overlap
ZGPAT 1 dataset
ChIP Hep-G2 ENCSR704IGU.ZGPAT.Hep-G2 199 bp overlap
ZIC1 1 dataset
Motif DE_12h DE_12h-ZIC1_MA0696.1 14 bp overlap
ZIC4 1 dataset
Motif DE_12h DE_12h-ZIC4_MA0751.2 14 bp overlap
ZNF189 1 dataset
Motif DE_12h DE_12h-ZNF189_MA1725.2 9 bp overlap
ZNF217 2 datasets
ChIP Hep-G2 ENCSR055FQB.ZNF217.Hep-G2 298 bp overlap
ChIP HepG2 ENCFF455XGO 339 bp overlap
ZNF219 1 dataset
ChIP HepG2 ENCFF266JIR 246 bp overlap
ZNF341 4 datasets
Motif DE_12h DE_12h-ZNF341_MA1655.2 8 bp overlap
ChIP HEK293 ENCFF944VMC 196 bp overlap
ChIP HEK293 ENCSR185FOY.ZNF341.HEK293 341 bp overlap
ChIP HEK293 GSE76494.ZNF341.HEK293 181 bp overlap
ZNF44 1 dataset
ChIP Hep-G2 ENCSR914HPP.ZNF44.Hep-G2 217 bp overlap
ZNF501 2 datasets
ChIP Hep-G2 ENCSR227PHM.ZNF501.Hep-G2 322 bp overlap
ChIP HepG2 ENCFF879XZR 150 bp overlap
ZNF503 1 dataset
ChIP HepG2 ENCFF923HZL 134 bp overlap
ZNF609 2 datasets
ChIP Hep-G2 ENCSR407BPU.ZNF609.Hep-G2 274 bp overlap
ChIP HepG2 ENCFF900FRP 262 bp overlap
ZNF652 2 datasets
ChIP Hep-G2 ENCSR502GAX.ZNF652.Hep-G2 405 bp overlap
ChIP HepG2 ENCFF331VPZ 227 bp overlap
ZNF708 1 dataset
Motif DE_12h DE_12h-ZNF708_MA1730.2 9 bp overlap
ZNF816 2 datasets
ChIP HepG2 ENCFF294VPD 405 bp overlap
ChIP HepG2 ENCFF294VPD 405 bp overlap
ZSCAN29 1 dataset
ChIP HepG2 ENCFF212SBM 405 bp overlap
Zfp809 1 dataset
Motif DE_12h DE_12h-Zfp809_MA2125.1 9 bp overlap