chr11 : 113,884,168 113,884,605
437 bp 114 TFs 1 linked gene
This 437 bp open chromatin element is linked to USP28 and is bound by 114 transcription factors.
Linked Genes
1 gene
Gene Expression Dist. to TSS Distance Link type
USP28 8.6 kb Proximal Proximity
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr11:113,879,168 – 113,889,605
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
114 transcription factors
Source
Cell type
ARID1A 2 datasets
ChIP RMG-I GSE120058.ARID1A.RMG-I 181 bp overlap
ChIP RMG-I GSE104545.ARID1A.RMG-I 152 bp overlap
ARID1B 2 datasets
ChIP K-562 ENCSR822CCM.ARID1B.K-562 206 bp overlap
ChIP K562 ENCFF938UXQ 329 bp overlap
ARID5B 1 dataset
ChIP HepG2 ENCFF964FWK 334 bp overlap
ARNT 1 dataset
ChIP K-562 ENCSR613NUC.ARNT.K-562 103 bp overlap
ASH2L 1 dataset
ChIP WA01 ENCSR850KIP.ASH2L.WA01 393 bp overlap
ATF2 10 datasets
ChIP H1 ENCFF295GZO 394 bp overlap
ChIP HEK293 ENCFF194VKZ 281 bp overlap
ChIP HEK293 ENCFF194VKZ 71 bp overlap
ChIP HEK293 ENCSR217HTK.ATF2.HEK293 237 bp overlap
ChIP Hep-G2 ENCSR047BUZ.ATF2.Hep-G2 215 bp overlap
ChIP HepG2 ENCFF578ZBI 173 bp overlap
ChIP HepG2 ENCFF955VER 289 bp overlap
ChIP K-562 ENCSR869IUD.ATF2.K-562 250 bp overlap
ChIP K562 ENCFF139ZZG 188 bp overlap
ChIP WA01 ENCSR000BQU.ATF2.WA01 150 bp overlap
ATF3 15 datasets
ChIP H1 ENCFF852GZY 61 bp overlap
ChIP HCT-116_CAMP GSE74355.ATF3.HCT-116_CAMP 126 bp overlap
ChIP Hep-G2 ENCSR402ZCY.ATF3.Hep-G2 173 bp overlap
ChIP Hep-G2 ENCSR000BKE.ATF3.Hep-G2 145 bp overlap
ChIP HepG2 ENCFF832LTU 193 bp overlap
ChIP HepG2 ENCFF928LDD 210 bp overlap
ChIP K-562 ENCSR028UIU.ATF3.K-562 350 bp overlap
ChIP K-562 ENCSR632DCH.ATF3.K-562 331 bp overlap
ChIP K562 ENCFF604FPV 376 bp overlap
ChIP K562 ENCFF921JQW 263 bp overlap
ChIP K562 ENCFF965VXT 209 bp overlap
ChIP WA01 ENCSR000BKC.ATF3.WA01 171 bp overlap
ChIP WTC11 ENCFF519QFH 276 bp overlap
ChIP liver ENCFF375GID 310 bp overlap
ChIP liver ENCFF867MFZ 301 bp overlap
ATF4 4 datasets
ChIP HepG2 ENCFF819ULE 267 bp overlap
ChIP HepG2 ENCFF903ADR 213 bp overlap
ChIP K-562 ENCSR145TSJ.ATF4.K-562 370 bp overlap
ChIP K562 ENCFF674KTF 375 bp overlap
ATF7 5 datasets
ChIP Hep-G2 ENCSR545FXC.ATF7.Hep-G2 267 bp overlap
ChIP HepG2 ENCFF470FKK 283 bp overlap
ChIP HepG2 ENCFF589EBD 339 bp overlap
ChIP K-562 ENCSR972ZBV.ATF7.K-562 359 bp overlap
ChIP K562 ENCFF308SKS 248 bp overlap
BRD4 3 datasets
ChIP K-562_DMSO GSE99178.BRD4.K-562_DMSO 71 bp overlap
ChIP Kelly_sensitive GSE103030.BRD4.Kelly_sensitive 117 bp overlap
ChIP SUM159PT_scramble GSE131097.BRD4.SUM159PT_scramble 245 bp overlap
CBX5 1 dataset
ChIP HepG2 ENCFF251YQZ 304 bp overlap
CEBPA 4 datasets
ChIP Hep-G2 ERP000209.CEBPA.Hep-G2 163 bp overlap
ChIP HepG2 ENCFF175DFS 210 bp overlap
ChIP T-47D_progesterone GSE132649.CEBPA.T-47D_progesterone 213 bp overlap
ChIP T-47D_siCtrl GSE132649.CEBPA.T-47D_siCtrl 207 bp overlap
CEBPB 19 datasets
ChIP H1 ENCFF871PTR 143 bp overlap
ChIP HCT-116 ENCSR000BSD.CEBPB.HCT-116 160 bp overlap
ChIP HCT116 ENCFF097OLY 290 bp overlap
ChIP HFOB_DIFF GSE82295.CEBPB.HFOB_DIFF 218 bp overlap
ChIP HeLa-S3 ENCFF722WEG 169 bp overlap
ChIP Hep-G2 ENCSR000EEX.CEBPB.Hep-G2 163 bp overlap
ChIP Hep-G2 ENCSR000BQI.CEBPB.Hep-G2 161 bp overlap
ChIP HepG2 ENCFF074JWB 94 bp overlap
ChIP HepG2 ENCFF536NTI 109 bp overlap
ChIP IMR-90 ENCFF468UGY 177 bp overlap
ChIP Ishikawa ENCFF010USJ 82 bp overlap
ChIP Ishikawa ENCSR000BTT.CEBPB.Ishikawa 149 bp overlap
ChIP K-562 ENCSR000EHE.CEBPB.K-562 154 bp overlap
ChIP K562 ENCFF189VBN 239 bp overlap
ChIP K562 ENCFF194QGF 262 bp overlap
ChIP K562 ENCFF584CTB 352 bp overlap
ChIP MCF-7 ENCFF772ZTQ 167 bp overlap
ChIP MCF-7 ENCSR000BSR.CEBPB.MCF-7 162 bp overlap
ChIP WA01 ENCSR000EBV.CEBPB.WA01 167 bp overlap
CEBPD 2 datasets
ChIP Hep-G2 ENCSR000BQJ.CEBPD.Hep-G2 133 bp overlap
ChIP HepG2 ENCFF345JDB 172 bp overlap
CEBPG 6 datasets
ChIP HepG2 ENCFF503XBC 225 bp overlap
ChIP K-562 ENCSR490LWA.CEBPG.K-562 327 bp overlap
ChIP K-562 ENCSR620VIC.CEBPG.K-562 240 bp overlap
ChIP K562 ENCFF651CMK 302 bp overlap
ChIP K562 ENCFF783ADE 213 bp overlap
ChIP K562 ENCFF956TPS 228 bp overlap
CREB1 2 datasets
ChIP GM23338 ENCFF432ZEW 264 bp overlap
ChIP GM23338 ENCSR214ZAV.CREB1.GM23338 191 bp overlap
CTBP1 2 datasets
ChIP K-562 ENCSR201NQZ.CTBP1.K-562 326 bp overlap
ChIP K562 ENCFF403WPG 178 bp overlap
CTCF 6 datasets
ChIP AG10803 ENCFF549AQK 181 bp overlap
ChIP Caco-2 ENCFF934QYS 157 bp overlap
ChIP HepG2 ENCFF757EKU 247 bp overlap
ChIP Kelly_resistant GSE103030.CTCF.Kelly_resistant 121 bp overlap
ChIP OVCAR-8 GSE70764.CTCF.OVCAR-8 130 bp overlap
ChIP fibroblast_PULMONARY_ARTERY ENCSR000DUX.CTCF.fibroblast_PULMONARY_ARTERY 116 bp overlap
CTCFL 2 datasets
ChIP FT282 GSE131931.CTCFL.FT282 153 bp overlap
ChIP OVCAR-8 GSE70764.CTCFL.OVCAR-8 169 bp overlap
CXXC5 1 dataset
ChIP K562 ENCFF497CZN 302 bp overlap
Cebpa 6 datasets
ChIP BLaER1 ENCFF031ISE 317 bp overlap
ChIP BLaER1 ENCFF093OYK 251 bp overlap
ChIP BLaER1 ENCFF274GAT 195 bp overlap
ChIP BLaER1 ENCFF364PUR 335 bp overlap
ChIP BLaER1 ENCFF844FIP 267 bp overlap
ChIP BLaER1 ENCFF896HSY 403 bp overlap
DPF2 2 datasets
ChIP K-562 ENCSR219BXP.DPF2.K-562 140 bp overlap
ChIP K562 ENCFF739JDE 281 bp overlap
DRAP1 1 dataset
ChIP Hep-G2 ENCSR765MKZ.DRAP1.Hep-G2 122 bp overlap
EGR1 1 dataset
ChIP Hep-G2 ENCSR026GSW.EGR1.Hep-G2 156 bp overlap
EP300 1 dataset
ChIP K-562 ENCSR000EGE.EP300.K-562 139 bp overlap
ERF 2 datasets
ChIP HepG2 ENCFF647PIT 320 bp overlap
ChIP HepG2 ENCFF647PIT 311 bp overlap
FOSL2 1 dataset
ChIP HepG2 ENCFF548CXY 274 bp overlap
FOXA1 4 datasets
ChIP T-47D_JC4743 GSE126004.FOXA1.T-47D_JC4743 80 bp overlap
ChIP T-47D_JC4745 GSE126004.FOXA1.T-47D_JC4745 57 bp overlap
ChIP T-47D_JC4748 GSE126004.FOXA1.T-47D_JC4748 89 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.FOXA1.pancreatic-progenitor_PP1 119 bp overlap
FOXA2 2 datasets
ChIP DE DE-FOXA2-1 343 bp overlap
ChIP DE DE-FOXA2-2 286 bp overlap
GABPB1 1 dataset
ChIP WTC11 ENCFF166QKI 203 bp overlap
GATA2 1 dataset
ChIP Hep-G2 ENCSR897LDT.GATA2.Hep-G2 114 bp overlap
GATA3 2 datasets
ChIP Kelly GSE94822.GATA3.Kelly 121 bp overlap
ChIP Kelly GSE65664.GATA3.Kelly 100 bp overlap
GATA4 2 datasets
ChIP DE DE-GATA4-1 201 bp overlap
ChIP DE DE-GATA4-2 437 bp overlap
GATA6 3 datasets
ChIP DE DE-GATA6-2 437 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 153 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 430 bp overlap
GATAD2A 1 dataset
ChIP HepG2 ENCFF252XNH 126 bp overlap
GLIS1 2 datasets
ChIP HEK293 ENCFF299RSE 239 bp overlap
ChIP HEK293 ENCSR482BBZ.GLIS1.HEK293 276 bp overlap
GLIS3 1 dataset
ChIP H9_plus GSE109562.GLIS3.H9_plus 143 bp overlap
GTF3C2 1 dataset
ChIP H9 GSE94418.GTF3C2.H9 204 bp overlap
HDAC1 1 dataset
ChIP K562 ENCFF968WBH 383 bp overlap
HLF 2 datasets
ChIP Hep-G2 ENCSR528PSI.HLF.Hep-G2 141 bp overlap
ChIP HepG2 ENCFF854JLR 219 bp overlap
IKZF1 3 datasets
ChIP K-562 ENCSR395HWC.IKZF1.K-562 357 bp overlap
ChIP K562 ENCFF348IBL 338 bp overlap
ChIP K562 ENCFF771OHZ 250 bp overlap
IKZF3 2 datasets
ChIP HEK293 ENCFF518OXG 255 bp overlap
ChIP HEK293 ENCSR304AMN.IKZF3.HEK293 172 bp overlap
IRF9 1 dataset
ChIP K562 ENCFF346PQL 209 bp overlap
ISL2 1 dataset
ChIP Hep-G2 ENCSR492IHH.ISL2.Hep-G2 121 bp overlap
JUN 7 datasets
ChIP DE_D1 S08-DE-d1-JUN-exp1 264 bp overlap
ChIP DE_D1 S40-DE-d1-JUN-exp2 321 bp overlap
ChIP DE_D2 S02-DE-d2-JUN-exp1 407 bp overlap
ChIP DE_D2 S54-DE-d2-JUN-exp2 280 bp overlap
ChIP ESC S24-ESC-d0-JUN-exp1 359 bp overlap
ChIP ESC S34-ESC-d0-JUN-exp2 195 bp overlap
ChIP HUES-8 GSE109524.JUN.HUES-8 357 bp overlap
JUND 10 datasets
ChIP H1 ENCFF010YXS 259 bp overlap
ChIP H1 ENCFF468JZD 229 bp overlap
ChIP HeLa-S3 ENCSR000EDH.JUND.HeLa-S3 142 bp overlap
ChIP Hep-G2 ENCSR000BGK.JUND.Hep-G2 123 bp overlap
ChIP HepG2 ENCFF172HFZ 237 bp overlap
ChIP HepG2 ENCFF869OPW 229 bp overlap
ChIP K-562 ENCSR000EGN.JUND.K-562 150 bp overlap
ChIP K562 ENCFF336RCR 319 bp overlap
ChIP K562 ENCFF830LVJ 214 bp overlap
ChIP WA01 ENCSR000EBZ.JUND.WA01 156 bp overlap
KLF5 1 dataset
ChIP HEK293_E419K GSE88976.KLF5.HEK293_E419K 184 bp overlap
L3MBTL2 2 datasets
ChIP K-562 ENCSR530XQI.L3MBTL2.K-562 329 bp overlap
ChIP K562 ENCFF320EQC 416 bp overlap
MAFF 2 datasets
ChIP HeLa-S3 ENCFF783SBT 205 bp overlap
ChIP HeLa-S3 ENCSR140DSL.MAFF.HeLa-S3 139 bp overlap
MED1 1 dataset
ChIP Hep-G2 ENCSR959XNY.MED1.Hep-G2 154 bp overlap
MTA3 1 dataset
ChIP K-562 ENCSR180NCY.MTA3.K-562 253 bp overlap
NELFE 2 datasets
ChIP HeLa GSE125534.NELFE.HeLa 107 bp overlap
ChIP HeLa_DOX GSE125534.NELFE.HeLa_DOX 114 bp overlap
NFIL3 1 dataset
ChIP HepG2 ENCFF686VLI 198 bp overlap
NKX2-1 1 dataset
ChIP H9_derived-cIN GSE99937.NKX2-1.H9_derived-cIN 61 bp overlap
NR3C1 2 datasets
ChIP BEAS-2B_TNF_IA1 GSE125623.NR3C1.BEAS-2B_TNF_IA1 207 bp overlap
ChIP BEAS-2B_Veh_IA1 GSE125623.NR3C1.BEAS-2B_Veh_IA1 217 bp overlap
OSR2 2 datasets
ChIP HEK293 ENCFF875BDB 362 bp overlap
ChIP HEK293 ENCSR324LTM.OSR2.HEK293 209 bp overlap
PATZ1 3 datasets
ChIP HEK293 ENCFF016MNJ 199 bp overlap
ChIP HEK293 ENCSR966ULI.PATZ1.HEK293 335 bp overlap
ChIP HepG2 ENCFF723PFC 171 bp overlap
POU5F1 2 datasets
ChIP hESC_NAIVE GSE69646.POU5F1.hESC_NAIVE 170 bp overlap
ChIP hiPSC GSE56567.POU5F1.hiPSC 114 bp overlap
PRDM10 2 datasets
ChIP HEK293 ENCFF145WQQ 246 bp overlap
ChIP Hep-G2 ENCSR289PSX.PRDM10.Hep-G2 145 bp overlap
PRDM14 1 dataset
ChIP primordial-germ-cell-like-cell GSE138674.PRDM14.primordial-germ-cell-like-cell 156 bp overlap
PSIP1 1 dataset
ChIP T-REx-293 GSE117155.PSIP1.T-REx-293 152 bp overlap
RAD21 1 dataset
ChIP Hep-G2 ENCSR054FKH.RAD21.Hep-G2 103 bp overlap
RARA 1 dataset
ChIP hiPSC_D5 GSE132532.RARA.hiPSC_D5 189 bp overlap
RBPJ 1 dataset
ChIP HepG2 ENCFF367CFI 147 bp overlap
RELA 1 dataset
ChIP 786-O GSE86092.RELA.786-O 191 bp overlap
REPIN1 2 datasets
ChIP HepG2 ENCFF598VSY 302 bp overlap
ChIP HepG2 ENCFF598VSY 103 bp overlap
REST 2 datasets
ChIP K-562 ENCSR000BMW.REST.K-562 115 bp overlap
ChIP K562 ENCFF430APM 192 bp overlap
RREB1 1 dataset
ChIP HepG2 ENCFF986CSN 113 bp overlap
SALL1 1 dataset
ChIP HepG2 ENCFF426MCK 365 bp overlap
SAP130 1 dataset
ChIP HepG2 ENCFF892EHZ 334 bp overlap
SMAD2-3 1 dataset
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 420 bp overlap
SMAD3 1 dataset
ChIP Hep-G2 ENCSR005GZH.SMAD3.Hep-G2 138 bp overlap
SMAD4 1 dataset
ChIP HepG2 ENCFF615GTE 251 bp overlap
SMARCA4 2 datasets
ChIP K-562 ENCSR587OQL.SMARCA4.K-562 205 bp overlap
ChIP K562 ENCFF506JCB 304 bp overlap
SMARCE1 2 datasets
ChIP K-562 ENCSR157TCS.SMARCE1.K-562 161 bp overlap
ChIP K562 ENCFF690CFF 294 bp overlap
SOX13 1 dataset
ChIP HepG2 ENCFF062VSQ 231 bp overlap
TAF1 1 dataset
ChIP WA01 ENCSR000BHO.TAF1.WA01 145 bp overlap
TAL1 1 dataset
ChIP K562 ENCFF620GMX 291 bp overlap
TBP 1 dataset
ChIP H1 ENCFF859IIO 289 bp overlap
TBX2 1 dataset
ChIP Kelly GSE94822.TBX2.Kelly 64 bp overlap
TCF3 2 datasets
ChIP K562 ENCFF319QZT 304 bp overlap
ChIP NPC GSE154479.TCF3.NPC 240 bp overlap
TEAD1 4 datasets
ChIP HEK293_siE1A GSE130135.TEAD1.HEK293_siE1A 131 bp overlap
ChIP HepG2 ENCFF661PNM 205 bp overlap
ChIP MSTO GSE68170.TEAD1.MSTO 220 bp overlap
ChIP pancreas_12D ERP008682.TEAD1.pancreas_12D 166 bp overlap
TEAD3 1 dataset
ChIP HepG2 ENCFF054UUL 189 bp overlap
TEAD4 13 datasets
ChIP ESC S26-ESC-d0-TEAD4-exp1 337 bp overlap
ChIP H1 ENCFF778PAX 141 bp overlap
ChIP HCT-116 ENCSR000BVJ.TEAD4.HCT-116 109 bp overlap
ChIP HepG2 ENCFF006QNB 304 bp overlap
ChIP HepG2 ENCFF250NXO 199 bp overlap
ChIP Ishikawa ENCFF772OTG 78 bp overlap
ChIP Ishikawa ENCSR000BSW.TEAD4.Ishikawa 135 bp overlap
ChIP K562 ENCFF673NIK 298 bp overlap
ChIP MCF-7 ENCFF710WPA 251 bp overlap
ChIP SK-MEL-147 GSE94488.TEAD4.SK-MEL-147 142 bp overlap
ChIP SNU-216 GSE44416.TEAD4.SNU-216 207 bp overlap
ChIP T-47D_Veh GSE125594.TEAD4.T-47D_Veh 204 bp overlap
ChIP hESC GSE99202.TEAD4.hESC 177 bp overlap
TEF 1 dataset
ChIP HepG2 ENCFF661AUQ 273 bp overlap
TFAP2A 1 dataset
ChIP WA09 GSE105081.TFAP2A.WA09 110 bp overlap
TFAP2C 3 datasets
ChIP UCLA1-hESCs_TFAP2C--_1d_TFAP2C_Induc GSE101074.TFAP2C.UCLA1-hESCs_TFAP2C--_1d_TFAP2C_Induc 357 bp overlap
ChIP UCLA1-hESCs_TFAP2C--_2d_TFAP2C_Induc GSE101074.TFAP2C.UCLA1-hESCs_TFAP2C--_2d_TFAP2C_Induc 251 bp overlap
ChIP WA09 GSE105081.TFAP2C.WA09 97 bp overlap
TFAP4 3 datasets
ChIP Hep-G2 ENCSR103SZL.TFAP4.Hep-G2 146 bp overlap
ChIP HepG2 ENCFF030SRU 252 bp overlap
ChIP HepG2 ENCFF932XOY 154 bp overlap
TFE3 1 dataset
ChIP Hep-G2 ENCSR589SNT.TFE3.Hep-G2 135 bp overlap
THAP11 1 dataset
ChIP HepG2 ENCFF272SWH 362 bp overlap
TRIM28 2 datasets
ChIP HEK293 ENCFF265CEM 354 bp overlap
ChIP HEK293 ENCFF582MWI 419 bp overlap
WT1 2 datasets
ChIP HEK293 ENCFF906HIR 191 bp overlap
ChIP HEK293 ENCSR966PJJ.WT1.HEK293 177 bp overlap
YY1 1 dataset
ChIP Hep-G2_RBM25-KD GSE120104.YY1.Hep-G2_RBM25-KD 360 bp overlap
YY1AP1 1 dataset
ChIP MCF-10A_DOX GSE115787.YY1AP1.MCF-10A_DOX 161 bp overlap
ZBTB17 1 dataset
ChIP HEK293 ENCFF865LIO 320 bp overlap
ZBTB44 2 datasets
ChIP HEK293 ENCFF560VPN 187 bp overlap
ChIP HEK293 ENCSR076STQ.ZBTB44.HEK293 229 bp overlap
ZEB2 7 datasets
ChIP HEK293 ENCFF847JIE 304 bp overlap
ChIP HEK293 ENCSR417VWF.ZEB2.HEK293 330 bp overlap
ChIP K-562 ENCSR322CFO.ZEB2.K-562 334 bp overlap
ChIP K-562 ENCSR004GKA.ZEB2.K-562 331 bp overlap
ChIP K562 ENCFF795CMH 343 bp overlap
ChIP K562 ENCFF975RXS 362 bp overlap
ChIP K562 ENCFF975RXS 162 bp overlap
ZNF2 1 dataset
ChIP HEK293 ENCSR011CKE.ZNF2.HEK293 242 bp overlap
ZNF281 1 dataset
ChIP K562 ENCFF594VNM 262 bp overlap
ZNF362 2 datasets
ChIP HEK293 ENCFF436CGE 66 bp overlap
ChIP HEK293 ENCSR715QNO.ZNF362.HEK293 64 bp overlap
ZNF384 4 datasets
ChIP HEK293T ENCFF019DZX 91 bp overlap
ChIP HEK293T ENCSR882ICT.ZNF384.HEK293T 85 bp overlap
ChIP Hep-G2 ENCSR101FJU.ZNF384.Hep-G2 81 bp overlap
ChIP K-562 ENCSR000EFP.ZNF384.K-562 118 bp overlap
ZNF48 1 dataset
ChIP HepG2 ENCFF362CDQ 376 bp overlap
ZNF600 1 dataset
ChIP HEK293 ENCFF785JSX 183 bp overlap
ZNF644 1 dataset
ChIP HepG2 ENCFF352VGJ 280 bp overlap
ZNF770 3 datasets
ChIP HEK293 ENCSR242BGR.ZNF770.HEK293 298 bp overlap
ChIP HEK293 GSE76494.ZNF770.HEK293 144 bp overlap
ChIP Hep-G2 ENCSR510GKB.ZNF770.Hep-G2 179 bp overlap