chr9 : 21,989,523 21,990,203
680 bp 120 TFs 2 linked genes
This 680 bp open chromatin element is linked to CDKN2B-AS1 and ENSG00000266446 and is bound by 120 transcription factors.
Linked Genes
2 genes
Gene Expression Dist. to TSS Distance Link type
CDKN2B-AS1 4.3 kb Proximal Proximity
ENSG00000266446 5.3 kb Proximal Proximity
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr9:21,984,523 – 21,995,203
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
120 transcription factors
Source
Cell type
AR 2 datasets
ChIP LNCaP GSE80256.AR.LNCaP 167 bp overlap
ChIP prostate_fetal-fibroblast GSE90772.AR.prostate_fetal-fibroblast 287 bp overlap
ASH2L 2 datasets
ChIP H1 ENCFF399KAM 680 bp overlap
ChIP WA01 ENCSR850KIP.ASH2L.WA01 680 bp overlap
ATF2 1 dataset
ChIP HEK293 ENCFF194VKZ 72 bp overlap
BACH1 2 datasets
ChIP WA01 ENCSR000EBQ.BACH1.WA01 130 bp overlap
ChIP WA01 ENCSR000EBQ.BACH1.WA01 159 bp overlap
BCL11A 3 datasets
ChIP HUDEP-2_30min GSE104676.BCL11A.HUDEP-2_30min 110 bp overlap
ChIP HUDEP-2_90min GSE104676.BCL11A.HUDEP-2_90min 68 bp overlap
ChIP HUDEP-2_90min GSE104676.BCL11A.HUDEP-2_90min 124 bp overlap
BCL6 1 dataset
ChIP OCI-Ly1 GSE29282.BCL6.OCI-Ly1 326 bp overlap
BCOR 4 datasets
ChIP WA01 GSE104690.BCOR.WA01 680 bp overlap
ChIP WA01_KDM2B_KO GSE104690.BCOR.WA01_KDM2B_KO 523 bp overlap
ChIP WA01_RNF2-R GSE104690.BCOR.WA01_RNF2-R 649 bp overlap
ChIP WA01_RNF2-R_Doxycyclin GSE104690.BCOR.WA01_RNF2-R_Doxycyclin 680 bp overlap
BRD1 1 dataset
ChIP HUES-64 GSE104059.BRD1.HUES-64 568 bp overlap
BRD4 9 datasets
ChIP CLL_patient1_4h_CpG GSE109411.BRD4.CLL_patient1_4h_CpG 302 bp overlap
ChIP CLL_patient1_4h_CpG GSE109411.BRD4.CLL_patient1_4h_CpG 168 bp overlap
ChIP CLL_patient2 GSE109411.BRD4.CLL_patient2 202 bp overlap
ChIP CLL_patient2 GSE109411.BRD4.CLL_patient2 296 bp overlap
ChIP CLL_patient2_4h_CpG_BET GSE109411.BRD4.CLL_patient2_4h_CpG_BET 170 bp overlap
ChIP CLL_patient4_4h_CpG GSE109411.BRD4.CLL_patient4_4h_CpG 320 bp overlap
ChIP HAP1 GSE108387.BRD4.HAP1 499 bp overlap
ChIP HEK293_Ctrl GSE129407.BRD4.HEK293_Ctrl 598 bp overlap
ChIP HEK293_sgNT GSE129407.BRD4.HEK293_sgNT 401 bp overlap
CBX7 1 dataset
ChIP lymphocyte_DMSO GSE110139.CBX7.lymphocyte_DMSO 630 bp overlap
CHD1 1 dataset
ChIP WA01 ENCSR000AQK.CHD1.WA01 548 bp overlap
CHD8 1 dataset
ChIP T-47D_ETOH_45 GSE62428.CHD8.T-47D_ETOH_45 155 bp overlap
CREB1 3 datasets
ChIP GM23338 ENCFF432ZEW 54 bp overlap
ChIP Hep-G2 ENCSR112ALD.CREB1.Hep-G2 93 bp overlap
ChIP WA01 ENCSR000BSN.CREB1.WA01 108 bp overlap
CTBP2 2 datasets
ChIP H1 ENCFF329MAX 445 bp overlap
ChIP WA01 ENCSR000EUO.CTBP2.WA01 645 bp overlap
CTNNB1 1 dataset
ChIP hiPSC_D2 GSE132532.CTNNB1.hiPSC_D2 188 bp overlap
CXXC4 1 dataset
ChIP HEK293T GSE42958.CXXC4.HEK293T 96 bp overlap
E2F6 2 datasets
ChIP H1 ENCFF785DWK 216 bp overlap
ChIP WA01 ENCSR000BSI.E2F6.WA01 134 bp overlap
EED 2 datasets
ChIP ProEs GSE59087.EED.ProEs 473 bp overlap
ChIP ProEs GSE59087.EED.ProEs 131 bp overlap
EGR1 1 dataset
ChIP H1 ENCFF451BLH 224 bp overlap
EGR2 1 dataset
ChIP HEK293 ENCFF336LFH 311 bp overlap
ESR1 2 datasets
ChIP Ishikawa_ETV4-KO2_Mili GSE129803.ESR1.Ishikawa_ETV4-KO2_Mili 299 bp overlap
ChIP Ishikawa_siNTC-96h GSE129803.ESR1.Ishikawa_siNTC-96h 229 bp overlap
ETS1 1 dataset
ChIP cardiomyocyte_D0 GSE129986.ETS1.cardiomyocyte_D0 199 bp overlap
EZH2 14 datasets
ChIP GM23338 ENCFF613YON 93 bp overlap
ChIP GM23338 ENCFF613YON 212 bp overlap
ChIP GM23338 ENCFF886DXX 357 bp overlap
ChIP H1 ENCFF232NZA 680 bp overlap
ChIP SU-DHL-5 GSE45982.EZH2.SU-DHL-5 76 bp overlap
ChIP endothelial_umbilical-vein ENCSR000ATA.EZH2.endothelial_umbilical-vein 680 bp overlap
ChIP hESC GSE13084.EZH2.hESC 276 bp overlap
ChIP hESC GSE113817.EZH2.hESC 294 bp overlap
ChIP hepatocyte ENCFF118DKH 357 bp overlap
ChIP hepatocyte ENCFF118DKH 348 bp overlap
ChIP hepatocyte ENCFF552DZB 535 bp overlap
ChIP neural ENCSR511CUH.EZH2.neural 467 bp overlap
ChIP neural progenitor cell ENCFF018MKA 663 bp overlap
ChIP neural progenitor cell ENCFF472NFV 680 bp overlap
FEZF1 1 dataset
ChIP HEK293 ENCFF528YED 62 bp overlap
FOXA1 1 dataset
ChIP PDAC_SHCTR GSE64557.FOXA1.PDAC_SHCTR 69 bp overlap
GATA6 4 datasets
ChIP DE_D1 S14-DE-d1-GATA6-exp1 392 bp overlap
ChIP DE_D1 S41-DE-d1-GATA6-exp2 499 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 573 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 555 bp overlap
GLIS1 2 datasets
ChIP HEK293 ENCFF299RSE 232 bp overlap
ChIP HEK293 ENCSR482BBZ.GLIS1.HEK293 563 bp overlap
GLIS2 2 datasets
ChIP HEK293 ENCFF446EIF 278 bp overlap
ChIP HEK293 ENCSR535DIA.GLIS2.HEK293 483 bp overlap
GTF3C2 1 dataset
ChIP H9 GSE94418.GTF3C2.H9 443 bp overlap
HDAC2 3 datasets
ChIP H1 ENCFF353UJQ 634 bp overlap
ChIP WA01 ENCSR000AVB.HDAC2.WA01 303 bp overlap
ChIP WA01 ENCSR000AVB.HDAC2.WA01 78 bp overlap
HIF1A 1 dataset
ChIP BEAS-2B_arsenic GSE145834.HIF1A.BEAS-2B_arsenic 409 bp overlap
HMGB2 1 dataset
ChIP HUVEC-C GSE98245.HMGB2.HUVEC-C 316 bp overlap
HNF4A 1 dataset
ChIP hiPSC GSE104613.HNF4A.hiPSC 361 bp overlap
IFNA1 1 dataset
ChIP Huh-7_GFP GSE110511.IFNA1.Huh-7_GFP 183 bp overlap
IKZF3 1 dataset
ChIP HEK293 ENCSR304AMN.IKZF3.HEK293 366 bp overlap
IRF2 1 dataset
ChIP keratinocyte_DOX GSE135677.IRF2.keratinocyte_DOX 147 bp overlap
JARID2 12 datasets
ChIP AMIPS6 GSE48516.JARID2.AMIPS6 532 bp overlap
ChIP AMIPS8 GSE48516.JARID2.AMIPS8 125 bp overlap
ChIP EDOMIPS2 GSE48516.JARID2.EDOMIPS2 382 bp overlap
ChIP EDOMIPS2 GSE48516.JARID2.EDOMIPS2 220 bp overlap
ChIP MRC-5_IPS25 GSE48516.JARID2.MRC-5_IPS25 231 bp overlap
ChIP MRC-5_IPS25_CTR GSE48515.JARID2.MRC-5_IPS25_CTR 680 bp overlap
ChIP MRC-5_IPS25_MEG3 GSE48515.JARID2.MRC-5_IPS25_MEG3 680 bp overlap
ChIP UTEIPS11 GSE48516.JARID2.UTEIPS11 433 bp overlap
ChIP UTEIPS11 GSE48516.JARID2.UTEIPS11 73 bp overlap
ChIP UTEIPS4 GSE48516.JARID2.UTEIPS4 656 bp overlap
ChIP UTEIPS6 GSE48516.JARID2.UTEIPS6 680 bp overlap
ChIP UTEIPS7 GSE48516.JARID2.UTEIPS7 680 bp overlap
JUN 7 datasets
ChIP DE_D1 S40-DE-d1-JUN-exp2 287 bp overlap
ChIP DE_D2 S02-DE-d2-JUN-exp1 272 bp overlap
ChIP ESC S24-ESC-d0-JUN-exp1 383 bp overlap
ChIP ESC S24-ESC-d0-JUN-exp1 192 bp overlap
ChIP ESC S34-ESC-d0-JUN-exp2 392 bp overlap
ChIP HUES-8 GSE109524.JUN.HUES-8 680 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.JUN.HUES-8_DE_ctrl 431 bp overlap
KDM4A 3 datasets
ChIP H1 ENCFF078LED 613 bp overlap
ChIP H1 ENCFF078LED 563 bp overlap
ChIP WA01 ENCSR000AVC.KDM4A.WA01 595 bp overlap
KLF17 1 dataset
ChIP HEK293 ENCSR065WUF.KLF17.HEK293 260 bp overlap
KLF8 2 datasets
ChIP HEK293 ENCFF929IAJ 260 bp overlap
ChIP HEK293 ENCSR635NOQ.KLF8.HEK293 257 bp overlap
KLF9 1 dataset
ChIP GBM1A GSE62211.KLF9.GBM1A 176 bp overlap
KMT2A 4 datasets
ChIP MV4-11 GSE83671.KMT2A.MV4-11 269 bp overlap
ChIP MV4-11 GSE83671.KMT2A.MV4-11 290 bp overlap
ChIP RCH-ACV GSE83671.KMT2A.RCH-ACV 640 bp overlap
ChIP RS4-11_DMSO-D3-180402 GSE127507.KMT2A.RS4-11_DMSO-D3-180402 363 bp overlap
MAX 2 datasets
ChIP H1 ENCFF914VQY 295 bp overlap
ChIP melanocyte GSE115845.MAX.melanocyte 153 bp overlap
MAZ 4 datasets
ChIP HEK293 ENCFF994GSG 202 bp overlap
ChIP HEK293 ENCFF994GSG 176 bp overlap
ChIP HEK293 ENCSR290SSQ.MAZ.HEK293 347 bp overlap
ChIP HEK293 GSE76494.MAZ.HEK293 196 bp overlap
MEN1 1 dataset
ChIP RS4-11_DMSO-D3-180110 GSE127507.MEN1.RS4-11_DMSO-D3-180110 680 bp overlap
MYC 1 dataset
ChIP SK-N-SH GSE138295.MYC.SK-N-SH 132 bp overlap
NANOG 1 dataset
ChIP HUES-8 GSE109524.NANOG.HUES-8 680 bp overlap
NCAPH2 1 dataset
ChIP IMR-90_Bethyl275_OIS GSE118494.NCAPH2.IMR-90_Bethyl275_OIS 205 bp overlap
OGG1 1 dataset
ChIP HEK293 GSE89017.OGG1.HEK293 345 bp overlap
PATZ1 2 datasets
ChIP HEK293 ENCFF016MNJ 288 bp overlap
ChIP HEK293 ENCSR966ULI.PATZ1.HEK293 608 bp overlap
PHF8 1 dataset
ChIP WA01 ENCSR000ATK.PHF8.WA01 134 bp overlap
POLR2A 1 dataset
ChIP HeLa-S3 ENCFF045HUU 526 bp overlap
POU5F1 4 datasets
ChIP BG03 GSE21614.POU5F1.BG03 222 bp overlap
ChIP DE_D1 DED1-OCT4_Batch_II 501 bp overlap
ChIP HUES-8 GSE109524.POU5F1.HUES-8 680 bp overlap
ChIP hESC ENCSR264RJX.POU5F1.hESC 113 bp overlap
POU5F1_M 1 dataset
ChIP DE_D1 DED1-OCT4-M_Batch_II 445 bp overlap
PRDM1 1 dataset
ChIP HEK293 ENCFF302TBP 257 bp overlap
PRDM10 1 dataset
ChIP HEK293 ENCFF145WQQ 399 bp overlap
RAD21 1 dataset
ChIP HEK293_siYAP GSE130135.RAD21.HEK293_siYAP 313 bp overlap
RBBP5 1 dataset
ChIP WA01 ENCSR000AQC.RBBP5.WA01 427 bp overlap
RNF2 1 dataset
ChIP H1 ENCFF239FFS 680 bp overlap
RYBP 1 dataset
ChIP WA01 GSE104690.RYBP.WA01 680 bp overlap
SETDB1 2 datasets
ChIP HEK293 ENCSR348AGV.SETDB1.HEK293 436 bp overlap
ChIP U2OS ENCSR000EYD.SETDB1.U2OS 427 bp overlap
SIN3A 1 dataset
ChIP WA01 ENCSR000EBO.SIN3A.WA01 375 bp overlap
SMAD3 2 datasets
ChIP H69_4TGF GSE145250.SMAD3.H69_4TGF 116 bp overlap
ChIP H69_4TGF GSE145250.SMAD3.H69_4TGF 296 bp overlap
SP1 2 datasets
ChIP H1 ENCFF263FUH 215 bp overlap
ChIP WA01 ENCSR000BIR.SP1.WA01 193 bp overlap
SP2 3 datasets
ChIP HEK293 ENCFF181QXT 205 bp overlap
ChIP HEK293 ENCSR807LQP.SP2.HEK293 284 bp overlap
ChIP HEK293 GSE76494.SP2.HEK293 186 bp overlap
SP3 1 dataset
ChIP HEK293 ENCSR141PZA.SP3.HEK293 185 bp overlap
SP7 2 datasets
ChIP HEK293 ENCFF733RBE 359 bp overlap
ChIP HEK293 ENCSR468IJT.SP7.HEK293 448 bp overlap
SREBP2 1 dataset
ChIP HCC70 GSE126380.SREBP2.HCC70 261 bp overlap
STAT3 1 dataset
ChIP monocyte_IFNg-LPS GSE120943.STAT3.monocyte_IFNg-LPS 251 bp overlap
SUZ12 5 datasets
ChIP H1 ENCFF507HGF 271 bp overlap
ChIP H1 ENCFF881NFR 680 bp overlap
ChIP NT2-D1 GSE101538.SUZ12.NT2-D1 545 bp overlap
ChIP WA01 ENCSR000ATS.SUZ12.WA01 680 bp overlap
ChIP WA01 ENCSR000EUQ.SUZ12.WA01 246 bp overlap
TBP 1 dataset
ChIP hiPSC_WTb_RNase-neg GSE128135.TBP.hiPSC_WTb_RNase-neg 243 bp overlap
TCF12 1 dataset
ChIP WA01 ENCSR000BIT.TCF12.WA01 177 bp overlap
TRIM28 1 dataset
ChIP AF22 GSE84259.TRIM28.AF22 221 bp overlap
TSHZ1 1 dataset
ChIP HEK293 ENCSR217WRC.TSHZ1.HEK293 258 bp overlap
YY1 2 datasets
ChIP H1 ENCFF524BTL 231 bp overlap
ChIP WA01 ENCSR000BKD.YY1.WA01 164 bp overlap
ZBTB10 2 datasets
ChIP HEK293 ENCFF679BCK 380 bp overlap
ChIP HEK293 ENCSR004PLU.ZBTB10.HEK293 388 bp overlap
ZBTB17 1 dataset
ChIP HEK293 ENCFF865LIO 671 bp overlap
ZBTB20 2 datasets
ChIP HEK293 ENCFF524ADK 235 bp overlap
ChIP HEK293 ENCSR460MBI.ZBTB20.HEK293 544 bp overlap
ZBTB21 2 datasets
ChIP HEK293 ENCFF509WYZ 111 bp overlap
ChIP HEK293 ENCSR321MSF.ZBTB21.HEK293 86 bp overlap
ZBTB26 4 datasets
ChIP HEK293 ENCFF752POA 379 bp overlap
ChIP HEK293 ENCFF752TCU 385 bp overlap
ChIP HEK293 ENCFF752TCU 345 bp overlap
ChIP HEK293 ENCSR229DYF.ZBTB26.HEK293 460 bp overlap
ZBTB48 1 dataset
ChIP U2OS_ZBTB48-KO GSE96776.ZBTB48.U2OS_ZBTB48-KO 507 bp overlap
ZBTB6 1 dataset
ChIP HEK293 GSE76494.ZBTB6.HEK293 390 bp overlap
ZBTB8A 2 datasets
ChIP HEK293 ENCFF303WRD 249 bp overlap
ChIP HEK293 ENCSR481FEC.ZBTB8A.HEK293 436 bp overlap
ZEB2 1 dataset
ChIP HEK293 ENCSR417VWF.ZEB2.HEK293 431 bp overlap
ZFHX2 1 dataset
ChIP HEK293 ENCFF167TUA 230 bp overlap
ZFP64 1 dataset
ChIP HEK293 GSE76494.ZFP64.HEK293 173 bp overlap
ZMYND8 1 dataset
ChIP HEK293_Flag-ZMYND8 GSE81696.ZMYND8.HEK293_Flag-ZMYND8 314 bp overlap
ZNF143 1 dataset
ChIP HeLa GSE39263.ZNF143.HeLa 204 bp overlap
ZNF189 2 datasets
ChIP HEK293 ENCFF638TIB 216 bp overlap
ChIP HEK293 ENCSR163RYW.ZNF189.HEK293 680 bp overlap
ZNF2 1 dataset
ChIP HEK293 ENCSR011CKE.ZNF2.HEK293 452 bp overlap
ZNF24 1 dataset
ChIP HEK293 ENCSR984MDV.ZNF24.HEK293 295 bp overlap
ZNF263 1 dataset
ChIP HEK293 ENCFF336CWQ 481 bp overlap
ZNF28 1 dataset
ChIP HEK293T GSE78099.ZNF28.HEK293T 182 bp overlap
ZNF320 1 dataset
ChIP HEK293 GSE76494.ZNF320.HEK293 246 bp overlap
ZNF333 1 dataset
ChIP HEK293T GSE78099.ZNF333.HEK293T 235 bp overlap
ZNF335 3 datasets
ChIP HEK293 ENCFF784SLD 671 bp overlap
ChIP HEK293 ENCFF784SLD 181 bp overlap
ChIP HEK293 ENCSR328SUD.ZNF335.HEK293 580 bp overlap
ZNF341 3 datasets
ChIP HEK293 ENCFF944VMC 271 bp overlap
ChIP HEK293 ENCFF944VMC 68 bp overlap
ChIP HEK293 ENCSR185FOY.ZNF341.HEK293 203 bp overlap
ZNF398 2 datasets
ChIP HEK293 ENCFF184XEW 271 bp overlap
ChIP HEK293 ENCSR676ZEF.ZNF398.HEK293 459 bp overlap
ZNF417 2 datasets
ChIP HEK293T GSE78099.ZNF417.HEK293T 261 bp overlap
ChIP WA01 GSE144149.ZNF417.WA01 137 bp overlap
ZNF445 1 dataset
ChIP HEK293T GSE78099.ZNF445.HEK293T 340 bp overlap
ZNF574 1 dataset
ChIP HEK293 GSE76494.ZNF574.HEK293 146 bp overlap
ZNF587 1 dataset
ChIP HEK293T GSE78099.ZNF587.HEK293T 147 bp overlap
ZNF600 2 datasets
ChIP HEK293 ENCFF785JSX 393 bp overlap
ChIP HEK293 ENCFF785JSX 169 bp overlap
ZNF623 2 datasets
ChIP HEK293 ENCFF505YHP 151 bp overlap
ChIP HEK293 ENCSR022IZK.ZNF623.HEK293 564 bp overlap
ZNF629 2 datasets
ChIP HEK293 ENCFF096ELQ 105 bp overlap
ChIP HEK293 ENCSR351NON.ZNF629.HEK293 514 bp overlap
ZNF692 2 datasets
ChIP HEK293 ENCSR418MKG.ZNF692.HEK293 343 bp overlap
ChIP HEK293 ENCSR418MKG.ZNF692.HEK293 163 bp overlap
ZNF711 1 dataset
ChIP HEK293T GSE145160.ZNF711.HEK293T 680 bp overlap
ZNF76 1 dataset
ChIP HEK293 ENCSR072LQF.ZNF76.HEK293 389 bp overlap
ZXDB 1 dataset
ChIP HEK293 ENCSR559IOZ.ZXDB.HEK293 540 bp overlap