chr7 : 30,984,342 30,985,023
681 bp 115 TFs 2 linked genes
This 681 bp open chromatin element is linked to GHRHR and MINDY4 and is bound by 115 transcription factors.
Linked Genes
2 genes
Distance
Link type
Gene Expression Dist. to TSS Distance Link type
GHRHR 262 bp At TSS Proximity
MINDY4 213.5 kb Distal Multiome
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr7:30,979,342 – 30,990,023
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
115 transcription factors
Source
Cell type
AR 1 dataset
ChIP A-375_SLNCR GSE116189.AR.A-375_SLNCR 219 bp overlap
ASH2L 3 datasets
ChIP H1 ENCFF399KAM 681 bp overlap
ChIP H1 ENCFF399KAM 525 bp overlap
ChIP WA01 ENCSR850KIP.ASH2L.WA01 661 bp overlap
ATF2 1 dataset
ChIP WA01 ENCSR000BQU.ATF2.WA01 140 bp overlap
ATF3 1 dataset
ChIP HCT-116_CAMP GSE74355.ATF3.HCT-116_CAMP 175 bp overlap
BCOR 4 datasets
ChIP WA01 GSE104690.BCOR.WA01 653 bp overlap
ChIP WA01_RNF2-R GSE104690.BCOR.WA01_RNF2-R 568 bp overlap
ChIP WA01_RNF2-R_Doxycyclin GSE104690.BCOR.WA01_RNF2-R_Doxycyclin 503 bp overlap
ChIP WA01_RNF2-R_Doxycyclin GSE104690.BCOR.WA01_RNF2-R_Doxycyclin 108 bp overlap
BRD1 1 dataset
ChIP HUES-64 GSE104059.BRD1.HUES-64 356 bp overlap
CBFA2T2 2 datasets
ChIP NCCIT GSE71675.CBFA2T2.NCCIT 198 bp overlap
ChIP NCCIT GSE71675.CBFA2T2.NCCIT 90 bp overlap
CDKN1B 1 dataset
ChIP MDA-MB-231_p27CK-DD GSE112444.CDKN1B.MDA-MB-231_p27CK-DD 258 bp overlap
CHD1 1 dataset
ChIP WA01 ENCSR000AQK.CHD1.WA01 347 bp overlap
CHD2 1 dataset
ChIP WA01 ENCSR000EBT.CHD2.WA01 136 bp overlap
CHD7 1 dataset
ChIP WA01 ENCSR000AVA.CHD7.WA01 157 bp overlap
CREB1 1 dataset
ChIP GM23338 ENCSR214ZAV.CREB1.GM23338 168 bp overlap
CTBP2 1 dataset
ChIP H1 ENCFF329MAX 591 bp overlap
CTCF 8 datasets
ChIP B-cell ENCSR000AUV.CTCF.B-cell 80 bp overlap
ChIP CD14 ENCSR000ATN.CTCF.CD14 208 bp overlap
ChIP T-47D_Y537S GSE148277.CTCF.T-47D_Y537S 395 bp overlap
ChIP adrenal-gland ENCSR770IWO.CTCF.adrenal-gland 122 bp overlap
ChIP erythroid GSE67783.CTCF.erythroid 302 bp overlap
ChIP lymphoblast_mut GSE155324.CTCF.lymphoblast_mut 232 bp overlap
ChIP nerve_tibial ENCSR434XLP.CTCF.nerve_tibial 67 bp overlap
ChIP prostate_epithelial ENCSR196HOM.CTCF.prostate_epithelial 100 bp overlap
Cebpa 1 dataset
ChIP BLaER1 ENCFF262VBH 414 bp overlap
E2F4 1 dataset
ChIP MCF-10A ENCSR000DOR.E2F4.MCF-10A 213 bp overlap
EBF1 1 dataset
Motif DE_12h DE_12h-EBF1_MA0154.5 11 bp overlap
EBF3 1 dataset
Motif DE_12h DE_12h-EBF3_MA1637.2 9 bp overlap
EP300 1 dataset
ChIP H1 ENCFF927IYK 270 bp overlap
ESR1 4 datasets
ChIP MCF-7_SIGATA_E2 GSE40129.ESR1.MCF-7_SIGATA_E2 140 bp overlap
ChIP T-47D_DMSO GSE148277.ESR1.T-47D_DMSO 238 bp overlap
ChIP U2OS_E2 GSE26110.ESR1.U2OS_E2 163 bp overlap
ChIP primary-endometrium-cancer_E1_DSG GSE114737.ESR1.primary-endometrium-cancer_E1_DSG 290 bp overlap
ETS1 5 datasets
ChIP HUVEC-C_12h GSE93030.ETS1.HUVEC-C_12h 164 bp overlap
ChIP HUVEC-C_VEGF_12H GSE41166.ETS1.HUVEC-C_VEGF_12H 174 bp overlap
ChIP HUVEC-C_VEGF_12h GSE109625.ETS1.HUVEC-C_VEGF_12h 164 bp overlap
ChIP HUVEC-C_VEGF_1H GSE41166.ETS1.HUVEC-C_VEGF_1H 186 bp overlap
ChIP HUVEC-C_VEGF_4H GSE41166.ETS1.HUVEC-C_VEGF_4H 319 bp overlap
EZH2 2 datasets
ChIP LNCaP-abl GSE39459.EZH2.LNCaP-abl 198 bp overlap
ChIP neural_progenitor ENCSR069DPL.EZH2.neural_progenitor 614 bp overlap
Ebf2 1 dataset
Motif DE_12h DE_12h-Ebf2_MA1604.2 9 bp overlap
Ebf4 1 dataset
Motif DE_12h DE_12h-Ebf4_MA2122.1 11 bp overlap
FEZF2 1 dataset
Motif DE_12h DE_12h-FEZF2_MA2341.1 8 bp overlap
FOS 4 datasets
ChIP CFPAC-1 GSE119930.FOS.CFPAC-1 241 bp overlap
ChIP CFPAC-1 GSE119930.FOS.CFPAC-1 266 bp overlap
ChIP HCT-116_A7A GSE152144.FOS.HCT-116_A7A 315 bp overlap
ChIP endothelial cell of umbilical vein ENCFF415XBG 302 bp overlap
FOSL2 2 datasets
ChIP MCF-7 ENCFF188KBZ 430 bp overlap
ChIP SK-N-SH ENCSR000BVB.FOSL2.SK-N-SH 112 bp overlap
FOXA1 7 datasets
ChIP CFPAC-1_FOXA2-KO GSE119930.FOXA1.CFPAC-1_FOXA2-KO 319 bp overlap
ChIP LNCaP_F266S_shFOXA1 GSE128883.FOXA1.LNCaP_F266S_shFOXA1 159 bp overlap
ChIP LNCaP_SHFOXA1_R1881 GSE37345.FOXA1.LNCaP_SHFOXA1_R1881 108 bp overlap
ChIP LS180 GSE140533.FOXA1.LS180 53 bp overlap
ChIP MCF-7 GSE72249.FOXA1.MCF-7 196 bp overlap
ChIP MCF-7_vehicle_ab2 GSE112969.FOXA1.MCF-7_vehicle_ab2 298 bp overlap
ChIP NCI-H3122 GSE39998.FOXA1.NCI-H3122 181 bp overlap
FOXA2 2 datasets
ChIP CFPAC-1_EV GSE119930.FOXA2.CFPAC-1_EV 259 bp overlap
ChIP KerCT GSE90454.FOXA2.KerCT 200 bp overlap
FOXK1 1 dataset
ChIP WTC11 ENCFF875IGU 365 bp overlap
FOXP1 1 dataset
ChIP H9 GSE31006.FOXP1.H9 162 bp overlap
GABPA 1 dataset
ChIP K-562 ENCSR000BLO.GABPA.K-562 133 bp overlap
HDAC2 2 datasets
ChIP WA01 ENCSR000AVB.HDAC2.WA01 440 bp overlap
ChIP WA01 ENCSR000AVB.HDAC2.WA01 153 bp overlap
HSF1 3 datasets
ChIP MCF-7_hyperthermia-15min GSE137558.HSF1.MCF-7_hyperthermia-15min 217 bp overlap
ChIP U2OS_HEAT GSE60984.HSF1.U2OS_HEAT 171 bp overlap
ChIP U2OS_HEAT_20 GSE60984.HSF1.U2OS_HEAT_20 198 bp overlap
IKZF1 1 dataset
Motif DE_12h DE_12h-IKZF1_MA1508.2 8 bp overlap
JUN 14 datasets
ChIP DE_D1 S08-DE-d1-JUN-exp1 493 bp overlap
ChIP DE_D1 S40-DE-d1-JUN-exp2 608 bp overlap
ChIP DE_D2 S02-DE-d2-JUN-exp1 523 bp overlap
ChIP DE_D2 S54-DE-d2-JUN-exp2 347 bp overlap
ChIP ESC S24-ESC-d0-JUN-exp1 681 bp overlap
ChIP ESC S34-ESC-d0-JUN-exp2 426 bp overlap
ChIP HAEC_IL1b_4h GSE89970.JUN.HAEC_IL1b_4h 179 bp overlap
ChIP HAEC_TNFa_4h GSE89970.JUN.HAEC_TNFa_4h 152 bp overlap
ChIP HUES-8 GSE109524.JUN.HUES-8 681 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.JUN.HUES-8_DE_ctrl 602 bp overlap
ChIP Karpas-299 GSE151413.JUN.Karpas-299 593 bp overlap
ChIP MCF-7_E2 GSE102410.JUN.MCF-7_E2 185 bp overlap
ChIP MCF-7_abemaciclib GSE157385.JUN.MCF-7_abemaciclib 277 bp overlap
ChIP MDA-BoM-1833 GSE112444.JUN.MDA-BoM-1833 347 bp overlap
JUNB 1 dataset
ChIP CFPAC-1 GSE119930.JUNB.CFPAC-1 479 bp overlap
JUND 3 datasets
ChIP H1 ENCFF468JZD 231 bp overlap
ChIP WA01 ENCSR000EBZ.JUND.WA01 220 bp overlap
ChIP WA01 ENCSR000BKP.JUND.WA01 136 bp overlap
KDM4A 5 datasets
ChIP H1 ENCFF078LED 172 bp overlap
ChIP H1 ENCFF078LED 502 bp overlap
ChIP WA01 ENCSR000AVC.KDM4A.WA01 323 bp overlap
ChIP hiPSC_IA12 GSE106870.KDM4A.hiPSC_IA12 165 bp overlap
ChIP hiPSC_IIA11 GSE106870.KDM4A.hiPSC_IIA11 391 bp overlap
KLF11 1 dataset
Motif DE_12h DE_12h-KLF11_MA1512.2 10 bp overlap
KLF16 1 dataset
Motif DE_12h DE_12h-KLF16_MA0741.1 11 bp overlap
KLF4 1 dataset
ChIP hiPSC GSE56567.KLF4.hiPSC 159 bp overlap
KLF6 1 dataset
Motif DE_12h DE_12h-KLF6_MA1517.2 9 bp overlap
KLF9 4 datasets
Motif DE_12h DE_12h-KLF9_MA1107.3 11 bp overlap
ChIP GBM1A GSE62211.KLF9.GBM1A 208 bp overlap
ChIP MCF-7 ENCFF618FCM 388 bp overlap
ChIP MCF-7 ENCSR125ZYC.KLF9.MCF-7 383 bp overlap
LIN54 2 datasets
Motif DE_12h DE_12h-LIN54_MA0619.2 7 bp overlap
Motif ES_0h ES_0h-LIN54_MA0619.2 7 bp overlap
MAX 2 datasets
ChIP NCI-H2171 GSE41105.MAX.NCI-H2171 128 bp overlap
ChIP WTC11 ENCFF223QFY 564 bp overlap
MED1 2 datasets
ChIP HCT-116_shLuc GSE121798.MED1.HCT-116_shLuc 192 bp overlap
ChIP HCT-116_thaps GSE121798.MED1.HCT-116_thaps 314 bp overlap
MED26 1 dataset
ChIP HCT-116_aux GSE121355.MED26.HCT-116_aux 303 bp overlap
MEIS1 1 dataset
Motif DE_12h DE_12h-MEIS1_MA0498.3 5 bp overlap
NANOG 4 datasets
ChIP GM23338 ENCFF065NZG 311 bp overlap
ChIP HUES-8 GSE109524.NANOG.HUES-8 427 bp overlap
ChIP WA01_3IL ERP004238.NANOG.WA01_3IL 196 bp overlap
ChIP WA09 GSE105028.NANOG.WA09 259 bp overlap
NELFE 1 dataset
ChIP HeLa_DOX GSE125534.NELFE.HeLa_DOX 131 bp overlap
NFE2L1 1 dataset
ChIP WTC11 ENCFF644BPU 377 bp overlap
NFIA 2 datasets
Motif DE_12h DE_12h-NFIA_MA0670.2 6 bp overlap
Motif ES_0h ES_0h-NFIA_MA0670.2 6 bp overlap
NFIX 2 datasets
Motif DE_12h DE_12h-NFIX_MA0671.2 6 bp overlap
Motif ES_0h ES_0h-NFIX_MA0671.2 6 bp overlap
OTX2 1 dataset
ChIP WTC11 ENCFF634NAO 245 bp overlap
POLR2A 2 datasets
ChIP H1 ENCFF566JSR 419 bp overlap
ChIP H1 ENCFF833NJP 291 bp overlap
POU2F3 2 datasets
Motif DE_12h DE_12h-POU2F3_MA0627.3 9 bp overlap
Motif ES_0h ES_0h-POU2F3_MA0627.3 9 bp overlap
POU5F1 8 datasets
ChIP DE_D1 DED1-OCT4_Batch_II 565 bp overlap
ChIP HUES-8 GSE109524.POU5F1.HUES-8 633 bp overlap
ChIP NCCIT_SNF5 GSE36134.POU5F1.NCCIT_SNF5 681 bp overlap
ChIP SKM-1 GSE93706.POU5F1.SKM-1 239 bp overlap
ChIP hESC ENCSR264RJX.POU5F1.hESC 136 bp overlap
ChIP hESC_PRIMED GSE69646.POU5F1.hESC_PRIMED 468 bp overlap
ChIP hiPSC GSE56567.POU5F1.hiPSC 158 bp overlap
ChIP hiPSC_3s2 GSE81899.POU5F1.hiPSC_3s2 168 bp overlap
POU5F1_M 1 dataset
ChIP DE_D1 DED1-OCT4-M_Batch_II 262 bp overlap
PROX1 1 dataset
ChIP SW480 GSE60390.PROX1.SW480 146 bp overlap
Pou5f1::Sox2 2 datasets
Motif DE_12h DE_12h-Pou5f1Sox2_MA0142.1 15 bp overlap
Motif ES_0h ES_0h-Pou5f1Sox2_MA0142.1 15 bp overlap
RAD21 2 datasets
ChIP HCT-116 GSE131606.RAD21.HCT-116 481 bp overlap
ChIP HeLa-S3_CTCF-s-OE GSE108869.RAD21.HeLa-S3_CTCF-s-OE 213 bp overlap
RBBP5 1 dataset
ChIP WA01 ENCSR000AQC.RBBP5.WA01 557 bp overlap
RELA 5 datasets
ChIP aortic-endothelial-cell_IL1B_D14 GSE139377.RELA.aortic-endothelial-cell_IL1B_D14 239 bp overlap
ChIP aortic-endothelial-cell_IL1B_D16 GSE139377.RELA.aortic-endothelial-cell_IL1B_D16 300 bp overlap
ChIP aortic-endothelial-cell_IL1B_D4 GSE139377.RELA.aortic-endothelial-cell_IL1B_D4 196 bp overlap
ChIP aortic-endothelial-cell_IL1B_D7 GSE139377.RELA.aortic-endothelial-cell_IL1B_D7 140 bp overlap
ChIP aortic-endothelial-cell_IL1B_D9 GSE139377.RELA.aortic-endothelial-cell_IL1B_D9 197 bp overlap
RNF2 1 dataset
ChIP WA01 ENCSR784VUY.RNF2.WA01 347 bp overlap
RREB1 1 dataset
Motif DE_12h DE_12h-RREB1_MA0073.2 19 bp overlap
RYBP 1 dataset
ChIP WA01 GSE104690.RYBP.WA01 468 bp overlap
SALL3 1 dataset
ChIP hiPSC GSE104863.SALL3.hiPSC 204 bp overlap
SAP30 1 dataset
ChIP WA01 ENCSR000ATR.SAP30.WA01 170 bp overlap
SIN3A 2 datasets
ChIP WA01 ENCSR000EBO.SIN3A.WA01 173 bp overlap
ChIP WA01 ENCSR000BIS.SIN3A.WA01 115 bp overlap
SIX4 1 dataset
ChIP WTC11 ENCFF891HYW 377 bp overlap
SMAD2-3 1 dataset
ChIP HUES-8_DE_ctrl GSE109524.SMAD2-3.HUES-8_DE_ctrl 404 bp overlap
SMAD3 2 datasets
ChIP BG03 GSE21614.SMAD3.BG03 190 bp overlap
ChIP BG03_DIFF_2H GSE36578.SMAD3.BG03_DIFF_2H 221 bp overlap
SMAD5 1 dataset
ChIP K-562 ENCSR000FCD.SMAD5.K-562 168 bp overlap
SMARCA4 5 datasets
ChIP A-549_AG15678 GSE132290.SMARCA4.A-549_AG15678 59 bp overlap
ChIP A-549_AG15721 GSE132290.SMARCA4.A-549_AG15721 63 bp overlap
ChIP hiPSC GSE124903.SMARCA4.hiPSC 178 bp overlap
ChIP hiPSC GSE124903.SMARCA4.hiPSC 358 bp overlap
ChIP hiPSC_WT GSE124903.SMARCA4.hiPSC_WT 393 bp overlap
SMARCB1 2 datasets
ChIP hiPSC GSE124903.SMARCB1.hiPSC 288 bp overlap
ChIP hiPSC_WT GSE124903.SMARCB1.hiPSC_WT 607 bp overlap
SMARCC1 4 datasets
ChIP ESC S25-ESC-d0-BAF155-exp1 381 bp overlap
ChIP hiPSC GSE124903.SMARCC1.hiPSC 163 bp overlap
ChIP hiPSC GSE124903.SMARCC1.hiPSC 278 bp overlap
ChIP hiPSC_WT GSE124903.SMARCC1.hiPSC_WT 406 bp overlap
SMC1 3 datasets
ChIP DKO GSE131606.SMC1.DKO 548 bp overlap
ChIP HCAEC GSE101921.SMC1.HCAEC 380 bp overlap
ChIP HCT-116 GSE131606.SMC1.HCT-116 389 bp overlap
SOX2 1 dataset
ChIP hiPSC GSE56567.SOX2.hiPSC 374 bp overlap
SP1 2 datasets
ChIP H1 ENCFF263FUH 270 bp overlap
ChIP WA01 ENCSR000BIR.SP1.WA01 130 bp overlap
SP3 1 dataset
Motif DE_12h DE_12h-SP3_MA0746.3 11 bp overlap
SP5 2 datasets
Motif DE_12h DE_12h-SP5_MA1965.2 6 bp overlap
Motif ES_0h ES_0h-SP5_MA1965.2 6 bp overlap
SP8 1 dataset
Motif DE_12h DE_12h-SP8_MA0747.2 11 bp overlap
STAT1 2 datasets
Motif DE_12h DE_12h-STAT1_MA0137.4 9 bp overlap
Motif ES_0h ES_0h-STAT1_MA0137.4 9 bp overlap
STAT3 4 datasets
ChIP HCC70 GSE152203.STAT3.HCC70 211 bp overlap
ChIP HCC70_EtOH GSE85579.STAT3.HCC70_EtOH 193 bp overlap
ChIP MCF-10A ENCSR000DOQ.STAT3.MCF-10A 168 bp overlap
ChIP MCF10A-Er-Src_TAM GSE115597.STAT3.MCF10A-Er-Src_TAM 163 bp overlap
SUZ12 1 dataset
ChIP CRL-7250_shSS18 GSE108025.SUZ12.CRL-7250_shSS18 189 bp overlap
TAF1 1 dataset
ChIP WA01 ENCSR000BHO.TAF1.WA01 199 bp overlap
TAF7 2 datasets
ChIP H1 ENCFF061XZZ 337 bp overlap
ChIP WA01 ENCSR000BLU.TAF7.WA01 264 bp overlap
TBP 6 datasets
ChIP H1 ENCFF859IIO 377 bp overlap
ChIP WA01 ENCSR000ECB.TBP.WA01 505 bp overlap
ChIP hESC GSE122298.TBP.hESC 305 bp overlap
ChIP hESC_10h GSE122298.TBP.hESC_10h 264 bp overlap
ChIP hESC_2h GSE122298.TBP.hESC_2h 246 bp overlap
ChIP hESC_8h GSE122298.TBP.hESC_8h 347 bp overlap
TEAD1 1 dataset
Motif DE_12h DE_12h-TEAD1_MA0090.4 9 bp overlap
TEAD2 1 dataset
Motif DE_12h DE_12h-TEAD2_MA1121.2 7 bp overlap
TEAD3 1 dataset
Motif DE_12h DE_12h-TEAD3_MA0808.1 8 bp overlap
TEAD4 4 datasets
ChIP BJ_fibroblast_CD13-pos GSE114367.TEAD4.BJ_fibroblast_CD13-pos 214 bp overlap
Motif DE_12h DE_12h-TEAD4_MA0809.3 8 bp overlap
ChIP H1 ENCFF778PAX 245 bp overlap
ChIP SK-MEL-147 GSE94488.TEAD4.SK-MEL-147 325 bp overlap
TFAP4 1 dataset
ChIP DLD-1 GSE46935.TFAP4.DLD-1 251 bp overlap
TP63 1 dataset
ChIP breast-organoid GSE113909.TP63.breast-organoid 116 bp overlap
TRIM28 1 dataset
ChIP hESC GSE115387.TRIM28.hESC 284 bp overlap
YY1 2 datasets
ChIP H1 ENCFF524BTL 285 bp overlap
ChIP WA01 ENCSR000BKD.YY1.WA01 187 bp overlap
ZBTB26 2 datasets
Motif DE_12h DE_12h-ZBTB26_MA1579.2 8 bp overlap
Motif ES_0h ES_0h-ZBTB26_MA1579.2 8 bp overlap
ZFP36 2 datasets
ChIP K-562 ENCSR776CYN.ZFP36.K-562 271 bp overlap
ChIP K562 ENCFF255RZG 243 bp overlap
ZNF143 2 datasets
ChIP WA01 ENCSR000EBW.ZNF143.WA01 237 bp overlap
ChIP WTC11 ENCFF249JUK 370 bp overlap
ZNF184 2 datasets
ChIP WTC11 ENCFF352POG 497 bp overlap
ChIP WTC11 ENCFF352POG 267 bp overlap
ZNF213 1 dataset
Motif DE_12h DE_12h-ZNF213_MA2121.1 12 bp overlap
ZNF214 1 dataset
Motif DE_12h DE_12h-ZNF214_MA1975.2 13 bp overlap
ZNF324 1 dataset
Motif DE_12h DE_12h-ZNF324_MA1977.2 14 bp overlap
ZNF382 1 dataset
Motif DE_12h DE_12h-ZNF382_MA1594.1 24 bp overlap
ZNF436 1 dataset
ChIP HEK293 GSE76494.ZNF436.HEK293 236 bp overlap
ZNF462 2 datasets
ChIP GM23338 ENCSR334UWP.ZNF462.GM23338 179 bp overlap
ChIP GM23338 ENCSR334UWP.ZNF462.GM23338 317 bp overlap
ZNF528 2 datasets
Motif DE_12h DE_12h-ZNF528_MA1597.1 17 bp overlap
Motif ES_0h ES_0h-ZNF528_MA1597.1 17 bp overlap
ZNF682 2 datasets
Motif DE_12h DE_12h-ZNF682_MA1599.2 11 bp overlap
Motif ES_0h ES_0h-ZNF682_MA1599.2 11 bp overlap
ZNF692 1 dataset
Motif DE_12h DE_12h-ZNF692_MA1986.2 8 bp overlap
ZNF70 1 dataset
ChIP SK-N-SH ENCFF833ACX 63 bp overlap
ZNF92 1 dataset
ChIP retina_pigment GSE60024.ZNF92.retina_pigment 155 bp overlap
Zfp335 1 dataset
Motif DE_12h DE_12h-Zfp335_MA2002.2 7 bp overlap