chr8 : 128,982,795 128,983,351
556 bp 116 TFs 1 linked gene
This 556 bp open chromatin element is linked to CCDC26 and is bound by 116 transcription factors.
Linked Genes
1 gene
Gene Expression Dist. to TSS Distance Link type
CCDC26 699.1 kb Distal Multiome+HiCAR
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr8:128,977,795 – 128,988,351
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
116 transcription factors
Source
Cell type
ARNTL 1 dataset
ChIP U2OS_DMOG GSE85096.ARNTL.U2OS_DMOG 209 bp overlap
ASCL1 2 datasets
Motif ES_0h ES_0h-ASCL1_MA1100.3 8 bp overlap
Motif ES_0h ES_0h-ASCL1_MA1631.2 9 bp overlap
ASH2L 3 datasets
ChIP H1 ENCFF399KAM 468 bp overlap
ChIP H1 ENCFF399KAM 556 bp overlap
ChIP WA01 ENCSR850KIP.ASH2L.WA01 469 bp overlap
ATF2 2 datasets
ChIP H1 ENCFF295GZO 477 bp overlap
ChIP WA01 ENCSR000BQU.ATF2.WA01 556 bp overlap
Ascl2 1 dataset
Motif ES_0h ES_0h-Ascl2_MA0816.1 10 bp overlap
BCL11A 2 datasets
ChIP WA01 ENCSR000BIP.BCL11A.WA01 127 bp overlap
ChIP WA01 ENCSR000BMJ.BCL11A.WA01 166 bp overlap
BCOR 3 datasets
ChIP WA01 GSE104690.BCOR.WA01 549 bp overlap
ChIP WA01_KDM2B_KO GSE104690.BCOR.WA01_KDM2B_KO 367 bp overlap
ChIP WA01_RNF2-R_Doxycyclin GSE104690.BCOR.WA01_RNF2-R_Doxycyclin 422 bp overlap
BRD4 5 datasets
ChIP Jurkat GSE83777.BRD4.Jurkat 450 bp overlap
ChIP KOPT-K1 GSE54379.BRD4.KOPT-K1 331 bp overlap
ChIP RH4 GSE83726.BRD4.RH4 410 bp overlap
ChIP RH4_shCHD4 GSE140115.BRD4.RH4_shCHD4 343 bp overlap
ChIP RH4_shscr GSE140115.BRD4.RH4_shscr 210 bp overlap
CASZ1 1 dataset
ChIP rhabdomyosarcoma_DOX GSE126142.CASZ1.rhabdomyosarcoma_DOX 469 bp overlap
CHD4 1 dataset
ChIP SCMC GSE155861.CHD4.SCMC 335 bp overlap
CHD7 2 datasets
ChIP H1 ENCFF126NLU 195 bp overlap
ChIP WA01 ENCSR000AVA.CHD7.WA01 305 bp overlap
CREB1 4 datasets
ChIP GM23338 ENCFF432ZEW 305 bp overlap
ChIP GM23338 ENCSR214ZAV.CREB1.GM23338 378 bp overlap
ChIP H1 ENCFF955PMP 321 bp overlap
ChIP WA01 ENCSR000BSN.CREB1.WA01 446 bp overlap
CRY1 2 datasets
ChIP U2OS GSE130602.CRY1.U2OS 506 bp overlap
ChIP U2OS_DMSO GSE130506.CRY1.U2OS_DMSO 506 bp overlap
CRY2 1 dataset
ChIP U2OS GSE130602.CRY2.U2OS 346 bp overlap
CTBP2 2 datasets
ChIP H1 ENCFF329MAX 391 bp overlap
ChIP WA01 ENCSR000EUO.CTBP2.WA01 556 bp overlap
CTNNB1 1 dataset
ChIP hESC_activinA_15h GSE99202.CTNNB1.hESC_activinA_15h 309 bp overlap
DUX4 1 dataset
Motif ES_0h ES_0h-DUX4_MA0468.1 11 bp overlap
E2F6 1 dataset
ChIP WA01 ENCSR000BSI.E2F6.WA01 255 bp overlap
EP300 3 datasets
ChIP H1 ENCFF927IYK 199 bp overlap
ChIP WA01 ENCSR000BKK.EP300.WA01 307 bp overlap
ChIP hESC GSE17917.EP300.hESC 336 bp overlap
ETV6 2 datasets
ChIP WTC11 ENCFF812SCD 300 bp overlap
ChIP WTC11 ENCFF812SCD 437 bp overlap
FOXA1 1 dataset
ChIP pancreatic-progenitor_PP1 GSE104840.FOXA1.pancreatic-progenitor_PP1 354 bp overlap
FOXA2 3 datasets
ChIP DE DE-FOXA2-1 556 bp overlap
ChIP DE DE-FOXA2-2 556 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.FOXA2.pancreatic-progenitor_PP1 437 bp overlap
FOXP1 1 dataset
ChIP H9 GSE31006.FOXP1.H9 530 bp overlap
FOXP4 2 datasets
ChIP WTC11 ENCFF708TAF 309 bp overlap
ChIP WTC11 ENCFF708TAF 377 bp overlap
GATA2 1 dataset
ChIP hiPSC_SLC6ebs GSE107639.GATA2.hiPSC_SLC6ebs 205 bp overlap
GATA4 6 datasets
ChIP DE DE-GATA4-1 367 bp overlap
ChIP DE DE-GATA4-2 555 bp overlap
Motif ES_0h ES_0h-GATA4_MA0482.3 8 bp overlap
ChIP foregut GSE117136.GATA4.foregut 460 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA4.pancreatic-progenitor 440 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA4.pancreatic-progenitor_PP1 556 bp overlap
GATA5 1 dataset
Motif ES_0h ES_0h-GATA5_MA0766.3 8 bp overlap
GATA6 10 datasets
ChIP DE DE-GATA6-1 424 bp overlap
ChIP DE DE-GATA6-2 518 bp overlap
ChIP DE_D1 S09-DE-d1-GATA6-exp1 349 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 353 bp overlap
Motif ES_0h ES_0h-GATA6_MA1104.3 8 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 415 bp overlap
ChIP foregut GSE117136.GATA6.foregut 451 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA6.pancreatic-progenitor 404 bp overlap
ChIP pancreatic-progenitor_KO GSE117136.GATA6.pancreatic-progenitor_KO 459 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA6.pancreatic-progenitor_PP1 487 bp overlap
GLIS3 1 dataset
ChIP H9_plus GSE109562.GLIS3.H9_plus 416 bp overlap
Gata3 1 dataset
Motif ES_0h ES_0h-Gata3_MA0037.5 8 bp overlap
HDAC1 1 dataset
ChIP K-562 ENCSR711VWL.HDAC1.K-562 203 bp overlap
HDAC2 3 datasets
ChIP RH4_DMSO-6H GSE116344.HDAC2.RH4_DMSO-6H 408 bp overlap
ChIP RH4_Entinostat-6H GSE116344.HDAC2.RH4_Entinostat-6H 257 bp overlap
ChIP WA01 ENCSR000BNR.HDAC2.WA01 205 bp overlap
HDAC3 1 dataset
ChIP RH4_DMSO-6H GSE116344.HDAC3.RH4_DMSO-6H 511 bp overlap
IRF2 1 dataset
Motif ES_0h ES_0h-IRF2_MA0051.2 16 bp overlap
IRF7 1 dataset
Motif ES_0h ES_0h-IRF7_MA0772.2 13 bp overlap
Irf1 1 dataset
Motif ES_0h ES_0h-Irf1_MA0050.4 11 bp overlap
JARID2 1 dataset
ChIP MRC-5_IPS25_CTR GSE48515.JARID2.MRC-5_IPS25_CTR 195 bp overlap
JUN 7 datasets
ChIP DE_D1 S40-DE-d1-JUN-exp2 286 bp overlap
ChIP DE_D2 S02-DE-d2-JUN-exp1 426 bp overlap
ChIP DE_D2 S54-DE-d2-JUN-exp2 489 bp overlap
ChIP ESC S24-ESC-d0-JUN-exp1 523 bp overlap
ChIP ESC S34-ESC-d0-JUN-exp2 514 bp overlap
ChIP HUES-8 GSE109524.JUN.HUES-8 488 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.JUN.HUES-8_DE_ctrl 301 bp overlap
JUND 2 datasets
ChIP H1 ENCFF010YXS 311 bp overlap
ChIP WA01 ENCSR000BKP.JUND.WA01 102 bp overlap
KLF4 2 datasets
ChIP WA09 GSE105028.KLF4.WA09 168 bp overlap
ChIP hiPSC GSE56567.KLF4.hiPSC 182 bp overlap
LEF1 1 dataset
ChIP hESC GSE64758.LEF1.hESC 197 bp overlap
LMO2 1 dataset
ChIP CCRF-CEM GSE33850.LMO2.CCRF-CEM 230 bp overlap
MAX 3 datasets
ChIP H1 ENCFF914VQY 357 bp overlap
ChIP WA01 ENCSR000EUP.MAX.WA01 167 bp overlap
ChIP WTC11 ENCFF223QFY 538 bp overlap
MED1 2 datasets
ChIP RH4 GSE83726.MED1.RH4 245 bp overlap
ChIP hESC_PRIMED GSE69646.MED1.hESC_PRIMED 456 bp overlap
MEIS1 1 dataset
Motif ES_0h ES_0h-MEIS1_MA0498.3 5 bp overlap
MTA2 1 dataset
ChIP RH4 GSE155861.MTA2.RH4 278 bp overlap
MYB 3 datasets
Motif ES_0h ES_0h-MYB_MA0100.4 6 bp overlap
ChIP Jurkat GSE59657.MYB.Jurkat 279 bp overlap
ChIP MOLT-3 GSE59657.MYB.MOLT-3 306 bp overlap
MYOD1 7 datasets
ChIP RD GSE137168.MYOD1.RD 452 bp overlap
ChIP RD_shSNAI2 GSE137168.MYOD1.RD_shSNAI2 515 bp overlap
ChIP RH30_DMSO GSE85169.MYOD1.RH30_DMSO 531 bp overlap
ChIP RH4 GSE83726.MYOD1.RH4 282 bp overlap
ChIP SMS-CTR GSE137168.MYOD1.SMS-CTR 325 bp overlap
ChIP myoblast GSE50413.MYOD1.myoblast 242 bp overlap
ChIP rhabdomyosarcoma GSE50413.MYOD1.rhabdomyosarcoma 329 bp overlap
MYOG 2 datasets
Motif ES_0h ES_0h-MYOG_MA0500.3 8 bp overlap
ChIP RH4_Entinostat-6H GSE116344.MYOG.RH4_Entinostat-6H 234 bp overlap
NANOG 13 datasets
ChIP GM23338 ENCFF065NZG 234 bp overlap
ChIP GM23338 ENCSR061DGF.NANOG.GM23338 223 bp overlap
ChIP H1 ENCFF747ZPQ 221 bp overlap
ChIP HUES-8 GSE109524.NANOG.HUES-8 556 bp overlap
ChIP WA01 ERP004238.NANOG.WA01 556 bp overlap
ChIP WA01 ENCSR000BMT.NANOG.WA01 195 bp overlap
ChIP WA01 ENCSR000BMT.NANOG.WA01 140 bp overlap
ChIP WA01_3IL ERP004238.NANOG.WA01_3IL 556 bp overlap
ChIP WA09 GSE105028.NANOG.WA09 556 bp overlap
ChIP WA09_heat-shock GSE105028.NANOG.WA09_heat-shock 391 bp overlap
ChIP hESC GSE18292.NANOG.hESC 152 bp overlap
ChIP hESC GSE20650.NANOG.hESC 556 bp overlap
ChIP hESC GSE18292.NANOG.hESC 128 bp overlap
NHLH1 1 dataset
Motif ES_0h ES_0h-NHLH1_MA0048.3 9 bp overlap
NIPBL 5 datasets
ChIP WA09 GSE105028.NIPBL.WA09 476 bp overlap
ChIP hESC GSE64758.NIPBL.hESC 321 bp overlap
ChIP hESC_WNT3A GSE64758.NIPBL.hESC_WNT3A 299 bp overlap
ChIP hESC_WNT3A_ACTIVIN GSE64758.NIPBL.hESC_WNT3A_ACTIVIN 266 bp overlap
ChIP hESC_activin GSE64758.NIPBL.hESC_activin 264 bp overlap
OTX2 1 dataset
ChIP WTC11 ENCFF634NAO 120 bp overlap
Olig2 1 dataset
Motif ES_0h ES_0h-Olig2_MA1997.2 6 bp overlap
PCGF1 1 dataset
ChIP WA01 GSE104690.PCGF1.WA01 162 bp overlap
PDX1 1 dataset
ChIP hiPSC_derived_PPs GSE106949.PDX1.hiPSC_derived_PPs 252 bp overlap
POLR2A 2 datasets
ChIP GM23338 ENCFF450WCS 204 bp overlap
ChIP H1 ENCFF833NJP 341 bp overlap
POU5F1 16 datasets
ChIP BG03 GSE21614.POU5F1.BG03 184 bp overlap
ChIP GM23338 ENCFF333SNB 228 bp overlap
ChIP H1 ENCFF698ZAP 191 bp overlap
ChIP HUES-8 GSE109524.POU5F1.HUES-8 556 bp overlap
ChIP NCCIT GSE36134.POU5F1.NCCIT 375 bp overlap
ChIP NCCIT_SNF5 GSE36134.POU5F1.NCCIT_SNF5 556 bp overlap
ChIP WA01 ENCSR000BMU.POU5F1.WA01 215 bp overlap
ChIP WA01_3IL ERP004238.POU5F1.WA01_3IL 200 bp overlap
ChIP WA09 GSE105028.POU5F1.WA09 270 bp overlap
ChIP WA09_heat-shock GSE105028.POU5F1.WA09_heat-shock 269 bp overlap
ChIP hESC ENCSR264RJX.POU5F1.hESC 252 bp overlap
ChIP hESC GSE20650.POU5F1.hESC 132 bp overlap
ChIP hESC_NAIVE GSE69646.POU5F1.hESC_NAIVE 269 bp overlap
ChIP hESC_PRIMED GSE69646.POU5F1.hESC_PRIMED 556 bp overlap
ChIP hiPSC GSE56567.POU5F1.hiPSC 287 bp overlap
ChIP hiPSC_3s2 GSE81899.POU5F1.hiPSC_3s2 337 bp overlap
PRDM1 1 dataset
Motif ES_0h ES_0h-PRDM1_MA0508.4 7 bp overlap
PRDM14 3 datasets
ChIP NCCIT GSE71675.PRDM14.NCCIT 270 bp overlap
ChIP hESC GSE22767.PRDM14.hESC 536 bp overlap
ChIP hESC GSE138674.PRDM14.hESC 540 bp overlap
PRDM9 1 dataset
Motif ES_0h ES_0h-PRDM9_MA1723.2 20 bp overlap
Prdm14 1 dataset
Motif ES_0h ES_0h-Prdm14_MA1998.2 8 bp overlap
RAD21 1 dataset
ChIP hiPSC_parental-line GSE106870.RAD21.hiPSC_parental-line 170 bp overlap
RBBP4 1 dataset
ChIP RH5 GSE155861.RBBP4.RH5 175 bp overlap
RNF2 1 dataset
ChIP WA09 GSE105028.RNF2.WA09 273 bp overlap
RXRA 1 dataset
ChIP WA01 ENCSR000BJW.RXRA.WA01 216 bp overlap
SALL3 1 dataset
ChIP hiPSC GSE104863.SALL3.hiPSC 129 bp overlap
SIN3A 1 dataset
ChIP WA01 ENCSR000BIS.SIN3A.WA01 176 bp overlap
SMAD1 2 datasets
ChIP BG03 GSE36578.SMAD1.BG03 96 bp overlap
ChIP BG03 GSE36578.SMAD1.BG03 96 bp overlap
SMAD2 1 dataset
ChIP hESC_activinA_15h GSE99202.SMAD2.hESC_activinA_15h 290 bp overlap
SMAD2-3 3 datasets
ChIP HUES-8 GSE109524.SMAD2-3.HUES-8 294 bp overlap
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 556 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.SMAD2-3.HUES-8_DE_ctrl 152 bp overlap
SMAD2_3 4 datasets
ChIP DE_D1 S07-DE-d1-SMAD2_3-exp1 138 bp overlap
ChIP DE_D2 S01-DE-d2-SMAD2_3-exp1 227 bp overlap
ChIP DE_D2 S53-DE-d2-SMAD2_3-exp2 341 bp overlap
ChIP ESC S33-ESC-d0-SMAD2_3-exp2 335 bp overlap
SMAD3 3 datasets
ChIP BG03 GSE21614.SMAD3.BG03 396 bp overlap
ChIP BG03_DIFF_2H GSE36578.SMAD3.BG03_DIFF_2H 243 bp overlap
ChIP hESC GSE29422.SMAD3.hESC 276 bp overlap
SMARCA4 5 datasets
ChIP HS-SY-2_HA-tagged GSE108926.SMARCA4.HS-SY-2_HA-tagged 370 bp overlap
ChIP WA09 GSE105028.SMARCA4.WA09 143 bp overlap
ChIP WA09_heat-shock GSE105028.SMARCA4.WA09_heat-shock 97 bp overlap
ChIP hiPSC GSE124903.SMARCA4.hiPSC 556 bp overlap
ChIP hiPSC_WT GSE124903.SMARCA4.hiPSC_WT 412 bp overlap
SMARCB1 3 datasets
ChIP hiPSC GSE124903.SMARCB1.hiPSC 307 bp overlap
ChIP hiPSC GSE124903.SMARCB1.hiPSC 157 bp overlap
ChIP hiPSC_WT GSE124903.SMARCB1.hiPSC_WT 556 bp overlap
SMARCC1 5 datasets
ChIP ESC S25-ESC-d0-BAF155-exp1 556 bp overlap
ChIP HS-SY-2 GSE108025.SMARCC1.HS-SY-2 229 bp overlap
ChIP HS-SY-2 GSE108025.SMARCC1.HS-SY-2 208 bp overlap
ChIP hiPSC GSE124903.SMARCC1.hiPSC 494 bp overlap
ChIP hiPSC_WT GSE124903.SMARCC1.hiPSC_WT 301 bp overlap
SNAI2 1 dataset
ChIP RD GSE137168.SNAI2.RD 496 bp overlap
SOX2 5 datasets
ChIP LK2_GFP GSE137459.SOX2.LK2_GFP 173 bp overlap
ChIP hESC GSE69479.SOX2.hESC 192 bp overlap
ChIP hESC GSE18292.SOX2.hESC 145 bp overlap
ChIP hiPSC GSE56567.SOX2.hiPSC 318 bp overlap
ChIP hiPSC_3s2 GSE81899.SOX2.hiPSC_3s2 84 bp overlap
SOX8 2 datasets
ChIP RH4_DMSO-6H GSE116344.SOX8.RH4_DMSO-6H 250 bp overlap
ChIP RH4_Entinostat-6H GSE116344.SOX8.RH4_Entinostat-6H 201 bp overlap
SP1 3 datasets
ChIP H1 ENCFF263FUH 256 bp overlap
ChIP WA01 ENCSR000BIR.SP1.WA01 318 bp overlap
ChIP WTC11 ENCFF688PEU 374 bp overlap
SPIC 1 dataset
Motif ES_0h ES_0h-SPIC_MA0687.2 13 bp overlap
SRF 2 datasets
ChIP H1 ENCFF036PEF 69 bp overlap
ChIP WA01 ENCSR000BIV.SRF.WA01 88 bp overlap
STAT3 1 dataset
ChIP WA01 ERP004237.STAT3.WA01 182 bp overlap
Stat2 1 dataset
Motif ES_0h ES_0h-Stat2_MA1623.2 10 bp overlap
TAF1 2 datasets
ChIP H1 ENCFF478SZO 434 bp overlap
ChIP WA01 ENCSR000BHO.TAF1.WA01 139 bp overlap
TAL1 1 dataset
ChIP PRIMA5 GSE33850.TAL1.PRIMA5 162 bp overlap
TCF12 2 datasets
ChIP H1 ENCFF203EBH 251 bp overlap
ChIP WA01 ENCSR000BIT.TCF12.WA01 449 bp overlap
TCF3 1 dataset
ChIP NPC GSE154479.TCF3.NPC 523 bp overlap
TEAD1 2 datasets
ChIP WTC11 ENCFF502QUV 379 bp overlap
ChIP pancreas_12D ERP008682.TEAD1.pancreas_12D 176 bp overlap
TEAD4 4 datasets
ChIP ESC S26-ESC-d0-TEAD4-exp1 541 bp overlap
ChIP H1 ENCFF778PAX 245 bp overlap
ChIP WTC11 ENCFF114TZS 341 bp overlap
ChIP hESC GSE99202.TEAD4.hESC 389 bp overlap
TP53 4 datasets
ChIP H9_ectoderm GSE142050.TP53.H9_ectoderm 467 bp overlap
ChIP H9_mesoderm GSE142050.TP53.H9_mesoderm 315 bp overlap
ChIP hESC_DAMAGED GSE39912.TP53.hESC_DAMAGED 464 bp overlap
ChIP hESC_DIFF GSE39912.TP53.hESC_DIFF 406 bp overlap
TRIM28 1 dataset
ChIP hESC GSE115387.TRIM28.hESC 365 bp overlap
TRPS1 1 dataset
Motif ES_0h ES_0h-TRPS1_MA1970.2 8 bp overlap
Tcf12 1 dataset
Motif ES_0h ES_0h-Tcf12_MA0521.3 6 bp overlap
Twist2 1 dataset
Motif ES_0h ES_0h-Twist2_MA0633.3 6 bp overlap
YY1 4 datasets
ChIP H1 ENCFF524BTL 307 bp overlap
ChIP RH4_DMSO-6H GSE116344.YY1.RH4_DMSO-6H 176 bp overlap
ChIP RH4_DMSO-6H GSE116344.YY1.RH4_DMSO-6H 127 bp overlap
ChIP WA01 ENCSR000BKD.YY1.WA01 178 bp overlap
ZBTB6 1 dataset
Motif ES_0h ES_0h-ZBTB6_MA1581.2 9 bp overlap
ZKSCAN8 1 dataset
ChIP WTC11 ENCFF666HNJ 345 bp overlap
ZNF114 1 dataset
ChIP GM23338 ENCFF631OSW 293 bp overlap
ZNF140 1 dataset
Motif ES_0h ES_0h-ZNF140_MA1589.2 19 bp overlap
ZNF143 1 dataset
ChIP WA09 GSE105028.ZNF143.WA09 173 bp overlap
ZNF184 1 dataset
Motif ES_0h ES_0h-ZNF184_MA2120.1 13 bp overlap
ZNF207 1 dataset
ChIP WA09 GSE118632.ZNF207.WA09 512 bp overlap
ZNF274 2 datasets
ChIP H1 ENCFF434LDY 108 bp overlap
ChIP WA01 ENCSR000EUN.ZNF274.WA01 202 bp overlap
ZNF331 1 dataset
Motif ES_0h ES_0h-ZNF331_MA1726.2 10 bp overlap
ZNF384 1 dataset
Motif ES_0h ES_0h-ZNF384_MA1125.2 8 bp overlap
ZNF423 1 dataset
ChIP WTC11 ENCFF574PBR 317 bp overlap
ZNF462 2 datasets
ChIP GM23338 ENCFF896CCA 251 bp overlap
ChIP GM23338 ENCSR334UWP.ZNF462.GM23338 466 bp overlap
ZNF530 2 datasets
Motif DE_12h DE_12h-ZNF530_MA1981.2 14 bp overlap
Motif ES_0h ES_0h-ZNF530_MA1981.2 14 bp overlap
ZNF85 1 dataset
Motif ES_0h ES_0h-ZNF85_MA1720.2 12 bp overlap