chr22 : 46,576,006 46,576,187
181 bp 120 TFs 1 linked gene
This 181 bp open chromatin element is linked to GRAMD4 and is bound by 120 transcription factors.
Linked Genes
1 gene
Gene Expression Dist. to TSS Distance Link type
GRAMD4 940 bp At TSS Proximity
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr22:46,571,006 – 46,581,187
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
120 transcription factors
Source
Cell type
AR 1 dataset
ChIP MCF-7 ERP001226.AR.MCF-7 179 bp overlap
ARID1A 1 dataset
ChIP MCF-7_4-hydroxytamoxifen GSE123284.ARID1A.MCF-7_4-hydroxytamoxifen 169 bp overlap
ARID3A 1 dataset
ChIP HepG2 ENCFF341DES 114 bp overlap
ARID4B 2 datasets
ChIP HepG2 ENCFF519OXJ 181 bp overlap
ChIP HepG2 ENCFF519OXJ 163 bp overlap
ASH2L 2 datasets
ChIP Hep-G2 ENCSR508LMH.ASH2L.Hep-G2 57 bp overlap
ChIP WA01 ENCSR850KIP.ASH2L.WA01 160 bp overlap
BRD1 1 dataset
ChIP HUES-64 GSE104059.BRD1.HUES-64 59 bp overlap
BRD4 13 datasets
ChIP BE2C GSE80151.BRD4.BE2C 181 bp overlap
ChIP HAP1 GSE108387.BRD4.HAP1 181 bp overlap
ChIP HCT-15 GSE73319.BRD4.HCT-15 67 bp overlap
ChIP HT29 GSE73319.BRD4.HT29 53 bp overlap
ChIP PC-3_GDC-R_SAHA GSE137207.BRD4.PC-3_GDC-R_SAHA 181 bp overlap
ChIP SK-N-BE2-C GSE80151.BRD4.SK-N-BE2-C 181 bp overlap
ChIP SK-N-BE2-C_Naive-veh GSE107706.BRD4.SK-N-BE2-C_Naive-veh 165 bp overlap
ChIP SK-N-BE2-C_Resistant-veh GSE107706.BRD4.SK-N-BE2-C_Resistant-veh 181 bp overlap
ChIP SW480 GSE73319.BRD4.SW480 91 bp overlap
ChIP SW480 GSE110473.BRD4.SW480 181 bp overlap
ChIP retina_AB1-FW15 GSE86981.BRD4.retina_AB1-FW15 181 bp overlap
ChIP retina_AB1-FW23 GSE86981.BRD4.retina_AB1-FW23 181 bp overlap
ChIP retina_AB1-RB GSE86981.BRD4.retina_AB1-RB 181 bp overlap
CDKN1B 1 dataset
ChIP MDA-MB-231 GSE112444.CDKN1B.MDA-MB-231 112 bp overlap
CHD1 1 dataset
ChIP HeLa-S3 ENCSR392EAA.CHD1.HeLa-S3 135 bp overlap
CREBBP 2 datasets
ChIP LS180_125 GSE39277.CREBBP.LS180_125 138 bp overlap
ChIP PC-3_GDC-resist GSE147455.CREBBP.PC-3_GDC-resist 181 bp overlap
CUX1 1 dataset
ChIP Hep-G2 ENCSR049KIZ.CUX1.Hep-G2 74 bp overlap
DRAP1 1 dataset
ChIP Hep-G2 ENCSR765MKZ.DRAP1.Hep-G2 181 bp overlap
ELF1 1 dataset
ChIP Hep-G2 ENCSR321VGW.ELF1.Hep-G2 99 bp overlap
ELF3 2 datasets
ChIP PDAC GSE64557.ELF3.PDAC 181 bp overlap
ChIP PDAC_KOKLF5 GSE64557.ELF3.PDAC_KOKLF5 181 bp overlap
EP300 2 datasets
ChIP HeLa-S3 ENCSR680OFU.EP300.HeLa-S3 172 bp overlap
ChIP PC-3 GSE147455.EP300.PC-3 128 bp overlap
ESR1 38 datasets
ChIP Ishikawa ENCSR000BIY.ESR1.Ishikawa 127 bp overlap
ChIP Ishikawa_E2 GSE132426.ESR1.Ishikawa_E2 181 bp overlap
ChIP Ishikawa_E2_D538G GSE132426.ESR1.Ishikawa_E2_D538G 143 bp overlap
ChIP Ishikawa_ETV4-KO2_Mili GSE129803.ESR1.Ishikawa_ETV4-KO2_Mili 181 bp overlap
ChIP MCF-7 GSE148277.ESR1.MCF-7 98 bp overlap
ChIP MCF-7 GSE103023.ESR1.MCF-7 181 bp overlap
ChIP MCF-7 GSE41561.ESR1.MCF-7 179 bp overlap
ChIP MCF-7 GSE48930.ESR1.MCF-7 165 bp overlap
ChIP MCF-7_ARID1A-KO GSE123284.ESR1.MCF-7_ARID1A-KO 181 bp overlap
ChIP MCF-7_ARID1A-KO_4-OHT GSE123284.ESR1.MCF-7_ARID1A-KO_4-OHT 155 bp overlap
ChIP MCF-7_ARID1A-KO_4-OHT_clone14 GSE123284.ESR1.MCF-7_ARID1A-KO_4-OHT_clone14 155 bp overlap
ChIP MCF-7_ARID1A-KO_clone14 GSE123284.ESR1.MCF-7_ARID1A-KO_clone14 181 bp overlap
ChIP MCF-7_AZD2014 GSE103023.ESR1.MCF-7_AZD2014 181 bp overlap
ChIP MCF-7_E2 GSE117941.ESR1.MCF-7_E2 181 bp overlap
ChIP MCF-7_E2 ERP000209.ESR1.MCF-7_E2 169 bp overlap
ChIP MCF-7_E2_90min GSE109820.ESR1.MCF-7_E2_90min 172 bp overlap
ChIP MCF-7_PROG GSE68355.ESR1.MCF-7_PROG 181 bp overlap
ChIP MCF-7_RAD001 GSE103023.ESR1.MCF-7_RAD001 181 bp overlap
ChIP MCF-7_SHCRT_E2 ERP000380.ESR1.MCF-7_SHCRT_E2 130 bp overlap
ChIP MCF-7_SICTR_E2 GSE40129.ESR1.MCF-7_SICTR_E2 151 bp overlap
ChIP MCF-7_SIGATA_E2 GSE40129.ESR1.MCF-7_SIGATA_E2 173 bp overlap
ChIP MCF-7_TamR GSE128445.ESR1.MCF-7_TamR 181 bp overlap
ChIP MCF-7_Y537S_DMSO GSE148277.ESR1.MCF-7_Y537S_DMSO 147 bp overlap
ChIP MCF-7_estradiol-DHT_4h GSE99626.ESR1.MCF-7_estradiol-DHT_4h 181 bp overlap
ChIP MCF-7_estradiol-Dex_4h GSE99626.ESR1.MCF-7_estradiol-Dex_4h 176 bp overlap
ChIP MCF-7_estradiol-progesterone_4h GSE99626.ESR1.MCF-7_estradiol-progesterone_4h 126 bp overlap
ChIP MCF-7_estradiol_4h GSE99626.ESR1.MCF-7_estradiol_4h 181 bp overlap
ChIP MCF-7_jc5850 GSE126004.ESR1.MCF-7_jc5850 181 bp overlap
ChIP MCF-7_parental GSE123284.ESR1.MCF-7_parental 181 bp overlap
ChIP MCF-7_parental_4-hydroxytamoxifen GSE123284.ESR1.MCF-7_parental_4-hydroxytamoxifen 177 bp overlap
ChIP MCF-7_shTEAD4 GSE125594.ESR1.MCF-7_shTEAD4 140 bp overlap
ChIP T-47D ENCSR000BQD.ESR1.T-47D 115 bp overlap
ChIP T-47D_siCont-IL6 GSE126004.ESR1.T-47D_siCont-IL6 181 bp overlap
ChIP T-47D_siFOXA1-Veh GSE126004.ESR1.T-47D_siFOXA1-Veh 138 bp overlap
ChIP U2OS_E2 GSE26110.ESR1.U2OS_E2 181 bp overlap
ChIP breast_tumor_Male_21 GSE104399.ESR1.breast_tumor_Male_21 181 bp overlap
ChIP breast_tumor_Male_26 GSE104399.ESR1.breast_tumor_Male_26 168 bp overlap
ChIP primary-endometrium-cancer_E1_DSG GSE114737.ESR1.primary-endometrium-cancer_E1_DSG 167 bp overlap
ESR1_Y537C 1 dataset
ChIP SUM44PE__ESR1_mutant_LTED GSE100074.ESR1_Y537C.SUM44PE__ESR1_mutant_LTED 181 bp overlap
ESR1_Y537N 2 datasets
ChIP MCF-7_dox GSE94493.ESR1_Y537N.MCF-7_dox 181 bp overlap
ChIP MCF-7_dox_E2 GSE94493.ESR1_Y537N.MCF-7_dox_E2 124 bp overlap
ESR1_Y537S 2 datasets
ChIP MCF-7_E2 GSE94493.ESR1_Y537S.MCF-7_E2 181 bp overlap
ChIP MCF-7_dox GSE94493.ESR1_Y537S.MCF-7_dox 181 bp overlap
ESR2 1 dataset
ChIP MDA-MB-231_estradiol GSE108979.ESR2.MDA-MB-231_estradiol 181 bp overlap
ESRRA 7 datasets
ChIP BT-474 GSE75876.ESRRA.BT-474 181 bp overlap
ChIP BT-474_AICAR GSE75876.ESRRA.BT-474_AICAR 181 bp overlap
ChIP BT-474_EGF GSE81651.ESRRA.BT-474_EGF 181 bp overlap
ChIP BT-474_HRG GSE81651.ESRRA.BT-474_HRG 181 bp overlap
ChIP SK-BR-3 GSE81651.ESRRA.SK-BR-3 181 bp overlap
ChIP SK-BR-3_EGF GSE81651.ESRRA.SK-BR-3_EGF 181 bp overlap
ChIP SK-BR-3_HRG GSE81651.ESRRA.SK-BR-3_HRG 181 bp overlap
ESRRG 2 datasets
ChIP BT-474 GSE144224.ESRRG.BT-474 181 bp overlap
ChIP cardiomyocyte GSE113760.ESRRG.cardiomyocyte 181 bp overlap
ETS1 2 datasets
ChIP SCC-25 GSE109884.ETS1.SCC-25 69 bp overlap
ChIP THP-6_shCtrl GSE138516.ETS1.THP-6_shCtrl 157 bp overlap
ETV4 1 dataset
ChIP T-47D GSE129803.ETV4.T-47D 181 bp overlap
EZH2 1 dataset
ChIP SU-DHL-6 GSE134136.EZH2.SU-DHL-6 181 bp overlap
FEZF1 2 datasets
ChIP HEK293 ENCSR827NWO.FEZF1.HEK293 181 bp overlap
ChIP HEK293 GSE76494.FEZF1.HEK293 181 bp overlap
FOXA1 1 dataset
ChIP breast-cancer_Veh-2856 GSE128018.FOXA1.breast-cancer_Veh-2856 141 bp overlap
FOXP4 1 dataset
ChIP Hep-G2 ENCSR232LLP.FOXP4.Hep-G2 163 bp overlap
GABPB1 1 dataset
ChIP HepG2 ENCFF315AWN 181 bp overlap
HDAC1 2 datasets
ChIP HepG2 ENCFF304IEJ 161 bp overlap
ChIP PC-3 GSE147455.HDAC1.PC-3 115 bp overlap
HDAC2 2 datasets
ChIP Hep-G2 ENCSR000BMC.HDAC2.Hep-G2 176 bp overlap
ChIP PC-3 GSE147455.HDAC2.PC-3 181 bp overlap
HDGF 1 dataset
ChIP K-562 ENCSR563YDA.HDGF.K-562 79 bp overlap
HMGXB4 3 datasets
ChIP Hep-G2 ENCSR174GOO.HMGXB4.Hep-G2 155 bp overlap
ChIP HepG2 ENCFF032DND 59 bp overlap
ChIP HepG2 ENCFF179TAD 94 bp overlap
HNF4A 2 datasets
ChIP Hep-G2 ENCSR000BLF.HNF4A.Hep-G2 144 bp overlap
ChIP KATO-III GSE114018.HNF4A.KATO-III 127 bp overlap
HOXA3 1 dataset
ChIP Hep-G2 ENCSR999JKC.HOXA3.Hep-G2 181 bp overlap
INO80 1 dataset
ChIP Huh-7 GSE97411.INO80.Huh-7 58 bp overlap
JUN 1 dataset
ChIP MDA-MB-231 GSE112444.JUN.MDA-MB-231 179 bp overlap
JUNB 1 dataset
ChIP HAEC GSE89970.JUNB.HAEC 95 bp overlap
KDM5B 1 dataset
ChIP T-47D GSE46055.KDM5B.T-47D 158 bp overlap
KLF1 3 datasets
ChIP HEK293 ENCFF159QSW 181 bp overlap
ChIP HEK293 ENCSR859BMR.KLF1.HEK293 181 bp overlap
ChIP HEK293 GSE76494.KLF1.HEK293 142 bp overlap
KLF17 1 dataset
ChIP HEK293 ENCSR065WUF.KLF17.HEK293 181 bp overlap
KLF4 1 dataset
ChIP WIBR3 GSE130417.KLF4.WIBR3 131 bp overlap
KLF5 3 datasets
ChIP ESO-26 GSE132680.KLF5.ESO-26 181 bp overlap
ChIP GP5D GSE51234.KLF5.GP5D 143 bp overlap
ChIP KATO-III GSE51705.KLF5.KATO-III 181 bp overlap
KLF9 1 dataset
ChIP HEK293 ENCSR076EZB.KLF9.HEK293 61 bp overlap
MAX 1 dataset
ChIP Hep-G2 ENCSR168DYA.MAX.Hep-G2 156 bp overlap
MAZ 4 datasets
ChIP HEK293 ENCFF994GSG 181 bp overlap
ChIP HEK293 ENCFF994GSG 140 bp overlap
ChIP HEK293 ENCSR290SSQ.MAZ.HEK293 181 bp overlap
ChIP HEK293 GSE76494.MAZ.HEK293 181 bp overlap
MED1 3 datasets
ChIP HUVEC-C_siCTL GSE124892.MED1.HUVEC-C_siCTL 181 bp overlap
ChIP HUVEC-C_siERG GSE124892.MED1.HUVEC-C_siERG 181 bp overlap
ChIP Hep-G2 ENCSR959XNY.MED1.Hep-G2 150 bp overlap
MED26 1 dataset
ChIP U2OS_SHEP400 GSE73742.MED26.U2OS_SHEP400 52 bp overlap
MYC 1 dataset
ChIP NB69 GSE138295.MYC.NB69 181 bp overlap
MYCN 2 datasets
ChIP COG-N-415 GSE138295.MYCN.COG-N-415 181 bp overlap
ChIP LA-N-5 GSE138295.MYCN.LA-N-5 71 bp overlap
MYNN 1 dataset
ChIP Hep-G2 ENCSR398TMP.MYNN.Hep-G2 181 bp overlap
NCOA2 2 datasets
ChIP HepG2 ENCFF853BJJ 181 bp overlap
ChIP MCF-7_E2 ERP000901.NCOA2.MCF-7_E2 130 bp overlap
NCOR1 1 dataset
ChIP LS180_125 GSE39277.NCOR1.LS180_125 68 bp overlap
NKX2-1 1 dataset
ChIP NCI-H3122 GSE39998.NKX2-1.NCI-H3122 174 bp overlap
NR2F2 1 dataset
ChIP MCF-7 ENCSR000BUY.NR2F2.MCF-7 153 bp overlap
NR5A1 2 datasets
ChIP Hep-G2 ENCSR310OZS.NR5A1.Hep-G2 168 bp overlap
ChIP HepG2 ENCFF970YZO 181 bp overlap
NR5A2 1 dataset
ChIP A-549 ENCSR190GIW.NR5A2.A-549 181 bp overlap
NRIP1 2 datasets
ChIP MCF-7 ERP005838.NRIP1.MCF-7 175 bp overlap
ChIP MCF-7_E2 ERP005838.NRIP1.MCF-7_E2 181 bp overlap
PATZ1 2 datasets
ChIP HEK293 ENCFF016MNJ 147 bp overlap
ChIP HEK293 ENCSR966ULI.PATZ1.HEK293 181 bp overlap
PHF5A 1 dataset
ChIP Hep-G2 ENCSR191TLD.PHF5A.Hep-G2 159 bp overlap
POLR2A 2 datasets
ChIP HeLa-S3 ENCFF224LWS 181 bp overlap
ChIP Peyer's patch ENCFF767HVN 181 bp overlap
POU5F1 1 dataset
ChIP DE_D1 DED1-OCT4_Batch_II 181 bp overlap
POU5F1_M 1 dataset
ChIP DE_D1 DED1-OCT4-M_Batch_II 181 bp overlap
RAD21 1 dataset
ChIP HeLa-S3_CTCF-s-OE GSE108869.RAD21.HeLa-S3_CTCF-s-OE 58 bp overlap
RARA 1 dataset
ChIP HepG2 ENCFF582XUA 181 bp overlap
RBBP5 1 dataset
ChIP K562 ENCFF070CVK 181 bp overlap
RBM39 2 datasets
ChIP Hep-G2 ENCSR339JTP.RBM39.Hep-G2 181 bp overlap
ChIP Hep-G2 GSE120104.RBM39.Hep-G2 181 bp overlap
RELA 1 dataset
ChIP FaDu_LPS GSE132018.RELA.FaDu_LPS 181 bp overlap
RXR 1 dataset
ChIP LS180_125 GSE31939.RXR.LS180_125 181 bp overlap
SKIL 1 dataset
ChIP HepG2 ENCFF823HPQ 181 bp overlap
SMAD1 2 datasets
ChIP Hep-G2 ENCSR213QOZ.SMAD1.Hep-G2 153 bp overlap
ChIP HepG2 ENCFF892OZT 181 bp overlap
SMAD3 2 datasets
ChIP NCI-H441 GSE51509.SMAD3.NCI-H441 181 bp overlap
ChIP NCI-H441_SIITTF1 GSE51509.SMAD3.NCI-H441_SIITTF1 181 bp overlap
SMAD4 2 datasets
ChIP Caco-2 GSE112946.SMAD4.Caco-2 181 bp overlap
ChIP HepG2 ENCFF615GTE 181 bp overlap
SMARCA4 3 datasets
ChIP HS-SY-2_HA-tagged GSE108926.SMARCA4.HS-SY-2_HA-tagged 181 bp overlap
ChIP J-Lat_GFP-Clone-A72_DMSO GSE100266.SMARCA4.J-Lat_GFP-Clone-A72_DMSO 181 bp overlap
ChIP endometrial-epithelial-cells_KO GSE106660.SMARCA4.endometrial-epithelial-cells_KO 181 bp overlap
SMARCB1 1 dataset
ChIP Hep-G2 GSE69566.SMARCB1.Hep-G2 181 bp overlap
SNAI2 1 dataset
ChIP PC-9_2DF GSE131687.SNAI2.PC-9_2DF 164 bp overlap
SOX2 1 dataset
ChIP LK2_GFP GSE137459.SOX2.LK2_GFP 181 bp overlap
SP1 1 dataset
ChIP HEK293 GSE76494.SP1.HEK293 160 bp overlap
SP2 3 datasets
ChIP HEK293 ENCFF181QXT 181 bp overlap
ChIP HEK293 ENCSR807LQP.SP2.HEK293 181 bp overlap
ChIP HEK293 GSE76494.SP2.HEK293 181 bp overlap
SP4 1 dataset
ChIP HEK293 GSE76494.SP4.HEK293 180 bp overlap
SP5 2 datasets
ChIP Hep-G2 ENCSR019NPF.SP5.Hep-G2 181 bp overlap
ChIP HepG2 ENCFF931FHV 181 bp overlap
STAT1 1 dataset
ChIP NCI-H358 GSE79707.STAT1.NCI-H358 181 bp overlap
STAT3 1 dataset
ChIP NCI-H358 GSE79707.STAT3.NCI-H358 175 bp overlap
SUPT5H 1 dataset
ChIP DLD-1_NELFCD-AID GSE144786.SUPT5H.DLD-1_NELFCD-AID 181 bp overlap
TCF7L2 4 datasets
ChIP HCT-116 ENCSR000EUV.TCF7L2.HCT-116 181 bp overlap
ChIP HCT116 ENCFF038POZ 181 bp overlap
ChIP HeLa-S3 ENCSR000EVE.TCF7L2.HeLa-S3 79 bp overlap
ChIP PANC-1 ENCSR000EXL.TCF7L2.PANC-1 102 bp overlap
THAP9 2 datasets
ChIP HepG2 ENCFF687WSR 111 bp overlap
ChIP HepG2 ENCFF687WSR 181 bp overlap
TP53 1 dataset
ChIP HCT-116_IR GSE60267.TP53.HCT-116_IR 121 bp overlap
U2AF1 1 dataset
ChIP Hep-G2 GSE120104.U2AF1.Hep-G2 55 bp overlap
U2AF1L5,U2AF1 1 dataset
ChIP HepG2 ENCFF548XGJ 181 bp overlap
VDR 4 datasets
ChIP LS180_125 GSE31939.VDR.LS180_125 181 bp overlap
ChIP LX2_CALCIPOTRIOL_TGFB1 GSE38103.VDR.LX2_CALCIPOTRIOL_TGFB1 175 bp overlap
ChIP THP-1_1d_1-25-OH-2D3 GSE89431.VDR.THP-1_1d_1-25-OH-2D3 181 bp overlap
ChIP primary-prostate-epithelial-cell GSE124576.VDR.primary-prostate-epithelial-cell 176 bp overlap
VEZF1 1 dataset
ChIP K562 ENCFF053XDV 61 bp overlap
XRCC5 2 datasets
ChIP K-562 ENCSR506KWJ.XRCC5.K-562 109 bp overlap
ChIP K-562 GSE120104.XRCC5.K-562 117 bp overlap
YY1 1 dataset
ChIP Huh-7 GSE97411.YY1.Huh-7 122 bp overlap
ZBTB6 1 dataset
ChIP HEK293 GSE76494.ZBTB6.HEK293 149 bp overlap
ZBTB7A 5 datasets
ChIP HUDEP-2 GSE103445.ZBTB7A.HUDEP-2 64 bp overlap
ChIP Hep-G2 ENCSR000BQA.ZBTB7A.Hep-G2 130 bp overlap
ChIP Ishikawa ENCFF191NFH 84 bp overlap
ChIP K-562 GSE103445.ZBTB7A.K-562 137 bp overlap
ChIP K-562 ENCSR000BME.ZBTB7A.K-562 124 bp overlap
ZEB1 1 dataset
ChIP Hep-G2 ENCSR000BVN.ZEB1.Hep-G2 74 bp overlap
ZEB2 1 dataset
ChIP K-562 ENCSR004GKA.ZEB2.K-562 148 bp overlap
ZFP91 1 dataset
ChIP HepG2 ENCFF012CME 181 bp overlap
ZGPAT 1 dataset
ChIP Hep-G2 ENCSR704IGU.ZGPAT.Hep-G2 181 bp overlap
ZMYM3 1 dataset
ChIP Hep-G2_Ab_JH39-2-2F10 GSE97661.ZMYM3.Hep-G2_Ab_JH39-2-2F10 135 bp overlap
ZNF221 1 dataset
ChIP HepG2 ENCFF374BUN 181 bp overlap
ZNF257 1 dataset
ChIP HEK293T GSE78099.ZNF257.HEK293T 146 bp overlap
ZNF430 1 dataset
ChIP HepG2 ENCFF967HQR 181 bp overlap
ZNF605 1 dataset
ChIP HepG2 ENCFF640NFJ 181 bp overlap
ZNF623 2 datasets
ChIP HEK293 ENCFF505YHP 181 bp overlap
ChIP HEK293 ENCSR022IZK.ZNF623.HEK293 181 bp overlap
ZSCAN25 1 dataset
ChIP HepG2 ENCFF265FLD 181 bp overlap