chr2 : 218,569,571 218,569,873
302 bp 106 TFs 2 linked genes
This 302 bp open chromatin element is linked to CNOT9 and USP37 and is bound by 106 transcription factors.
Linked Genes
2 genes
Distance
Gene Expression Dist. to TSS Distance Link type
CNOT9 992 bp At TSS Proximity
USP37 1.2 kb Proximal Proximity
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr2:218,564,571 – 218,574,873
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
106 transcription factors
Source
Cell type
ARID1A 2 datasets
ChIP RMG-I GSE120058.ARID1A.RMG-I 179 bp overlap
ChIP RMG-I GSE104545.ARID1A.RMG-I 220 bp overlap
ARID2 2 datasets
ChIP BIN-67_lentivirus-SMARCA4-K785R GSE117734.ARID2.BIN-67_lentivirus-SMARCA4-K785R 50 bp overlap
ChIP HepG2 ENCFF317ZHO 225 bp overlap
ARID4B 1 dataset
ChIP HepG2 ENCFF519OXJ 113 bp overlap
ASH2L 2 datasets
ChIP GM12878 ENCSR849WCQ.ASH2L.GM12878 302 bp overlap
ChIP WA01 ENCSR850KIP.ASH2L.WA01 247 bp overlap
ATF3 1 dataset
ChIP A-549 ENCSR000BPS.ATF3.A-549 153 bp overlap
BCL6 1 dataset
ChIP SU-DHL-4 GSE119038.BCL6.SU-DHL-4 130 bp overlap
BCOR 2 datasets
ChIP B-cell_GERMINAL_CENTER GSE43350.BCOR.B-cell_GERMINAL_CENTER 157 bp overlap
ChIP WA01_RNF2-R GSE104690.BCOR.WA01_RNF2-R 302 bp overlap
BRD1 2 datasets
ChIP HUES-64 GSE104059.BRD1.HUES-64 277 bp overlap
ChIP RKO GSE47190.BRD1.RKO 302 bp overlap
BRD2 7 datasets
ChIP HCC1806_NTsiRNA GSE116879.BRD2.HCC1806_NTsiRNA 57 bp overlap
ChIP MDA-MB-231_BAZ2AsiRNA GSE116879.BRD2.MDA-MB-231_BAZ2AsiRNA 173 bp overlap
ChIP MDA-MB-231_BAZ2BsiRNA GSE116879.BRD2.MDA-MB-231_BAZ2BsiRNA 191 bp overlap
ChIP MDA-MB-231_DMSO GSE116879.BRD2.MDA-MB-231_DMSO 154 bp overlap
ChIP MDA-MB-231_DMSO_rDNA GSE116879.BRD2.MDA-MB-231_DMSO_rDNA 154 bp overlap
ChIP MDA-MB-231_MGSK2801 GSE116879.BRD2.MDA-MB-231_MGSK2801 154 bp overlap
ChIP MDA-MB-231_MGSK2801_rDNA GSE116879.BRD2.MDA-MB-231_MGSK2801_rDNA 154 bp overlap
BRD3 2 datasets
ChIP MV4-11_DMSO GSE120715.BRD3.MV4-11_DMSO 302 bp overlap
ChIP MV4-11_IBET151_50nM GSE120715.BRD3.MV4-11_IBET151_50nM 302 bp overlap
BRD4 15 datasets
ChIP CHL-1 GSE95585.BRD4.CHL-1 72 bp overlap
ChIP CLL_patient4 GSE109411.BRD4.CLL_patient4 100 bp overlap
ChIP HEK293T GSE39579.BRD4.HEK293T 197 bp overlap
ChIP HT29 GSE73319.BRD4.HT29 262 bp overlap
ChIP LNCaP-C4-2_F133V GSE88871.BRD4.LNCaP-C4-2_F133V 302 bp overlap
ChIP LNCaP-C4-2_F133V_JQ1 GSE88871.BRD4.LNCaP-C4-2_F133V_JQ1 161 bp overlap
ChIP LNCaP-C4-2_OE GSE88871.BRD4.LNCaP-C4-2_OE 228 bp overlap
ChIP MDA-MB-231_DMSO GSE116879.BRD4.MDA-MB-231_DMSO 79 bp overlap
ChIP MDA-MB-231_DMSO_rDNA GSE116879.BRD4.MDA-MB-231_DMSO_rDNA 79 bp overlap
ChIP MOLM-14_DMSO GSE65138.BRD4.MOLM-14_DMSO 146 bp overlap
ChIP MOLM-14_IBET GSE65138.BRD4.MOLM-14_IBET 272 bp overlap
ChIP MV4-11_DMSO GSE120715.BRD4.MV4-11_DMSO 205 bp overlap
ChIP PC-3_GDC-R GSE137207.BRD4.PC-3_GDC-R 153 bp overlap
ChIP PC-3_GDC-R_SAHA GSE137207.BRD4.PC-3_GDC-R_SAHA 269 bp overlap
ChIP THP-1_iBET-BD2 GSE138084.BRD4.THP-1_iBET-BD2 50 bp overlap
CDK8 1 dataset
ChIP MOLM-14_IBET GSE65138.CDK8.MOLM-14_IBET 111 bp overlap
CHD1 2 datasets
ChIP K562 ENCFF118VJV 90 bp overlap
ChIP hMSC-TERT GSE89179.CHD1.hMSC-TERT 302 bp overlap
CREBBP 3 datasets
ChIP PC-3 GSE147455.CREBBP.PC-3 129 bp overlap
ChIP PC-3_GDC-resist GSE147455.CREBBP.PC-3_GDC-resist 302 bp overlap
ChIP tonsil_GCBc_p4 GSE89688.CREBBP.tonsil_GCBc_p4 247 bp overlap
CTCF 1 dataset
ChIP KMS-11 ENCFF853JKX 302 bp overlap
DRAP1 1 dataset
ChIP Hep-G2 ENCSR765MKZ.DRAP1.Hep-G2 191 bp overlap
EHF 1 dataset
ChIP RWPE-1 GSE114241.EHF.RWPE-1 302 bp overlap
ELF4 1 dataset
ChIP HepG2 ENCFF752OAT 62 bp overlap
EP300 1 dataset
ChIP PC-3 GSE147455.EP300.PC-3 145 bp overlap
ESR1 1 dataset
ChIP breast_tumor_Male_26 GSE104399.ESR1.breast_tumor_Male_26 134 bp overlap
ESR2 1 dataset
ChIP MDA-MB-231_estradiol GSE108979.ESR2.MDA-MB-231_estradiol 88 bp overlap
ETS1 1 dataset
ChIP HEY-A8 GSE101832.ETS1.HEY-A8 203 bp overlap
FEZF1 1 dataset
ChIP HEK293 ENCSR827NWO.FEZF1.HEK293 100 bp overlap
FOXA1 2 datasets
ChIP PDAC_KOKLF5 GSE64557.FOXA1.PDAC_KOKLF5 73 bp overlap
ChIP breast_tumor_Male_14 GSE104399.FOXA1.breast_tumor_Male_14 198 bp overlap
FOXA2 1 dataset
ChIP DE DE-FOXA2-1 104 bp overlap
FOXJ2 1 dataset
ChIP K562 ENCFF457GZC 142 bp overlap
FOXO1-PAX3 2 datasets
ChIP RH4_DMSO-6H GSE116344.FOXO1-PAX3.RH4_DMSO-6H 218 bp overlap
ChIP RH4_Entinostat-6H GSE116344.FOXO1-PAX3.RH4_Entinostat-6H 127 bp overlap
FOXP1 1 dataset
ChIP LNCaP GSE62492.FOXP1.LNCaP 184 bp overlap
GATA1 2 datasets
ChIP erythroblast ENCFF867JAR 88 bp overlap
ChIP erythroblast ENCSR000EXP.GATA1.erythroblast 302 bp overlap
GTF2E2 1 dataset
ChIP K562 ENCFF741URT 302 bp overlap
HDAC1 7 datasets
ChIP HepG2 ENCFF750ZWM 302 bp overlap
ChIP K-562 ENCSR711VWL.HDAC1.K-562 166 bp overlap
ChIP K-562 ENCSR387UWP.HDAC1.K-562 125 bp overlap
ChIP MCF-7_ARID1A-KO GSE123284.HDAC1.MCF-7_ARID1A-KO 139 bp overlap
ChIP MCF-7_ARID1A-KO_clone14 GSE123284.HDAC1.MCF-7_ARID1A-KO_clone14 82 bp overlap
ChIP MCF-7_parental GSE123284.HDAC1.MCF-7_parental 259 bp overlap
ChIP NB4 GSE126720.HDAC1.NB4 230 bp overlap
HDAC2 2 datasets
ChIP PC-3 GSE147455.HDAC2.PC-3 258 bp overlap
ChIP pre-B-cell GSE107886.HDAC2.pre-B-cell 149 bp overlap
HDGF 2 datasets
ChIP GM12878 ENCFF653WYI 302 bp overlap
ChIP K-562 ENCSR563YDA.HDGF.K-562 302 bp overlap
HOXB13 2 datasets
ChIP G-401 GSE65381.HOXB13.G-401 152 bp overlap
ChIP prostate_2030 GSE130408.HOXB13.prostate_2030 59 bp overlap
IKZF1 2 datasets
ChIP GM12878 ENCFF753XDO 81 bp overlap
ChIP pre-B-cell GSE107886.IKZF1.pre-B-cell 104 bp overlap
INO80 1 dataset
ChIP Hep-G2 GSE97411.INO80.Hep-G2 128 bp overlap
ISX 1 dataset
ChIP HepG2 ENCFF878QAY 83 bp overlap
JUNB 1 dataset
ChIP HAEC GSE89970.JUNB.HAEC 215 bp overlap
KDM1A 1 dataset
ChIP HepG2 ENCFF240UWG 293 bp overlap
KLF15 1 dataset
ChIP HepG2 ENCFF282HUB 60 bp overlap
KLF17 2 datasets
ChIP HEK293 ENCFF658MHR 172 bp overlap
ChIP HEK293 ENCSR065WUF.KLF17.HEK293 117 bp overlap
KMT2A 11 datasets
ChIP CCRF-CEM GSE83671.KMT2A.CCRF-CEM 302 bp overlap
ChIP HEK293T_N-term GSE90762.KMT2A.HEK293T_N-term 302 bp overlap
ChIP HEK293T_N-term_shCtrl GSE90762.KMT2A.HEK293T_N-term_shCtrl 154 bp overlap
ChIP HEK293T_N-term_shCtrl_TASP1-KO GSE90762.KMT2A.HEK293T_N-term_shCtrl_TASP1-KO 302 bp overlap
ChIP IMS-M2_DMSO GSE129636.KMT2A.IMS-M2_DMSO 302 bp overlap
ChIP MOLM-13 GSE114981.KMT2A.MOLM-13 219 bp overlap
ChIP MOLM-13_CBS79-KO-VP-HOTTIP GSE114981.KMT2A.MOLM-13_CBS79-KO-VP-HOTTIP 302 bp overlap
ChIP MV4-11 GSE83671.KMT2A.MV4-11 302 bp overlap
ChIP OCI-AML-3_DMSO GSE129636.KMT2A.OCI-AML-3_DMSO 302 bp overlap
ChIP RS4-11_DMSO-D3-180402 GSE127507.KMT2A.RS4-11_DMSO-D3-180402 265 bp overlap
ChIP SEM GSE83671.KMT2A.SEM 302 bp overlap
MAX 1 dataset
ChIP MDA-MB-468 GSE81381.MAX.MDA-MB-468 79 bp overlap
MAZ 1 dataset
ChIP HepG2 ENCFF068NYH 132 bp overlap
MED1 8 datasets
ChIP AML GSE154985.MED1.AML 147 bp overlap
ChIP HCT-116 GSE121798.MED1.HCT-116 113 bp overlap
ChIP HCT-116_shLuc GSE121798.MED1.HCT-116_shLuc 251 bp overlap
ChIP HCT-116_shLuc_thaps GSE121798.MED1.HCT-116_shLuc_thaps 245 bp overlap
ChIP HCT-116_thaps GSE121798.MED1.HCT-116_thaps 275 bp overlap
ChIP HUVEC-C_siERG GSE124892.MED1.HUVEC-C_siERG 302 bp overlap
ChIP MOLM-13 GSE154985.MED1.MOLM-13 230 bp overlap
ChIP cardiomyocyte_7 GSE85628.MED1.cardiomyocyte_7 74 bp overlap
MEF2B 1 dataset
ChIP tonsil GSE110682.MEF2B.tonsil 185 bp overlap
MEIS1 1 dataset
ChIP HepG2 ENCFF706DID 302 bp overlap
MEN1 2 datasets
ChIP OCI-AML-3_DMSO GSE129636.MEN1.OCI-AML-3_DMSO 302 bp overlap
ChIP SEM GSE83671.MEN1.SEM 185 bp overlap
MIER1 1 dataset
ChIP K-562 ENCSR426MDV.MIER1.K-562 199 bp overlap
MLLT1 2 datasets
ChIP K-562 ENCSR107GRP.MLLT1.K-562 265 bp overlap
ChIP K-562 ENCSR675LRO.MLLT1.K-562 203 bp overlap
MLX 1 dataset
ChIP Hep-G2 ENCSR125DAD.MLX.Hep-G2 145 bp overlap
MNT 1 dataset
ChIP K-562 ENCSR390VGH.MNT.K-562 64 bp overlap
MXI1 1 dataset
ChIP IMR-90 ENCFF040YVH 97 bp overlap
MYCN 3 datasets
ChIP Kelly GSE94822.MYCN.Kelly 138 bp overlap
ChIP LA-N-5 GSE138295.MYCN.LA-N-5 149 bp overlap
ChIP LNCaP_EtOH GSE117304.MYCN.LNCaP_EtOH 251 bp overlap
NANOG 1 dataset
ChIP WA01_3IL ERP004238.NANOG.WA01_3IL 177 bp overlap
NBN 3 datasets
ChIP GM12878 ENCFF213ZNN 252 bp overlap
ChIP GM12878 ENCSR278SQL.NBN.GM12878 284 bp overlap
ChIP K-562 ENCSR085QEV.NBN.K-562 127 bp overlap
NFATC1 1 dataset
ChIP HUVEC-C_VEGF GSE49428.NFATC1.HUVEC-C_VEGF 244 bp overlap
NIPBL 1 dataset
ChIP HCT-116_RAD21-mAC_500uM_auxin GSE104888.NIPBL.HCT-116_RAD21-mAC_500uM_auxin 165 bp overlap
NRF1 1 dataset
ChIP HeLa-S3 GSE108856.NRF1.HeLa-S3 202 bp overlap
NUTM1 1 dataset
ChIP embryonic-kidney_induced GSE133122.NUTM1.embryonic-kidney_induced 302 bp overlap
PGR 2 datasets
ChIP breast_tumor_Male_26 GSE104399.PGR.breast_tumor_Male_26 289 bp overlap
ChIP breast_tumor_Male_30 GSE104399.PGR.breast_tumor_Male_30 61 bp overlap
PHIP 4 datasets
ChIP HCT-116_MLL1-KO__ab833 GSE101646.PHIP.HCT-116_MLL1-KO__ab833 302 bp overlap
ChIP HCT-116_MLL4-SET-KO_ab833 GSE101646.PHIP.HCT-116_MLL4-SET-KO_ab833 302 bp overlap
ChIP HCT-116_ab833 GSE101646.PHIP.HCT-116_ab833 302 bp overlap
ChIP HCT-116_ab834 GSE101646.PHIP.HCT-116_ab834 302 bp overlap
POLR2A 12 datasets
ChIP HeLa-S3 ENCFF773DNG 135 bp overlap
ChIP IMR-90 ENCFF672YWV 302 bp overlap
ChIP K562 ENCFF419GHN 77 bp overlap
ChIP body of pancreas ENCFF675RCN 141 bp overlap
ChIP breast epithelium ENCFF045XXN 73 bp overlap
ChIP breast epithelium ENCFF955FMX 159 bp overlap
ChIP endothelial cell of umbilical vein ENCFF091YHT 105 bp overlap
ChIP erythroblast ENCFF498VMR 202 bp overlap
ChIP gastrocnemius medialis ENCFF145VIB 153 bp overlap
ChIP gastroesophageal sphincter ENCFF826EZZ 202 bp overlap
ChIP prostate gland ENCFF832RQK 81 bp overlap
ChIP vagina ENCFF305NWS 76 bp overlap
RBBP5 1 dataset
ChIP K-562 ENCSR000AQI.RBBP5.K-562 302 bp overlap
RELA 3 datasets
ChIP SW480_16h_TNFa GSE102796.RELA.SW480_16h_TNFa 273 bp overlap
ChIP aortic-endothelial-cell_IL1B_D25 GSE139377.RELA.aortic-endothelial-cell_IL1B_D25 137 bp overlap
ChIP aortic-endothelial-cell_IL1B_D47 GSE139377.RELA.aortic-endothelial-cell_IL1B_D47 95 bp overlap
RFX1 1 dataset
ChIP HepG2 ENCFF144SCF 126 bp overlap
SAP30 1 dataset
ChIP WA01 ENCSR000ATR.SAP30.WA01 302 bp overlap
SIN3A 7 datasets
ChIP A-549 ENCSR513XQX.SIN3A.A-549 188 bp overlap
ChIP A549 ENCFF752ATT 203 bp overlap
ChIP K-562 ENCSR920BLG.SIN3A.K-562 181 bp overlap
ChIP Panc1 ENCFF898EEQ 248 bp overlap
ChIP SK-N-SH ENCSR000BPB.SIN3A.SK-N-SH 160 bp overlap
ChIP WA01 ENCSR000BIS.SIN3A.WA01 165 bp overlap
ChIP WA01 ENCSR000EBO.SIN3A.WA01 141 bp overlap
SMARCA4 13 datasets
ChIP 22Rv1_PTEN-KO GSE115615.SMARCA4.22Rv1_PTEN-KO 183 bp overlap
ChIP 501-mel_SHCTR GSE61965.SMARCA4.501-mel_SHCTR 238 bp overlap
ChIP A-549_AG15684 GSE132290.SMARCA4.A-549_AG15684 273 bp overlap
ChIP A-549_AG15685 GSE132290.SMARCA4.A-549_AG15685 167 bp overlap
ChIP Aska-SS_shSSX GSE108025.SMARCA4.Aska-SS_shSSX 302 bp overlap
ChIP BIN-67_lentivirus-SMARCA4 GSE117734.SMARCA4.BIN-67_lentivirus-SMARCA4 132 bp overlap
ChIP BIN-67_lentivirus-SMARCA4-K785R GSE117734.SMARCA4.BIN-67_lentivirus-SMARCA4-K785R 302 bp overlap
ChIP MCF-7 GSE140185.SMARCA4.MCF-7 174 bp overlap
ChIP NPC_K755R-siCT GSE122631.SMARCA4.NPC_K755R-siCT 118 bp overlap
ChIP TOV-21G GSE110448.SMARCA4.TOV-21G 122 bp overlap
ChIP TTC-1240_SMARCB1-FL GSE124903.SMARCA4.TTC-1240_SMARCB1-FL 239 bp overlap
ChIP TTC-1240_delC GSE124903.SMARCA4.TTC-1240_delC 246 bp overlap
ChIP hiPSC_WT GSE124903.SMARCA4.hiPSC_WT 189 bp overlap
SMARCB1 1 dataset
ChIP proliferating-human-fibroblast GSE131711.SMARCB1.proliferating-human-fibroblast 135 bp overlap
SMARCC1 2 datasets
ChIP Aska-SS_shSSX GSE108025.SMARCC1.Aska-SS_shSSX 302 bp overlap
ChIP HeLa-S3 ENCFF971JGA 199 bp overlap
SNAI2 1 dataset
ChIP PC-9_2DF GSE131687.SNAI2.PC-9_2DF 302 bp overlap
SP7 2 datasets
ChIP HEK293 ENCFF733RBE 237 bp overlap
ChIP HEK293 ENCSR468IJT.SP7.HEK293 302 bp overlap
SPI1 1 dataset
ChIP CTV-1_delQ GSE128835.SPI1.CTV-1_delQ 161 bp overlap
TAF1 5 datasets
ChIP HeLa-S3 ENCFF556LCN 107 bp overlap
ChIP HeLa-S3 ENCSR000BHT.TAF1.HeLa-S3 157 bp overlap
ChIP K-562 ENCSR000BKS.TAF1.K-562 137 bp overlap
ChIP SK-N-SH ENCSR000BQF.TAF1.SK-N-SH 81 bp overlap
ChIP WA01 ENCSR000BHO.TAF1.WA01 285 bp overlap
TBX5 2 datasets
ChIP G296S GSE85628.TBX5.G296S 199 bp overlap
ChIP G296S_2 GSE85628.TBX5.G296S_2 199 bp overlap
TCF3 1 dataset
ChIP Kasumi-1 GSE43834.TCF3.Kasumi-1 187 bp overlap
TP53 1 dataset
ChIP HepG2 ENCFF687JDU 55 bp overlap
UBN1 1 dataset
ChIP HeLa GSE45024.UBN1.HeLa 241 bp overlap
UBTF 1 dataset
ChIP K562 ENCFF174SPM 191 bp overlap
WDR5 3 datasets
ChIP K-562_DMSO GSE115377.WDR5.K-562_DMSO 53 bp overlap
ChIP breast-cancer_shLuc GSE113279.WDR5.breast-cancer_shLuc 302 bp overlap
ChIP breast-cancer_shWDR5 GSE113279.WDR5.breast-cancer_shWDR5 206 bp overlap
WT1 2 datasets
ChIP HEK293 ENCFF906HIR 247 bp overlap
ChIP HEK293 ENCSR966PJJ.WT1.HEK293 198 bp overlap
YY1 2 datasets
ChIP ALL GSE145549.YY1.ALL 200 bp overlap
ChIP Hep-G2_RBM25-KD GSE120104.YY1.Hep-G2_RBM25-KD 55 bp overlap
ZBED4 1 dataset
ChIP HepG2 ENCFF157CDZ 148 bp overlap
ZBTB17 1 dataset
ChIP HEK293 ENCFF865LIO 221 bp overlap
ZEB2 2 datasets
ChIP HEK293 ENCFF847JIE 282 bp overlap
ChIP HEK293 ENCSR417VWF.ZEB2.HEK293 204 bp overlap
ZFP64 1 dataset
ChIP HEK293 GSE76494.ZFP64.HEK293 202 bp overlap
ZNF133 1 dataset
ChIP HEK293T GSE78099.ZNF133.HEK293T 153 bp overlap
ZNF2 1 dataset
ChIP HEK293 ENCSR011CKE.ZNF2.HEK293 148 bp overlap
ZNF263 2 datasets
ChIP HEK293 ENCFF336CWQ 79 bp overlap
ChIP HEK293T GSE78099.ZNF263.HEK293T 181 bp overlap
ZNF394 3 datasets
ChIP HEK293 ENCFF236OPX 129 bp overlap
ChIP HEK293 ENCFF236OPX 189 bp overlap
ChIP HEK293 ENCSR125DNC.ZNF394.HEK293 302 bp overlap
ZNF44 1 dataset
ChIP Hep-G2 ENCSR914HPP.ZNF44.Hep-G2 188 bp overlap
ZNF501 1 dataset
ChIP HEK293 ENCFF066RAQ 187 bp overlap
ZNF614 1 dataset
ChIP HepG2 ENCFF677IUD 70 bp overlap
ZNF629 2 datasets
ChIP HEK293 ENCFF096ELQ 81 bp overlap
ChIP HEK293 ENCSR351NON.ZNF629.HEK293 189 bp overlap
ZSCAN5C 2 datasets
ChIP HEK293 ENCFF343DTU 97 bp overlap
ChIP HEK293 ENCSR731AGO.ZSCAN5C.HEK293 184 bp overlap
ZXDB 1 dataset
ChIP HEK293 ENCSR559IOZ.ZXDB.HEK293 100 bp overlap