chr5 : 67,780,166 – 67,780,843
677 bp 136 TFs 0 linked genes
This 677 bp open chromatin element has no linked target genes and is bound by 136 transcription factors.
Linked Genes
No linked genes
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr5:67,775,166 – 67,785,843
This element Other elements ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
136 transcription factors
Source
Cell type
ARID5B 1 dataset
ChIP HepG2 ENCFF964FWK 220 bp overlap
ASH2L 1 dataset
ChIP H1 ENCFF399KAM 649 bp overlap
BRD4 1 dataset
ChIP hESC GSE33281.BRD4.hESC 66 bp overlap
CTCF 2 datasets
Motif DE_12h DE_12h-CTCF_MA1929.2 31 bp overlap
ChIP HEK293 ENCSR617IFZ.CTCF.HEK293 131 bp overlap
CTCFL 1 dataset
Motif DE_12h DE_12h-CTCFL_MA1102.3 8 bp overlap
EGR2 1 dataset
ChIP HEK293 ENCFF336LFH 147 bp overlap
ETV1 2 datasets
Motif DE_12h DE_12h-ETV1_MA0761.3 9 bp overlap
Motif ES_0h ES_0h-ETV1_MA0761.3 9 bp overlap
ETV6 2 datasets
ChIP WTC11 ENCFF812SCD 228 bp overlap
ChIP WTC11 ENCFF812SCD 437 bp overlap
FEZF2 2 datasets
Motif DE_12h DE_12h-FEZF2_MA2341.1 8 bp overlap
Motif ES_0h ES_0h-FEZF2_MA2341.1 8 bp overlap
FOXA1 5 datasets
Motif DE_12h DE_12h-FOXA1_MA0148.5 8 bp overlap
Motif ES_0h ES_0h-FOXA1_MA0148.5 8 bp overlap
ChIP HepG2 ENCFF207NVJ 98 bp overlap
ChIP HepG2 ENCFF361KNY 82 bp overlap
ChIP HepG2 ENCFF740VZW 87 bp overlap
FOXA2 7 datasets
ChIP DE DE-FOXA2-1 531 bp overlap
ChIP DE DE-FOXA2-2 445 bp overlap
Motif DE_12h DE_12h-FOXA2_MA0047.4 8 bp overlap
Motif ES_0h ES_0h-FOXA2_MA0047.4 8 bp overlap
ChIP HepG2 ENCFF533COJ 297 bp overlap
ChIP HepG2 ENCFF570ABM 203 bp overlap
ChIP HepG2 ENCFF894AYY 113 bp overlap
FOXA3 4 datasets
Motif DE_12h DE_12h-FOXA3_MA1683.2 7 bp overlap
Motif ES_0h ES_0h-FOXA3_MA1683.2 7 bp overlap
ChIP HepG2 ENCFF005KGL 156 bp overlap
ChIP K562 ENCFF781VSC 341 bp overlap
FOXB1 2 datasets
Motif DE_12h DE_12h-FOXB1_MA0845.1 11 bp overlap
Motif ES_0h ES_0h-FOXB1_MA0845.1 11 bp overlap
FOXC1 2 datasets
Motif DE_12h DE_12h-FOXC1_MA0032.2 11 bp overlap
Motif ES_0h ES_0h-FOXC1_MA0032.2 11 bp overlap
FOXC2 2 datasets
Motif DE_12h DE_12h-FOXC2_MA0846.2 11 bp overlap
Motif ES_0h ES_0h-FOXC2_MA0846.2 11 bp overlap
FOXD1 2 datasets
Motif DE_12h DE_12h-FOXD1_MA0031.2 7 bp overlap
Motif ES_0h ES_0h-FOXD1_MA0031.2 7 bp overlap
FOXD2 2 datasets
Motif DE_12h DE_12h-FOXD2_MA0847.4 11 bp overlap
Motif ES_0h ES_0h-FOXD2_MA0847.4 11 bp overlap
FOXD3 2 datasets
Motif DE_12h DE_12h-FOXD3_MA0041.3 14 bp overlap
Motif ES_0h ES_0h-FOXD3_MA0041.3 14 bp overlap
FOXE1 2 datasets
Motif DE_12h DE_12h-FOXE1_MA1487.3 12 bp overlap
Motif ES_0h ES_0h-FOXE1_MA1487.3 12 bp overlap
FOXF2 2 datasets
Motif DE_12h DE_12h-FOXF2_MA0030.2 9 bp overlap
Motif ES_0h ES_0h-FOXF2_MA0030.2 9 bp overlap
FOXG1 2 datasets
Motif DE_12h DE_12h-FOXG1_MA0613.1 8 bp overlap
Motif ES_0h ES_0h-FOXG1_MA0613.1 8 bp overlap
FOXI1 2 datasets
Motif DE_12h DE_12h-FOXI1_MA0042.2 7 bp overlap
Motif ES_0h ES_0h-FOXI1_MA0042.2 7 bp overlap
FOXK1 3 datasets
Motif DE_12h DE_12h-FOXK1_MA0852.3 7 bp overlap
Motif ES_0h ES_0h-FOXK1_MA0852.3 7 bp overlap
ChIP WTC11 ENCFF875IGU 391 bp overlap
FOXK2 2 datasets
Motif DE_12h DE_12h-FOXK2_MA1103.3 7 bp overlap
Motif ES_0h ES_0h-FOXK2_MA1103.3 7 bp overlap
FOXL1 2 datasets
Motif DE_12h DE_12h-FOXL1_MA0033.2 7 bp overlap
Motif ES_0h ES_0h-FOXL1_MA0033.2 7 bp overlap
FOXN3 2 datasets
Motif DE_12h DE_12h-FOXN3_MA1489.1 8 bp overlap
Motif ES_0h ES_0h-FOXN3_MA1489.1 8 bp overlap
FOXO4 2 datasets
Motif DE_12h DE_12h-FOXO4_MA0848.1 7 bp overlap
Motif ES_0h ES_0h-FOXO4_MA0848.1 7 bp overlap
FOXO6 2 datasets
Motif DE_12h DE_12h-FOXO6_MA0849.1 7 bp overlap
Motif ES_0h ES_0h-FOXO6_MA0849.1 7 bp overlap
FOXP1 2 datasets
Motif DE_12h DE_12h-FOXP1_MA0481.4 7 bp overlap
Motif ES_0h ES_0h-FOXP1_MA0481.4 7 bp overlap
FOXP2 2 datasets
Motif DE_12h DE_12h-FOXP2_MA0593.2 9 bp overlap
Motif ES_0h ES_0h-FOXP2_MA0593.2 9 bp overlap
FOXP3 2 datasets
Motif DE_12h DE_12h-FOXP3_MA0850.1 7 bp overlap
Motif ES_0h ES_0h-FOXP3_MA0850.1 7 bp overlap
FOXP4 2 datasets
Motif DE_12h DE_12h-FOXP4_MA2117.1 7 bp overlap
Motif ES_0h ES_0h-FOXP4_MA2117.1 7 bp overlap
FOXS1 2 datasets
Motif DE_12h DE_12h-FOXS1_MA2118.1 8 bp overlap
Motif ES_0h ES_0h-FOXS1_MA2118.1 8 bp overlap
Foxf1 2 datasets
Motif DE_12h DE_12h-Foxf1_MA1606.2 7 bp overlap
Motif ES_0h ES_0h-Foxf1_MA1606.2 7 bp overlap
Foxj2 2 datasets
Motif DE_12h DE_12h-Foxj2_MA0614.1 8 bp overlap
Motif ES_0h ES_0h-Foxj2_MA0614.1 8 bp overlap
Foxj3 2 datasets
Motif DE_12h DE_12h-Foxj3_MA0851.2 9 bp overlap
Motif ES_0h ES_0h-Foxj3_MA0851.2 9 bp overlap
Foxl2 2 datasets
Motif DE_12h DE_12h-Foxl2_MA1607.2 10 bp overlap
Motif ES_0h ES_0h-Foxl2_MA1607.2 10 bp overlap
Foxo1 2 datasets
Motif DE_12h DE_12h-Foxo1_MA0480.3 7 bp overlap
Motif ES_0h ES_0h-Foxo1_MA0480.3 7 bp overlap
Foxo3 2 datasets
Motif DE_12h DE_12h-Foxo3_MA0157.4 7 bp overlap
Motif ES_0h ES_0h-Foxo3_MA0157.4 7 bp overlap
GATAD2A 1 dataset
ChIP HepG2 ENCFF252XNH 465 bp overlap
GLIS1 1 dataset
ChIP HEK293 ENCFF299RSE 195 bp overlap
GLIS2 1 dataset
ChIP HEK293 ENCFF446EIF 511 bp overlap
IKZF2 2 datasets
Motif DE_12h DE_12h-IKZF2_MA2326.1 6 bp overlap
Motif ES_0h ES_0h-IKZF2_MA2326.1 6 bp overlap
IKZF3 1 dataset
ChIP HEK293 ENCFF518OXG 215 bp overlap
INSM1 3 datasets
Motif DE_12h DE_12h-INSM1_MA0155.1 12 bp overlap
Motif DE_24h DE_24h-INSM1_MA0155.1 12 bp overlap
Motif ES_0h ES_0h-INSM1_MA0155.1 12 bp overlap
Ikzf3 2 datasets
Motif DE_12h DE_12h-Ikzf3_MA1992.2 9 bp overlap
Motif ES_0h ES_0h-Ikzf3_MA1992.2 9 bp overlap
JUN 1 dataset
ChIP ESC S34-ESC-d0-JUN-exp2 313 bp overlap
KLF1 1 dataset
ChIP HEK293 ENCFF159QSW 241 bp overlap
KLF10 3 datasets
Motif DE_12h DE_12h-KLF10_MA1511.2 9 bp overlap
Motif ES_0h ES_0h-KLF10_MA1511.2 9 bp overlap
ChIP HEK293 ENCFF326EGX 491 bp overlap
KLF12 2 datasets
Motif DE_12h DE_12h-KLF12_MA0742.2 9 bp overlap
Motif ES_0h ES_0h-KLF12_MA0742.2 9 bp overlap
KLF13 1 dataset
Motif DE_12h DE_12h-KLF13_MA0657.2 17 bp overlap
KLF14 2 datasets
Motif DE_12h DE_12h-KLF14_MA0740.2 9 bp overlap
Motif ES_0h ES_0h-KLF14_MA0740.2 9 bp overlap
KLF16 2 datasets
Motif DE_12h DE_12h-KLF16_MA0741.1 11 bp overlap
Motif ES_0h ES_0h-KLF16_MA0741.1 11 bp overlap
KLF17 1 dataset
ChIP HEK293 ENCFF658MHR 306 bp overlap
KLF4 2 datasets
Motif DE_12h DE_12h-KLF4_MA0039.5 8 bp overlap
Motif ES_0h ES_0h-KLF4_MA0039.5 8 bp overlap
KLF5 4 datasets
Motif DE_12h DE_12h-KLF5_MA0599.1 10 bp overlap
Motif DE_12h DE_12h-KLF5_MA0599.1 10 bp overlap
Motif ES_0h ES_0h-KLF5_MA0599.1 10 bp overlap
Motif ES_0h ES_0h-KLF5_MA0599.1 10 bp overlap
KLF7 1 dataset
ChIP HEK293 ENCFF599UKL 357 bp overlap
MAF 2 datasets
Motif DE_12h DE_12h-MAF_MA1520.2 13 bp overlap
Motif DE_24h DE_24h-MAF_MA1520.2 13 bp overlap
MAZ 2 datasets
Motif DE_12h DE_12h-MAZ_MA1522.2 8 bp overlap
Motif ES_0h ES_0h-MAZ_MA1522.2 8 bp overlap
MSANTD3 1 dataset
Motif DE_12h DE_12h-MSANTD3_MA1523.2 7 bp overlap
MYB 2 datasets
Motif DE_12h DE_12h-MYB_MA0100.4 6 bp overlap
Motif ES_0h ES_0h-MYB_MA0100.4 6 bp overlap
NCOA2 1 dataset
ChIP HepG2 ENCFF853BJJ 451 bp overlap
NFIC 1 dataset
ChIP K562 ENCFF167YID 68 bp overlap
NRF1 1 dataset
ChIP K562 ENCFF791UHF 561 bp overlap
Nanog 2 datasets
Motif DE_12h DE_12h-Nanog_MA2339.1 7 bp overlap
Motif DE_24h DE_24h-Nanog_MA2339.1 7 bp overlap
OSR2 1 dataset
ChIP HEK293 ENCFF875BDB 421 bp overlap
PATZ1 3 datasets
Motif DE_12h DE_12h-PATZ1_MA1961.2 11 bp overlap
Motif ES_0h ES_0h-PATZ1_MA1961.2 11 bp overlap
ChIP HEK293 ENCFF016MNJ 521 bp overlap
PAXIP1 1 dataset
ChIP HepG2 ENCFF526NOJ 551 bp overlap
PKNOX1 2 datasets
ChIP HEK293T ENCFF174WDB 91 bp overlap
ChIP K562 ENCFF236IUS 81 bp overlap
POGZ 1 dataset
ChIP HepG2 ENCFF153UUK 517 bp overlap
POU5F1 1 dataset
ChIP DE_D1 DED1-OCT4_Batch_II 350 bp overlap
POU5F1_M 1 dataset
ChIP DE_D1 DED1-OCT4-M_Batch_II 520 bp overlap
PRDM1 1 dataset
ChIP HEK293 ENCFF302TBP 263 bp overlap
PRDM10 1 dataset
ChIP HEK293 ENCFF145WQQ 269 bp overlap
REST 2 datasets
ChIP HEK293 ENCFF073DOT 394 bp overlap
ChIP MCF-7 ENCFF893RRD 345 bp overlap
SALL1 1 dataset
ChIP HepG2 ENCFF426MCK 497 bp overlap
SMAD2_3 3 datasets
ChIP DE_D1 S39-DE-d1-SMAD2_3-exp2 352 bp overlap
ChIP DE_D2 S01-DE-d2-SMAD2_3-exp1 260 bp overlap
ChIP DE_D2 S01-DE-d2-SMAD2_3-exp1 221 bp overlap
SOX10 2 datasets
Motif DE_12h DE_12h-SOX10_MA0442.3 6 bp overlap
Motif ES_0h ES_0h-SOX10_MA0442.3 6 bp overlap
SOX13 1 dataset
ChIP HepG2 ENCFF062VSQ 357 bp overlap
SOX17_M 1 dataset
ChIP DE_D2 DED2-SOX17-M_Batch_II 584 bp overlap
SOX4 2 datasets
Motif DE_12h DE_12h-SOX4_MA0867.3 8 bp overlap
Motif ES_0h ES_0h-SOX4_MA0867.3 8 bp overlap
SP1 3 datasets
Motif DE_12h DE_12h-SP1_MA0079.5 9 bp overlap
Motif ES_0h ES_0h-SP1_MA0079.5 9 bp overlap
ChIP H1 ENCFF263FUH 321 bp overlap
SP2 2 datasets
Motif DE_12h DE_12h-SP2_MA0516.3 9 bp overlap
Motif ES_0h ES_0h-SP2_MA0516.3 9 bp overlap
SP3 2 datasets
Motif DE_12h DE_12h-SP3_MA0746.3 11 bp overlap
Motif ES_0h ES_0h-SP3_MA0746.3 11 bp overlap
SP4 2 datasets
Motif DE_12h DE_12h-SP4_MA0685.2 9 bp overlap
Motif ES_0h ES_0h-SP4_MA0685.2 9 bp overlap
SP5 3 datasets
Motif DE_12h DE_12h-SP5_MA1965.2 6 bp overlap
Motif ES_0h ES_0h-SP5_MA1965.2 6 bp overlap
ChIP HepG2 ENCFF931FHV 351 bp overlap
SP8 3 datasets
Motif DE_12h DE_12h-SP8_MA0747.2 11 bp overlap
Motif DE_24h DE_24h-SP8_MA0747.2 11 bp overlap
Motif ES_0h ES_0h-SP8_MA0747.2 11 bp overlap
Sox11 2 datasets
Motif DE_12h DE_12h-Sox11_MA0869.3 8 bp overlap
Motif ES_0h ES_0h-Sox11_MA0869.3 8 bp overlap
TCF12 1 dataset
ChIP WA01 ENCSR000BIT.TCF12.WA01 111 bp overlap
TCF7L1 2 datasets
Motif DE_12h DE_12h-TCF7L1_MA1421.1 12 bp overlap
Motif DE_24h DE_24h-TCF7L1_MA1421.1 12 bp overlap
TFAP4 1 dataset
ChIP HepG2 ENCFF932XOY 397 bp overlap
THAP1 1 dataset
Motif DE_12h DE_12h-THAP1_MA0597.3 8 bp overlap
WT1 1 dataset
ChIP HEK293 ENCFF906HIR 287 bp overlap
Wt1 3 datasets
Motif DE_12h DE_12h-Wt1_MA1627.2 10 bp overlap
Motif DE_24h DE_24h-Wt1_MA1627.2 10 bp overlap
Motif ES_0h ES_0h-Wt1_MA1627.2 10 bp overlap
YY1 3 datasets
ChIP HEK293 ENCFF734SBY 249 bp overlap
ChIP Hep-G2_NC GSE120104.YY1.Hep-G2_NC 111 bp overlap
ChIP Hep-G2_RBM25-KD GSE120104.YY1.Hep-G2_RBM25-KD 54 bp overlap
ZBTB17 2 datasets
ChIP HEK293 ENCFF865LIO 361 bp overlap
ChIP HEK293 ENCFF865LIO 361 bp overlap
ZBTB26 3 datasets
ChIP HEK293 ENCFF752POA 677 bp overlap
ChIP HEK293 ENCFF752POA 491 bp overlap
ChIP HEK293 ENCFF752TCU 675 bp overlap
ZBTB33 2 datasets
ChIP HepG2 ENCFF778UKV 337 bp overlap
ChIP HepG2 ENCFF778UKV 280 bp overlap
ZBTB48 2 datasets
ChIP HEK293 ENCFF809BPK 324 bp overlap
ChIP HEK293 ENCSR781EQJ.ZBTB48.HEK293 177 bp overlap
ZBTB6 2 datasets
Motif DE_12h DE_12h-ZBTB6_MA1581.2 9 bp overlap
Motif ES_0h ES_0h-ZBTB6_MA1581.2 9 bp overlap
ZBTB8A 3 datasets
ChIP HEK293 ENCFF303WRD 282 bp overlap
ChIP HEK293 ENCFF303WRD 367 bp overlap
ChIP HEK293 ENCSR481FEC.ZBTB8A.HEK293 182 bp overlap
ZEB2 2 datasets
ChIP HEK293 ENCFF847JIE 446 bp overlap
ChIP HEK293 ENCFF847JIE 551 bp overlap
ZFHX2 1 dataset
ChIP HEK293 ENCFF167TUA 184 bp overlap
ZFP69B 1 dataset
ChIP HEK293 ENCFF942LFP 201 bp overlap
ZGPAT 1 dataset
ChIP HepG2 ENCFF055YSO 252 bp overlap
ZIC2 1 dataset
ChIP HEK293 ENCFF033NQQ 422 bp overlap
ZKSCAN3 2 datasets
Motif DE_12h DE_12h-ZKSCAN3_MA1973.2 14 bp overlap
Motif ES_0h ES_0h-ZKSCAN3_MA1973.2 14 bp overlap
ZMYM4 1 dataset
ChIP HepG2 ENCFF567SQY 551 bp overlap
ZNF135 1 dataset
Motif DE_12h DE_12h-ZNF135_MA1587.1 14 bp overlap
ZNF148 3 datasets
Motif DE_12h DE_12h-ZNF148_MA1653.2 10 bp overlap
Motif ES_0h ES_0h-ZNF148_MA1653.2 10 bp overlap
ChIP K562 ENCFF352SDL 590 bp overlap
ZNF189 1 dataset
ChIP HEK293 ENCFF638TIB 422 bp overlap
ZNF213 2 datasets
Motif DE_12h DE_12h-ZNF213_MA2121.1 12 bp overlap
Motif ES_0h ES_0h-ZNF213_MA2121.1 12 bp overlap
ZNF281 6 datasets
Motif DE_12h DE_12h-ZNF281_MA1630.3 10 bp overlap
Motif DE_12h DE_12h-ZNF281_MA1630.3 10 bp overlap
Motif ES_0h ES_0h-ZNF281_MA1630.3 10 bp overlap
ChIP HepG2 ENCFF585QNU 246 bp overlap
ChIP K562 ENCFF594VNM 390 bp overlap
ChIP WTC11 ENCFF551GAV 212 bp overlap
ZNF324 2 datasets
ChIP HEK293 ENCFF062DPE 260 bp overlap
ChIP HEK293 ENCFF062DPE 405 bp overlap
ZNF335 2 datasets
ChIP HEK293 ENCFF784SLD 324 bp overlap
ChIP HEK293 ENCSR328SUD.ZNF335.HEK293 158 bp overlap
ZNF341 2 datasets
ChIP HEK293 ENCFF944VMC 196 bp overlap
ChIP HEK293 ENCSR185FOY.ZNF341.HEK293 62 bp overlap
ZNF366 2 datasets
ChIP HEK293 ENCFF799ATK 195 bp overlap
ChIP HEK293 ENCSR106EBH.ZNF366.HEK293 59 bp overlap
ZNF423 1 dataset
ChIP HEK293 ENCFF937QHI 274 bp overlap
ZNF501 1 dataset
ChIP HEK293 ENCFF066RAQ 311 bp overlap
ZNF518A 1 dataset
ChIP HEK293 ENCFF892ULS 398 bp overlap
ZNF530 3 datasets
Motif DE_12h DE_12h-ZNF530_MA1981.2 14 bp overlap
Motif DE_24h DE_24h-ZNF530_MA1981.2 14 bp overlap
Motif ES_0h ES_0h-ZNF530_MA1981.2 14 bp overlap
ZNF549 4 datasets
Motif DE_12h DE_12h-ZNF549_MA1728.2 8 bp overlap
Motif DE_24h DE_24h-ZNF549_MA1728.2 8 bp overlap
Motif ES_0h ES_0h-ZNF549_MA1728.2 8 bp overlap
ChIP HEK293 ENCFF528IUI 337 bp overlap
ZNF561 1 dataset
ChIP HEK293 ENCFF399XKF 57 bp overlap
ZNF574 1 dataset
Motif DE_12h DE_12h-ZNF574_MA1982.2 14 bp overlap
ZNF582 1 dataset
Motif DE_12h DE_12h-ZNF582_MA1983.2 19 bp overlap
ZNF600 1 dataset
ChIP HEK293 ENCFF785JSX 262 bp overlap
ZNF610 2 datasets
Motif DE_12h DE_12h-ZNF610_MA1713.2 10 bp overlap
Motif ES_0h ES_0h-ZNF610_MA1713.2 10 bp overlap
ZNF687 2 datasets
ChIP HepG2 ENCFF653WIX 527 bp overlap
ChIP HepG2 ENCFF653WIX 351 bp overlap
ZNF692 1 dataset
ChIP HEK293 ENCFF040AZE 228 bp overlap
ZNF766 2 datasets
ChIP HepG2 ENCFF774VLV 501 bp overlap
ChIP K562 ENCFF348LDO 533 bp overlap
ZNF93 3 datasets
Motif DE_12h DE_12h-ZNF93_MA1721.2 14 bp overlap
Motif DE_24h DE_24h-ZNF93_MA1721.2 14 bp overlap
Motif ES_0h ES_0h-ZNF93_MA1721.2 14 bp overlap
ZSCAN30 1 dataset
ChIP HEK293 ENCFF082YBI 337 bp overlap
Zfp335 1 dataset
Motif DE_12h DE_12h-Zfp335_MA2002.2 7 bp overlap
Zfp961 2 datasets
Motif DE_12h DE_12h-Zfp961_MA2126.1 8 bp overlap
Motif ES_0h ES_0h-Zfp961_MA2126.1 8 bp overlap
Zic1::Zic2 3 datasets
Motif DE_12h DE_12h-Zic1Zic2_MA1628.2 7 bp overlap
Motif DE_24h DE_24h-Zic1Zic2_MA1628.2 7 bp overlap
Motif ES_0h ES_0h-Zic1Zic2_MA1628.2 7 bp overlap
Zic3 3 datasets
Motif DE_12h DE_12h-Zic3_MA0697.3 7 bp overlap
Motif DE_24h DE_24h-Zic3_MA0697.3 7 bp overlap
Motif ES_0h ES_0h-Zic3_MA0697.3 7 bp overlap
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