chr15 : 97,177,029 – 97,177,985
956 bp 149 TFs 0 linked genes
This 956 bp open chromatin element has no linked target genes and is bound by 149 transcription factors.
Linked Genes
No linked genes
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr15:97,172,029 – 97,182,985
This element Other elements ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
149 transcription factors
Source
Cell type
AR 1 dataset
ChIP MCF-7 GSE48930.AR.MCF-7 279 bp overlap
ASH2L 3 datasets
ChIP H1 ENCFF399KAM 785 bp overlap
ChIP H1 ENCFF399KAM 671 bp overlap
ChIP WA01 ENCSR850KIP.ASH2L.WA01 956 bp overlap
Ahr::Arnt 1 dataset
Motif DE_12h DE_12h-AhrArnt_MA0006.2 5 bp overlap
Ar 2 datasets
Motif DE_12h DE_12h-Ar_MA0007.4 16 bp overlap
Motif ES_0h ES_0h-Ar_MA0007.4 16 bp overlap
BCL6B 2 datasets
Motif DE_12h DE_12h-BCL6B_MA0731.1 17 bp overlap
Motif ES_0h ES_0h-BCL6B_MA0731.1 17 bp overlap
BCOR 2 datasets
ChIP WA01_RNF2-R GSE104690.BCOR.WA01_RNF2-R 293 bp overlap
ChIP WA01_RNF2-R_Doxycyclin GSE104690.BCOR.WA01_RNF2-R_Doxycyclin 338 bp overlap
BHLHE22 1 dataset
Motif DE_12h DE_12h-BHLHE22_MA1635.2 6 bp overlap
BRD2 2 datasets
ChIP MDA-MB-231_BAZ2AsiRNA GSE116879.BRD2.MDA-MB-231_BAZ2AsiRNA 348 bp overlap
ChIP MDA-MB-231_BAZ2BsiRNA GSE116879.BRD2.MDA-MB-231_BAZ2BsiRNA 388 bp overlap
BRD4 3 datasets
ChIP Kelly_resistant GSE103030.BRD4.Kelly_resistant 313 bp overlap
ChIP RH4_shCHD4 GSE140115.BRD4.RH4_shCHD4 204 bp overlap
ChIP hESC GSE33281.BRD4.hESC 70 bp overlap
Bhlha15 1 dataset
Motif DE_12h DE_12h-Bhlha15_MA1472.3 8 bp overlap
CDX1 1 dataset
Motif DE_12h DE_12h-CDX1_MA0878.3 10 bp overlap
CDX2 1 dataset
Motif DE_12h DE_12h-CDX2_MA0465.3 8 bp overlap
CDX4 1 dataset
Motif DE_12h DE_12h-CDX4_MA1473.2 9 bp overlap
CEBPA 1 dataset
Motif DE_12h DE_12h-CEBPA_MA0102.5 10 bp overlap
CHD7 1 dataset
ChIP hiPSC_derived_lt-NES GSE108506.CHD7.hiPSC_derived_lt-NES 199 bp overlap
CTCF 4 datasets
Motif DE_12h DE_12h-CTCF_MA1930.2 33 bp overlap
Motif DE_12h DE_12h-CTCF_MA1930.2 33 bp overlap
Motif ES_0h ES_0h-CTCF_MA1930.2 33 bp overlap
ChIP erythroid GSE67783.CTCF.erythroid 270 bp overlap
CTNNB1 1 dataset
ChIP hiPSC_D2 GSE132532.CTNNB1.hiPSC_D2 315 bp overlap
Cebpa 2 datasets
ChIP BLaER1 ENCFF262VBH 251 bp overlap
ChIP BLaER1 ENCFF364PUR 251 bp overlap
DMRTA1 2 datasets
Motif DE_12h DE_12h-DMRTA1_MA1707.2 10 bp overlap
Motif ES_0h ES_0h-DMRTA1_MA1707.2 10 bp overlap
DMRTA2 2 datasets
Motif DE_12h DE_12h-DMRTA2_MA1478.2 6 bp overlap
Motif ES_0h ES_0h-DMRTA2_MA1478.2 6 bp overlap
Ddit3::Cebpa 1 dataset
Motif DE_12h DE_12h-Ddit3Cebpa_MA0019.2 10 bp overlap
E2F6 2 datasets
ChIP WA01 ENCSR000BSI.E2F6.WA01 116 bp overlap
ChIP WA01 ENCSR000BSI.E2F6.WA01 371 bp overlap
EED 1 dataset
ChIP ProEs GSE59087.EED.ProEs 154 bp overlap
EHF 1 dataset
Motif DE_12h DE_12h-EHF_MA0598.4 9 bp overlap
ELF1 1 dataset
Motif DE_12h DE_12h-ELF1_MA0473.4 9 bp overlap
ELF3 1 dataset
Motif DE_12h DE_12h-ELF3_MA0640.3 9 bp overlap
ELK4 1 dataset
Motif DE_12h DE_12h-ELK4_MA0076.3 9 bp overlap
ERG 1 dataset
ChIP VCaP_DHT GSE79128.ERG.VCaP_DHT 365 bp overlap
ETV1 1 dataset
Motif DE_12h DE_12h-ETV1_MA0761.3 9 bp overlap
ETV5::FIGLA 1 dataset
Motif DE_12h DE_12h-ETV5FIGLA_MA1945.2 14 bp overlap
EZH1 2 datasets
ChIP ProEs GSE59087.EZH1.ProEs 162 bp overlap
ChIP ProEs_SHCTR GSE59087.EZH1.ProEs_SHCTR 127 bp overlap
Ebf4 2 datasets
Motif DE_12h DE_12h-Ebf4_MA2122.1 11 bp overlap
Motif ES_0h ES_0h-Ebf4_MA2122.1 11 bp overlap
Erg 1 dataset
Motif DE_12h DE_12h-Erg_MA0474.4 10 bp overlap
FIGLA 1 dataset
Motif DE_12h DE_12h-FIGLA_MA0820.2 6 bp overlap
FOXA1 5 datasets
ChIP HEK293_i176m_TFS GSE123618.FOXA1.HEK293_i176m_TFS 311 bp overlap
ChIP HEK293_r261g_TFS GSE123618.FOXA1.HEK293_r261g_TFS 316 bp overlap
ChIP MCF-7_TNF GSE59530.FOXA1.MCF-7_TNF 72 bp overlap
ChIP T-47D_JC4747 GSE126004.FOXA1.T-47D_JC4747 176 bp overlap
ChIP breast_tumor_Male_3 GSE104399.FOXA1.breast_tumor_Male_3 409 bp overlap
FOXA2 1 dataset
ChIP pancreatic-progenitor_PP1 GSE104840.FOXA2.pancreatic-progenitor_PP1 204 bp overlap
FOXD3 1 dataset
Motif DE_12h DE_12h-FOXD3_MA0041.3 14 bp overlap
FOXP2 2 datasets
Motif DE_12h DE_12h-FOXP2_MA0593.2 9 bp overlap
Motif ES_0h ES_0h-FOXP2_MA0593.2 9 bp overlap
GABPA 1 dataset
Motif DE_12h DE_12h-GABPA_MA0062.4 10 bp overlap
GATA2 4 datasets
Motif DE_12h DE_12h-GATA2_MA0036.4 7 bp overlap
Motif ES_0h ES_0h-GATA2_MA0036.4 7 bp overlap
ChIP hiPSC_SLC6ebs GSE107639.GATA2.hiPSC_SLC6ebs 195 bp overlap
ChIP hiPSC_SLC9ebs GSE107639.GATA2.hiPSC_SLC9ebs 222 bp overlap
GATA4 3 datasets
Motif DE_12h DE_12h-GATA4_MA0482.3 8 bp overlap
Motif ES_0h ES_0h-GATA4_MA0482.3 8 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA4.pancreatic-progenitor_PP1 494 bp overlap
GATA5 2 datasets
Motif DE_12h DE_12h-GATA5_MA0766.3 8 bp overlap
Motif ES_0h ES_0h-GATA5_MA0766.3 8 bp overlap
GATA6 5 datasets
Motif DE_12h DE_12h-GATA6_MA1104.3 8 bp overlap
Motif ES_0h ES_0h-GATA6_MA1104.3 8 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 259 bp overlap
ChIP pancreatic-progenitor_KO GSE117136.GATA6.pancreatic-progenitor_KO 363 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA6.pancreatic-progenitor_PP1 462 bp overlap
Gata3 2 datasets
Motif DE_12h DE_12h-Gata3_MA0037.5 8 bp overlap
Motif ES_0h ES_0h-Gata3_MA0037.5 8 bp overlap
HDAC2 1 dataset
ChIP RH4_DMSO-6H GSE116344.HDAC2.RH4_DMSO-6H 205 bp overlap
HDAC3 1 dataset
ChIP RH4_DMSO-6H GSE116344.HDAC3.RH4_DMSO-6H 193 bp overlap
HOXB4 1 dataset
Motif DE_12h DE_12h-HOXB4_MA1499.2 6 bp overlap
HOXC13 1 dataset
Motif DE_12h DE_12h-HOXC13_MA0907.2 9 bp overlap
HOXC4 1 dataset
Motif DE_12h DE_12h-HOXC4_MA1504.2 6 bp overlap
HOXD4 1 dataset
Motif DE_12h DE_12h-HOXD4_MA1507.2 6 bp overlap
HOXD9 1 dataset
Motif DE_12h DE_12h-HOXD9_MA0913.3 9 bp overlap
HSF1 1 dataset
ChIP MCF-7_hyperthermia-15min GSE137558.HSF1.MCF-7_hyperthermia-15min 261 bp overlap
IKZF1 1 dataset
Motif DE_12h DE_12h-IKZF1_MA1508.2 8 bp overlap
INTS11 2 datasets
ChIP HL-60 GSE106359.INTS11.HL-60 165 bp overlap
ChIP HL-60 GSE106359.INTS11.HL-60 185 bp overlap
IRF3 1 dataset
Motif DE_12h DE_12h-IRF3_MA1418.2 17 bp overlap
Ikzf3 1 dataset
Motif DE_12h DE_12h-Ikzf3_MA1992.2 9 bp overlap
KLF10 1 dataset
Motif DE_12h DE_12h-KLF10_MA1511.2 9 bp overlap
KLF12 1 dataset
Motif DE_12h DE_12h-KLF12_MA0742.2 9 bp overlap
KLF14 1 dataset
Motif DE_12h DE_12h-KLF14_MA0740.2 9 bp overlap
KLF16 1 dataset
Motif DE_12h DE_12h-KLF16_MA0741.1 11 bp overlap
KLF5 2 datasets
Motif DE_12h DE_12h-KLF5_MA0599.1 10 bp overlap
ChIP HEK293_E419Q GSE88976.KLF5.HEK293_E419Q 376 bp overlap
MAFK 2 datasets
Motif DE_12h DE_12h-MAFK_MA0496.4 10 bp overlap
Motif ES_0h ES_0h-MAFK_MA0496.4 10 bp overlap
MAX 1 dataset
ChIP H1 ENCFF914VQY 357 bp overlap
MAZ 3 datasets
Motif DE_12h DE_12h-MAZ_MA1522.2 8 bp overlap
Motif DE_12h DE_12h-MAZ_MA1522.2 8 bp overlap
Motif ES_0h ES_0h-MAZ_MA1522.2 8 bp overlap
MEF2A 1 dataset
Motif DE_12h DE_12h-MEF2A_MA0052.5 10 bp overlap
MEF2C 1 dataset
Motif DE_12h DE_12h-MEF2C_MA0497.2 11 bp overlap
MRTFB 1 dataset
ChIP A-673-clone-Asp114 GSE92738.MRTFB.A-673-clone-Asp114 303 bp overlap
MYB 1 dataset
Motif DE_12h DE_12h-MYB_MA0100.4 6 bp overlap
MYF5 1 dataset
Motif DE_12h DE_12h-MYF5_MA1641.2 8 bp overlap
MYOG 1 dataset
ChIP RH4_DMSO-6H GSE116344.MYOG.RH4_DMSO-6H 245 bp overlap
Mecom 2 datasets
Motif DE_12h DE_12h-Mecom_MA0029.2 11 bp overlap
Motif ES_0h ES_0h-Mecom_MA0029.2 11 bp overlap
NANOG 1 dataset
ChIP WA09 GSE105028.NANOG.WA09 254 bp overlap
NEUROD1 1 dataset
Motif DE_12h DE_12h-NEUROD1_MA1109.2 8 bp overlap
NHLH2 2 datasets
Motif DE_12h DE_12h-NHLH2_MA1529.2 16 bp overlap
Motif ES_0h ES_0h-NHLH2_MA1529.2 16 bp overlap
NR3C1 2 datasets
Motif DE_12h DE_12h-NR3C1_MA0113.4 15 bp overlap
Motif ES_0h ES_0h-NR3C1_MA0113.4 15 bp overlap
NR3C2 2 datasets
Motif DE_12h DE_12h-NR3C2_MA0727.2 15 bp overlap
Motif ES_0h ES_0h-NR3C2_MA0727.2 15 bp overlap
Neurod2 2 datasets
Motif DE_12h DE_12h-Neurod2_MA0668.3 8 bp overlap
Motif DE_12h DE_12h-Neurod2_MA1993.2 6 bp overlap
Nr1h3::Rxra 2 datasets
Motif DE_12h DE_12h-Nr1h3Rxra_MA0494.2 16 bp overlap
Motif ES_0h ES_0h-Nr1h3Rxra_MA0494.2 16 bp overlap
Olig2 1 dataset
Motif DE_12h DE_12h-Olig2_MA1997.2 6 bp overlap
PATZ1 3 datasets
Motif DE_12h DE_12h-PATZ1_MA1961.2 11 bp overlap
Motif DE_12h DE_12h-PATZ1_MA1961.2 11 bp overlap
Motif ES_0h ES_0h-PATZ1_MA1961.2 11 bp overlap
PAX3-FOXO1 2 datasets
ChIP RH4 GSE140115.PAX3-FOXO1.RH4 169 bp overlap
ChIP RH4 GSE140115.PAX3-FOXO1.RH4 241 bp overlap
PLAGL2 2 datasets
Motif DE_12h DE_12h-PLAGL2_MA1548.2 8 bp overlap
Motif ES_0h ES_0h-PLAGL2_MA1548.2 8 bp overlap
POU5F1 1 dataset
ChIP BG03 GSE21614.POU5F1.BG03 153 bp overlap
PRDM9 1 dataset
Motif DE_12h DE_12h-PRDM9_MA1723.2 20 bp overlap
Pgr 2 datasets
Motif DE_12h DE_12h-Pgr_MA2323.1 17 bp overlap
Motif ES_0h ES_0h-Pgr_MA2323.1 17 bp overlap
Ptf1A 1 dataset
Motif DE_12h DE_12h-Ptf1A_MA1619.2 8 bp overlap
RAD21 1 dataset
ChIP HUVEC-C_hypoxia GSE94872.RAD21.HUVEC-C_hypoxia 145 bp overlap
RBBP5 1 dataset
ChIP WA01 ENCSR000AQC.RBBP5.WA01 463 bp overlap
RELA 1 dataset
ChIP KB_IL GSE52469.RELA.KB_IL 184 bp overlap
Runx1 2 datasets
Motif DE_12h DE_12h-Runx1_MA0002.3 9 bp overlap
Motif ES_0h ES_0h-Runx1_MA0002.3 9 bp overlap
SALL2 1 dataset
ChIP HEK293 GSE145940.SALL2.HEK293 198 bp overlap
SETDB1 2 datasets
ChIP HEK293 ENCSR348AGV.SETDB1.HEK293 163 bp overlap
ChIP U2OS ENCSR000EYD.SETDB1.U2OS 163 bp overlap
SIN3A 1 dataset
ChIP hiPSC_IID12 GSE106870.SIN3A.hiPSC_IID12 162 bp overlap
SIX1 1 dataset
Motif DE_12h DE_12h-SIX1_MA1118.2 9 bp overlap
SIX2 5 datasets
Motif DE_12h DE_12h-SIX2_MA1119.2 11 bp overlap
Motif ES_0h ES_0h-SIX2_MA1119.2 11 bp overlap
ChIP HEK GSE73865.SIX2.HEK 730 bp overlap
ChIP kidney_fetal_16w GSE75948.SIX2.kidney_fetal_16w 525 bp overlap
ChIP kidney_fetal_17w GSE75948.SIX2.kidney_fetal_17w 597 bp overlap
SKI 3 datasets
ChIP HL-60 GSE107553.SKI.HL-60 157 bp overlap
ChIP HL-60 GSE107553.SKI.HL-60 144 bp overlap
ChIP HL-60_CRISPRCas9_ctrl GSE107553.SKI.HL-60_CRISPRCas9_ctrl 236 bp overlap
SMAD2 1 dataset
ChIP hESC_YAP-_activinA_15h GSE99202.SMAD2.hESC_YAP-_activinA_15h 335 bp overlap
SMAD2-3 1 dataset
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 325 bp overlap
SMAD3 1 dataset
ChIP BG03 GSE21614.SMAD3.BG03 155 bp overlap
SNAI2 1 dataset
ChIP SMS-CTR_D48 GSE137168.SNAI2.SMS-CTR_D48 271 bp overlap
SOX12 2 datasets
Motif DE_12h DE_12h-SOX12_MA1561.2 10 bp overlap
Motif ES_0h ES_0h-SOX12_MA1561.2 10 bp overlap
SOX2 1 dataset
ChIP RENVM GSE49404.SOX2.RENVM 240 bp overlap
SP1 1 dataset
Motif DE_12h DE_12h-SP1_MA0079.5 9 bp overlap
SP2 1 dataset
Motif DE_12h DE_12h-SP2_MA0516.3 9 bp overlap
SP4 1 dataset
Motif DE_12h DE_12h-SP4_MA0685.2 9 bp overlap
SP5 2 datasets
Motif DE_12h DE_12h-SP5_MA1965.2 6 bp overlap
Motif DE_12h DE_12h-SP5_MA1965.2 6 bp overlap
SP8 1 dataset
Motif DE_12h DE_12h-SP8_MA0747.2 11 bp overlap
SPIB 1 dataset
Motif DE_12h DE_12h-SPIB_MA0081.3 13 bp overlap
SUZ12 1 dataset
ChIP LNCaP GSE39459.SUZ12.LNCaP 214 bp overlap
Spi1 2 datasets
Motif DE_12h DE_12h-Spi1_MA0080.7 13 bp overlap
Motif DE_12h DE_12h-Spi1_MA0080.7 13 bp overlap
Stat5b 2 datasets
Motif DE_12h DE_12h-Stat5b_MA1625.2 9 bp overlap
Motif ES_0h ES_0h-Stat5b_MA1625.2 9 bp overlap
Stat6 2 datasets
Motif DE_12h DE_12h-Stat6_MA0520.2 10 bp overlap
Motif ES_0h ES_0h-Stat6_MA0520.2 10 bp overlap
TAF1 2 datasets
ChIP H1 ENCFF478SZO 485 bp overlap
ChIP WA01 ENCSR000BHO.TAF1.WA01 393 bp overlap
TCF4 1 dataset
Motif DE_12h DE_12h-TCF4_MA0830.3 8 bp overlap
TEAD1 1 dataset
Motif DE_12h DE_12h-TEAD1_MA0090.4 9 bp overlap
TEAD2 1 dataset
Motif DE_12h DE_12h-TEAD2_MA1121.2 7 bp overlap
TEAD3 1 dataset
Motif DE_12h DE_12h-TEAD3_MA0808.1 8 bp overlap
TEAD4 2 datasets
Motif DE_12h DE_12h-TEAD4_MA0809.3 8 bp overlap
ChIP H1 ENCFF778PAX 245 bp overlap
TFAP4::FLI1 1 dataset
Motif DE_12h DE_12h-TFAP4FLI1_MA1967.2 14 bp overlap
THAP1 1 dataset
Motif DE_12h DE_12h-THAP1_MA0597.3 8 bp overlap
THRB 1 dataset
Motif DE_12h DE_12h-THRB_MA1575.2 17 bp overlap
TRIM28 1 dataset
ChIP HEK293 ENCFF582MWI 671 bp overlap
TRPS1 2 datasets
Motif DE_12h DE_12h-TRPS1_MA1970.2 8 bp overlap
Motif ES_0h ES_0h-TRPS1_MA1970.2 8 bp overlap
Tcf12 1 dataset
Motif DE_12h DE_12h-Tcf12_MA0521.3 6 bp overlap
Twist2 1 dataset
Motif DE_12h DE_12h-Twist2_MA0633.3 6 bp overlap
Wt1 1 dataset
Motif DE_12h DE_12h-Wt1_MA1627.2 10 bp overlap
YY1 3 datasets
ChIP H1 ENCFF524BTL 337 bp overlap
ChIP RH4_DMSO-6H GSE116344.YY1.RH4_DMSO-6H 213 bp overlap
ChIP WA01 ENCSR000BKD.YY1.WA01 212 bp overlap
ZBTB26 3 datasets
Motif DE_12h DE_12h-ZBTB26_MA1579.2 8 bp overlap
Motif DE_12h DE_12h-ZBTB26_MA1579.2 8 bp overlap
Motif ES_0h ES_0h-ZBTB26_MA1579.2 8 bp overlap
ZEB1 1 dataset
Motif DE_12h DE_12h-ZEB1_MA0103.4 6 bp overlap
ZFP14 1 dataset
Motif DE_12h DE_12h-ZFP14_MA1972.1 15 bp overlap
ZIM3 1 dataset
Motif DE_12h DE_12h-ZIM3_MA1709.2 11 bp overlap
ZKSCAN1 2 datasets
Motif DE_12h DE_12h-ZKSCAN1_MA1585.2 9 bp overlap
Motif ES_0h ES_0h-ZKSCAN1_MA1585.2 9 bp overlap
ZNF140 1 dataset
Motif DE_12h DE_12h-ZNF140_MA1589.2 19 bp overlap
ZNF148 1 dataset
Motif DE_12h DE_12h-ZNF148_MA1653.2 10 bp overlap
ZNF281 1 dataset
Motif DE_12h DE_12h-ZNF281_MA1630.3 10 bp overlap
ZNF320 2 datasets
Motif DE_12h DE_12h-ZNF320_MA1976.2 20 bp overlap
Motif ES_0h ES_0h-ZNF320_MA1976.2 20 bp overlap
ZNF416 1 dataset
Motif DE_12h DE_12h-ZNF416_MA1979.2 10 bp overlap
ZNF449 2 datasets
Motif DE_12h DE_12h-ZNF449_MA1656.2 10 bp overlap
Motif ES_0h ES_0h-ZNF449_MA1656.2 10 bp overlap
ZNF479 1 dataset
ChIP HEK293T GSE78099.ZNF479.HEK293T 85 bp overlap
ZNF528 1 dataset
Motif DE_12h DE_12h-ZNF528_MA1597.1 17 bp overlap
ZNF549 2 datasets
Motif DE_12h DE_12h-ZNF549_MA1728.2 8 bp overlap
Motif ES_0h ES_0h-ZNF549_MA1728.2 8 bp overlap
ZNF558 1 dataset
Motif DE_12h DE_12h-ZNF558_MA2335.1 29 bp overlap
ZNF573 1 dataset
ChIP HEK293T GSE78099.ZNF573.HEK293T 185 bp overlap
ZNF708 1 dataset
Motif DE_12h DE_12h-ZNF708_MA1730.2 9 bp overlap
ZNF778 1 dataset
ChIP HEK293T GSE78099.ZNF778.HEK293T 257 bp overlap
ZNF93 2 datasets
Motif DE_12h DE_12h-ZNF93_MA1721.2 14 bp overlap
Motif ES_0h ES_0h-ZNF93_MA1721.2 14 bp overlap
Zfp335 3 datasets
Motif DE_12h DE_12h-Zfp335_MA2002.2 7 bp overlap
Motif DE_12h DE_12h-Zfp335_MA2002.2 7 bp overlap
Motif ES_0h ES_0h-Zfp335_MA2002.2 7 bp overlap
Zfp809 1 dataset
Motif DE_12h DE_12h-Zfp809_MA2125.1 9 bp overlap
Zic2 1 dataset
Motif DE_12h DE_12h-Zic2_MA1629.2 9 bp overlap
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