chr14 : 85,546,295 – 85,546,973
678 bp 116 TFs 1 linked gene
This 678 bp open chromatin element is linked to FLRT2 and is bound by 116 transcription factors.
Linked Genes
1 gene
Gene↕ Expression Dist. to TSS↑ Distance↕ Link type↕
FLRT2 13.4 kb Distal Multiome
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr14:85,541,295 – 85,551,973
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
116 transcription factors
Source
Cell type
ARID1A 1 dataset
ChIP HAP1 GSE108387.ARID1A.HAP1 256 bp overlap
ARNTL 1 dataset
ChIP U2OS_DMOG GSE85096.ARNTL.U2OS_DMOG 262 bp overlap
ASH2L 3 datasets
ChIP H1 ENCFF399KAM 678 bp overlap
ChIP H1 ENCFF399KAM 492 bp overlap
ChIP WA01 ENCSR850KIP.ASH2L.WA01 574 bp overlap
ATF2 1 dataset
ChIP WA01 ENCSR000BQU.ATF2.WA01 331 bp overlap
BCL6 2 datasets
Motif DE_12h DE_12h-BCL6_MA0463.3 13 bp overlap
Motif ES_0h ES_0h-BCL6_MA0463.3 13 bp overlap
BCOR 3 datasets
ChIP WA01 GSE104690.BCOR.WA01 213 bp overlap
ChIP WA01_KDM2B_KO GSE104690.BCOR.WA01_KDM2B_KO 234 bp overlap
ChIP WA01_RNF2-R_Doxycyclin GSE104690.BCOR.WA01_RNF2-R_Doxycyclin 200 bp overlap
CBFB 1 dataset
ChIP WTC11 ENCFF113HIY 486 bp overlap
CHD7 1 dataset
ChIP WA01 ENCSR000AVA.CHD7.WA01 197 bp overlap
CREB1 1 dataset
ChIP GM23338 ENCSR214ZAV.CREB1.GM23338 454 bp overlap
CTBP2 1 dataset
ChIP WA01 ENCSR000EUO.CTBP2.WA01 492 bp overlap
CTNNB1 2 datasets
ChIP hESC_YAP-_activinA_15h GSE99202.CTNNB1.hESC_YAP-_activinA_15h 295 bp overlap
ChIP hESC_activinA_15h GSE99202.CTNNB1.hESC_activinA_15h 287 bp overlap
E2F6 4 datasets
Motif DE_12h DE_12h-E2F6_MA0471.3 8 bp overlap
Motif ES_0h ES_0h-E2F6_MA0471.3 8 bp overlap
ChIP H1 ENCFF785DWK 445 bp overlap
ChIP WA01 ENCSR000BSI.E2F6.WA01 146 bp overlap
ELF2 2 datasets
Motif DE_12h DE_12h-ELF2_MA1483.3 10 bp overlap
Motif ES_0h ES_0h-ELF2_MA1483.3 10 bp overlap
EP300 2 datasets
ChIP H1 ENCFF927IYK 297 bp overlap
ChIP WA01 ENCSR000BKK.EP300.WA01 230 bp overlap
ERF::HOXB13 1 dataset
Motif DE_12h DE_12h-ERFHOXB13_MA1937.2 13 bp overlap
ESR1 1 dataset
ChIP MCF-7_RUNX2_DOX GSE86538.ESR1.MCF-7_RUNX2_DOX 161 bp overlap
ETV1 2 datasets
Motif DE_12h DE_12h-ETV1_MA0761.3 9 bp overlap
Motif ES_0h ES_0h-ETV1_MA0761.3 9 bp overlap
ETV2::DRGX 2 datasets
Motif DE_12h DE_12h-ETV2DRGX_MA1940.2 12 bp overlap
Motif ES_0h ES_0h-ETV2DRGX_MA1940.2 12 bp overlap
ETV2::FOXI1 2 datasets
Motif DE_12h DE_12h-ETV2FOXI1_MA1942.2 11 bp overlap
Motif ES_0h ES_0h-ETV2FOXI1_MA1942.2 11 bp overlap
ETV5::DRGX 4 datasets
Motif DE_12h DE_12h-ETV5DRGX_MA1944.2 12 bp overlap
Motif DE_12h DE_12h-ETV5DRGX_MA1944.2 12 bp overlap
Motif ES_0h ES_0h-ETV5DRGX_MA1944.2 12 bp overlap
Motif ES_0h ES_0h-ETV5DRGX_MA1944.2 12 bp overlap
ETV5::HOXA2 2 datasets
Motif DE_12h DE_12h-ETV5HOXA2_MA1948.2 12 bp overlap
Motif ES_0h ES_0h-ETV5HOXA2_MA1948.2 12 bp overlap
EWSR1-FLI1 2 datasets
Motif DE_12h DE_12h-EWSR1-FLI1_MA0149.1 18 bp overlap
Motif ES_0h ES_0h-EWSR1-FLI1_MA0149.1 18 bp overlap
Elf5 4 datasets
Motif DE_12h DE_12h-Elf5_MA0136.4 8 bp overlap
Motif DE_12h DE_12h-Elf5_MA0136.4 8 bp overlap
Motif ES_0h ES_0h-Elf5_MA0136.4 8 bp overlap
Motif ES_0h ES_0h-Elf5_MA0136.4 8 bp overlap
FOXJ2::ELF1 2 datasets
Motif DE_12h DE_12h-FOXJ2ELF1_MA1952.2 11 bp overlap
Motif ES_0h ES_0h-FOXJ2ELF1_MA1952.2 11 bp overlap
FOXK1 1 dataset
ChIP WTC11 ENCFF875IGU 391 bp overlap
FOXP4 1 dataset
ChIP WTC11 ENCFF708TAF 377 bp overlap
GATA4 1 dataset
Motif DE_12h DE_12h-GATA4_MA0482.3 8 bp overlap
GATA6 3 datasets
ChIP DE_D1 S41-DE-d1-GATA6-exp2 124 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 156 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 139 bp overlap
HNF1A 1 dataset
Motif DE_12h DE_12h-HNF1A_MA0046.3 13 bp overlap
HOXB4 1 dataset
Motif DE_12h DE_12h-HOXB4_MA1499.2 6 bp overlap
HOXC4 1 dataset
Motif DE_12h DE_12h-HOXC4_MA1504.2 6 bp overlap
HOXD12::ELK1 2 datasets
Motif DE_12h DE_12h-HOXD12ELK1_MA1958.2 13 bp overlap
Motif ES_0h ES_0h-HOXD12ELK1_MA1958.2 13 bp overlap
HOXD4 1 dataset
Motif DE_12h DE_12h-HOXD4_MA1507.2 6 bp overlap
Hnf1A 2 datasets
Motif DE_12h DE_12h-Hnf1A_MA1991.2 10 bp overlap
Motif ES_0h ES_0h-Hnf1A_MA1991.2 10 bp overlap
Hoxa13 2 datasets
Motif DE_12h DE_12h-Hoxa13_MA0650.4 8 bp overlap
Motif ES_0h ES_0h-Hoxa13_MA0650.4 8 bp overlap
IKZF2 6 datasets
Motif DE_12h DE_12h-IKZF2_MA2326.1 6 bp overlap
Motif DE_12h DE_12h-IKZF2_MA2326.1 6 bp overlap
Motif DE_12h DE_12h-IKZF2_MA2326.1 6 bp overlap
Motif ES_0h ES_0h-IKZF2_MA2326.1 6 bp overlap
Motif ES_0h ES_0h-IKZF2_MA2326.1 6 bp overlap
Motif ES_0h ES_0h-IKZF2_MA2326.1 6 bp overlap
IRF3 1 dataset
Motif DE_12h DE_12h-IRF3_MA1418.2 17 bp overlap
Ikzf3 2 datasets
Motif DE_12h DE_12h-Ikzf3_MA1992.2 9 bp overlap
Motif ES_0h ES_0h-Ikzf3_MA1992.2 9 bp overlap
Isl1 1 dataset
Motif DE_12h DE_12h-Isl1_MA1608.2 7 bp overlap
JARID2 1 dataset
ChIP MRC-5_IPS25_CTR GSE48515.JARID2.MRC-5_IPS25_CTR 226 bp overlap
JUN 2 datasets
ChIP ESC S34-ESC-d0-JUN-exp2 312 bp overlap
ChIP HUES-8 GSE109524.JUN.HUES-8 307 bp overlap
KDM4A 1 dataset
ChIP WA01 ENCSR000AVC.KDM4A.WA01 216 bp overlap
KDM5B 1 dataset
ChIP WA01 ENCSR000AUR.KDM5B.WA01 224 bp overlap
KLF4 1 dataset
ChIP hiPSC GSE56567.KLF4.hiPSC 277 bp overlap
MAX 2 datasets
ChIP WA01 ENCSR000EUP.MAX.WA01 352 bp overlap
ChIP WTC11 ENCFF223QFY 585 bp overlap
MEIS1 1 dataset
Motif DE_12h DE_12h-MEIS1_MA0498.3 5 bp overlap
MEIS2 1 dataset
Motif DE_12h DE_12h-MEIS2_MA0774.1 8 bp overlap
MEIS3 1 dataset
Motif DE_12h DE_12h-MEIS3_MA0775.2 7 bp overlap
MGA::EVX1 2 datasets
Motif DE_12h DE_12h-MGAEVX1_MA1960.2 11 bp overlap
Motif ES_0h ES_0h-MGAEVX1_MA1960.2 11 bp overlap
MXI1 2 datasets
Motif DE_12h DE_12h-MXI1_MA1108.3 6 bp overlap
Motif ES_0h ES_0h-MXI1_MA1108.3 6 bp overlap
MYOD1 1 dataset
ChIP RD_shSNAI2 GSE137168.MYOD1.RD_shSNAI2 238 bp overlap
Msgn1 2 datasets
Motif DE_12h DE_12h-Msgn1_MA1524.3 10 bp overlap
Motif ES_0h ES_0h-Msgn1_MA1524.3 10 bp overlap
NANOG 11 datasets
ChIP GM23338 ENCFF065NZG 311 bp overlap
ChIP GM23338 ENCSR061DGF.NANOG.GM23338 206 bp overlap
ChIP H1 ENCFF747ZPQ 241 bp overlap
ChIP HUES-8 GSE109524.NANOG.HUES-8 678 bp overlap
ChIP WA01 ERP004238.NANOG.WA01 407 bp overlap
ChIP WA01 ENCSR000BMT.NANOG.WA01 157 bp overlap
ChIP WA01_3IL ERP004238.NANOG.WA01_3IL 678 bp overlap
ChIP WA09 GSE105028.NANOG.WA09 678 bp overlap
ChIP WA09_heat-shock GSE105028.NANOG.WA09_heat-shock 678 bp overlap
ChIP hESC GSE20650.NANOG.hESC 156 bp overlap
ChIP hESC GSE18292.NANOG.hESC 172 bp overlap
NCOA2 1 dataset
ChIP MCF-7 ERP000901.NCOA2.MCF-7 188 bp overlap
NIPBL 4 datasets
ChIP hESC GSE64758.NIPBL.hESC 204 bp overlap
ChIP hESC_WNT3A GSE64758.NIPBL.hESC_WNT3A 219 bp overlap
ChIP hESC_WNT3A_ACTIVIN GSE64758.NIPBL.hESC_WNT3A_ACTIVIN 298 bp overlap
ChIP hESC_activin GSE64758.NIPBL.hESC_activin 214 bp overlap
NKX6-3 1 dataset
Motif DE_12h DE_12h-NKX6-3_MA1530.2 8 bp overlap
NR2C2 2 datasets
Motif DE_12h DE_12h-NR2C2_MA0504.2 14 bp overlap
Motif ES_0h ES_0h-NR2C2_MA0504.2 14 bp overlap
Nfatc2 1 dataset
Motif DE_12h DE_12h-Nfatc2_MA0152.3 8 bp overlap
POU2F1::SOX2 2 datasets
Motif DE_12h DE_12h-POU2F1SOX2_MA1962.1 17 bp overlap
Motif ES_0h ES_0h-POU2F1SOX2_MA1962.1 17 bp overlap
POU4F1 1 dataset
Motif DE_12h DE_12h-POU4F1_MA0790.2 12 bp overlap
POU4F3 1 dataset
Motif DE_12h DE_12h-POU4F3_MA0791.2 12 bp overlap
POU5F1 14 datasets
ChIP BG03 GSE21614.POU5F1.BG03 175 bp overlap
ChIP GM23338 ENCFF333SNB 311 bp overlap
ChIP HUES-8 GSE109524.POU5F1.HUES-8 600 bp overlap
ChIP NCCIT GSE36134.POU5F1.NCCIT 313 bp overlap
ChIP NCCIT_SNF5 GSE36134.POU5F1.NCCIT_SNF5 678 bp overlap
ChIP SKM-1 GSE93706.POU5F1.SKM-1 229 bp overlap
ChIP WA01 ENCSR000BMU.POU5F1.WA01 139 bp overlap
ChIP WA09 GSE105028.POU5F1.WA09 238 bp overlap
ChIP WA09_heat-shock GSE105028.POU5F1.WA09_heat-shock 312 bp overlap
ChIP hESC ENCSR264RJX.POU5F1.hESC 250 bp overlap
ChIP hESC_NAIVE GSE69646.POU5F1.hESC_NAIVE 678 bp overlap
ChIP hESC_PRIMED GSE69646.POU5F1.hESC_PRIMED 599 bp overlap
ChIP hiPSC GSE56567.POU5F1.hiPSC 302 bp overlap
ChIP hiPSC_3s2 GSE81899.POU5F1.hiPSC_3s2 270 bp overlap
POU6F1 1 dataset
Motif DE_12h DE_12h-POU6F1_MA1549.2 7 bp overlap
POU6F2 1 dataset
Motif DE_12h DE_12h-POU6F2_MA0793.2 9 bp overlap
PPARG 3 datasets
Motif DE_12h DE_12h-PPARG_MA0066.2 19 bp overlap
Motif DE_12h DE_12h-PPARG_MA0066.2 19 bp overlap
Motif ES_0h ES_0h-PPARG_MA0066.2 19 bp overlap
PRDM14 1 dataset
ChIP hESC GSE22767.PRDM14.hESC 260 bp overlap
Pou5f1::Sox2 6 datasets
Motif DE_12h DE_12h-Pou5f1Sox2_MA0142.1 15 bp overlap
Motif DE_12h DE_12h-Pou5f1Sox2_MA0142.1 15 bp overlap
Motif DE_12h DE_12h-Pou5f1Sox2_MA0142.1 15 bp overlap
Motif ES_0h ES_0h-Pou5f1Sox2_MA0142.1 15 bp overlap
Motif ES_0h ES_0h-Pou5f1Sox2_MA0142.1 15 bp overlap
Motif ES_0h ES_0h-Pou5f1Sox2_MA0142.1 15 bp overlap
RAD21 5 datasets
ChIP H1 ENCFF967OJF 245 bp overlap
ChIP HAP1 GSE126634.RAD21.HAP1 570 bp overlap
ChIP HAP1 GSE152721.RAD21.HAP1 209 bp overlap
ChIP HAP1_CTCF GSE126634.RAD21.HAP1_CTCF 433 bp overlap
ChIP HAP1_clone25 GSE152721.RAD21.HAP1_clone25 182 bp overlap
REST 1 dataset
ChIP WA01 ENCSR000BHM.REST.WA01 103 bp overlap
RNF2 1 dataset
ChIP WA09_heat-shock GSE105028.RNF2.WA09_heat-shock 267 bp overlap
SIX2 2 datasets
Motif DE_12h DE_12h-SIX2_MA1119.2 11 bp overlap
Motif ES_0h ES_0h-SIX2_MA1119.2 11 bp overlap
SMAD1 1 dataset
ChIP BG03 GSE36578.SMAD1.BG03 96 bp overlap
SMAD2 4 datasets
Motif DE_12h DE_12h-SMAD2_MA1964.2 6 bp overlap
Motif ES_0h ES_0h-SMAD2_MA1964.2 6 bp overlap
ChIP hESC_YAP-_activinA_15h GSE99202.SMAD2.hESC_YAP-_activinA_15h 313 bp overlap
ChIP hESC_activinA_15h GSE99202.SMAD2.hESC_activinA_15h 248 bp overlap
SMAD2-3 3 datasets
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 254 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.SMAD2-3.HUES-8_DE_JNKi 275 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.SMAD2-3.HUES-8_DE_ctrl 452 bp overlap
SMAD2_3 2 datasets
ChIP DE_D1 S39-DE-d1-SMAD2_3-exp2 426 bp overlap
ChIP ESC S33-ESC-d0-SMAD2_3-exp2 366 bp overlap
SMAD3 3 datasets
ChIP BG03 GSE21614.SMAD3.BG03 281 bp overlap
Motif DE_12h DE_12h-SMAD3_MA0795.1 10 bp overlap
Motif ES_0h ES_0h-SMAD3_MA0795.1 10 bp overlap
SMAD5 2 datasets
Motif DE_12h DE_12h-SMAD5_MA1557.1 10 bp overlap
Motif ES_0h ES_0h-SMAD5_MA1557.1 10 bp overlap
SMARCA4 3 datasets
ChIP WA09_heat-shock GSE105028.SMARCA4.WA09_heat-shock 484 bp overlap
ChIP hiPSC GSE124903.SMARCA4.hiPSC 263 bp overlap
ChIP hiPSC_WT GSE124903.SMARCA4.hiPSC_WT 372 bp overlap
SMARCB1 2 datasets
ChIP hiPSC GSE124903.SMARCB1.hiPSC 391 bp overlap
ChIP hiPSC_WT GSE124903.SMARCB1.hiPSC_WT 334 bp overlap
SMARCC1 2 datasets
ChIP hiPSC GSE124903.SMARCC1.hiPSC 219 bp overlap
ChIP hiPSC_WT GSE124903.SMARCC1.hiPSC_WT 535 bp overlap
SOX2 2 datasets
ChIP LK2_GFP GSE137459.SOX2.LK2_GFP 671 bp overlap
ChIP hiPSC GSE56567.SOX2.hiPSC 363 bp overlap
SP1 1 dataset
ChIP WA01 ENCSR000BIR.SP1.WA01 160 bp overlap
SP4 1 dataset
ChIP WA01 ENCSR000BQV.SP4.WA01 134 bp overlap
SPI1 1 dataset
ChIP CTV-1_FLAG_15perc GSE128835.SPI1.CTV-1_FLAG_15perc 173 bp overlap
SRF 2 datasets
Motif DE_12h DE_12h-SRF_MA0083.3 16 bp overlap
Motif ES_0h ES_0h-SRF_MA0083.3 16 bp overlap
SS18-SSX 1 dataset
ChIP fibroblast_7aa GSE139053.SS18-SSX.fibroblast_7aa 99 bp overlap
TBX5 2 datasets
Motif DE_12h DE_12h-TBX5_MA0807.1 8 bp overlap
Motif ES_0h ES_0h-TBX5_MA0807.1 8 bp overlap
TCF12 1 dataset
ChIP WA01 ENCSR000BIT.TCF12.WA01 150 bp overlap
TCF7L2 3 datasets
Motif DE_12h DE_12h-TCF7L2_MA0523.2 9 bp overlap
Motif ES_0h ES_0h-TCF7L2_MA0523.2 9 bp overlap
ChIP MDA-MB-453 GSE45201.TCF7L2.MDA-MB-453 201 bp overlap
TEAD1 3 datasets
Motif DE_12h DE_12h-TEAD1_MA0090.4 9 bp overlap
Motif ES_0h ES_0h-TEAD1_MA0090.4 9 bp overlap
ChIP WTC11 ENCFF502QUV 405 bp overlap
TEAD3 2 datasets
Motif DE_12h DE_12h-TEAD3_MA0808.1 8 bp overlap
Motif ES_0h ES_0h-TEAD3_MA0808.1 8 bp overlap
TFAP2A 1 dataset
Motif DE_12h DE_12h-TFAP2A_MA0810.2 11 bp overlap
TFAP2B 1 dataset
Motif DE_12h DE_12h-TFAP2B_MA0811.2 11 bp overlap
TFAP2C 1 dataset
Motif DE_12h DE_12h-TFAP2C_MA0524.3 11 bp overlap
THAP1 1 dataset
Motif DE_12h DE_12h-THAP1_MA0597.3 8 bp overlap
TP53 1 dataset
ChIP H9_ectoderm GSE142050.TP53.H9_ectoderm 575 bp overlap
TRIM28 2 datasets
ChIP WIBR3_NAIVE GSE84382.TRIM28.WIBR3_NAIVE 291 bp overlap
ChIP hESC GSE115387.TRIM28.hESC 290 bp overlap
USF1 3 datasets
ChIP H1 ENCFF090WVU 154 bp overlap
ChIP WA01 ENCSR000BIU.USF1.WA01 133 bp overlap
ChIP WA01 ENCSR000BIU.USF1.WA01 121 bp overlap
YY1 3 datasets
ChIP H1 ENCFF524BTL 337 bp overlap
ChIP NT2-D1 ENCSR000EXG.YY1.NT2-D1 311 bp overlap
ChIP WA01 ENCSR000BKD.YY1.WA01 387 bp overlap
ZBTB21 1 dataset
ChIP WTC11 ENCFF677ZYY 401 bp overlap
ZEB1 1 dataset
ChIP neuron_bipolar_doxy_4d ENCSR418KUS.ZEB1.neuron_bipolar_doxy_4d 112 bp overlap
ZFP42 2 datasets
Motif DE_12h DE_12h-ZFP42_MA1651.2 13 bp overlap
Motif ES_0h ES_0h-ZFP42_MA1651.2 13 bp overlap
ZNF114 1 dataset
ChIP GM23338 ENCSR555KFE.ZNF114.GM23338 232 bp overlap
ZNF136 1 dataset
Motif DE_12h DE_12h-ZNF136_MA1588.1 15 bp overlap
ZNF207 1 dataset
ChIP WA09 GSE118632.ZNF207.WA09 368 bp overlap
ZNF317 2 datasets
Motif DE_12h DE_12h-ZNF317_MA1593.2 8 bp overlap
Motif ES_0h ES_0h-ZNF317_MA1593.2 8 bp overlap
ZNF331 1 dataset
ChIP GM23338 ENCSR918LRB.ZNF331.GM23338 156 bp overlap
ZNF341 2 datasets
Motif DE_12h DE_12h-ZNF341_MA1655.2 8 bp overlap
Motif ES_0h ES_0h-ZNF341_MA1655.2 8 bp overlap
ZNF382 2 datasets
Motif DE_12h DE_12h-ZNF382_MA1594.1 24 bp overlap
Motif ES_0h ES_0h-ZNF382_MA1594.1 24 bp overlap
ZNF418 2 datasets
Motif DE_12h DE_12h-ZNF418_MA1980.1 15 bp overlap
Motif ES_0h ES_0h-ZNF418_MA1980.1 15 bp overlap
ZNF449 2 datasets
Motif DE_12h DE_12h-ZNF449_MA1656.2 10 bp overlap
Motif ES_0h ES_0h-ZNF449_MA1656.2 10 bp overlap
ZNF462 2 datasets
ChIP GM23338 ENCFF896CCA 251 bp overlap
ChIP GM23338 ENCSR334UWP.ZNF462.GM23338 522 bp overlap
ZNF528 2 datasets
Motif DE_12h DE_12h-ZNF528_MA1597.1 17 bp overlap
Motif ES_0h ES_0h-ZNF528_MA1597.1 17 bp overlap
ZNF582 2 datasets
Motif DE_12h DE_12h-ZNF582_MA1983.2 19 bp overlap
Motif ES_0h ES_0h-ZNF582_MA1983.2 19 bp overlap
ZNF766 3 datasets
Motif DE_12h DE_12h-ZNF766_MA2098.1 9 bp overlap
Motif DE_12h DE_12h-ZNF766_MA2098.1 9 bp overlap
Motif ES_0h ES_0h-ZNF766_MA2098.1 9 bp overlap
ZSCAN31 1 dataset
Motif DE_12h DE_12h-ZSCAN31_MA1722.2 18 bp overlap
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