chr4 : 184,167,643 184,168,130
487 bp 103 TFs 1 linked gene
This 487 bp open chromatin element is linked to STOX2 and is bound by 103 transcription factors.
Linked Genes
1 gene
Gene Expression Dist. to TSS Distance Link type
STOX2 370.0 kb Distal Multiome
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr4:184,162,643 – 184,173,130
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
103 transcription factors
Source
Cell type
AR 1 dataset
ChIP WTC11 ENCFF267GQJ 265 bp overlap
ASCL1 1 dataset
ChIP NCI-H82 GSE69394.ASCL1.NCI-H82 221 bp overlap
ASH2L 3 datasets
ChIP H1 ENCFF399KAM 487 bp overlap
ChIP H1 ENCFF399KAM 262 bp overlap
ChIP WA01 ENCSR850KIP.ASH2L.WA01 487 bp overlap
ATF2 1 dataset
ChIP WA01 ENCSR000BQU.ATF2.WA01 366 bp overlap
ATOH7 1 dataset
Motif DE_12h DE_12h-ATOH7_MA1468.1 10 bp overlap
Atoh1 1 dataset
Motif DE_12h DE_12h-Atoh1_MA1467.3 7 bp overlap
BATF 1 dataset
ChIP GM12878 ENCFF954REE 160 bp overlap
BCL11A 2 datasets
ChIP WA01 ENCSR000BMJ.BCL11A.WA01 121 bp overlap
ChIP WA01 ENCSR000BIP.BCL11A.WA01 102 bp overlap
BCL6 1 dataset
ChIP B-cell_GERMINAL_CENTER GSE43350.BCL6.B-cell_GERMINAL_CENTER 102 bp overlap
BCOR 3 datasets
ChIP WA01 GSE104690.BCOR.WA01 487 bp overlap
ChIP WA01_RNF2-R GSE104690.BCOR.WA01_RNF2-R 417 bp overlap
ChIP WA01_RNF2-R_Doxycyclin GSE104690.BCOR.WA01_RNF2-R_Doxycyclin 436 bp overlap
BHLHA15 1 dataset
Motif DE_12h DE_12h-BHLHA15_MA0607.2 10 bp overlap
CHD7 1 dataset
ChIP WA01 ENCSR000AVA.CHD7.WA01 189 bp overlap
CREB1 8 datasets
ChIP GM12878 ENCFF870CVH 147 bp overlap
ChIP GM12878 ENCSR000BUF.CREB1.GM12878 163 bp overlap
ChIP GM23338 ENCFF432ZEW 172 bp overlap
ChIP GM23338 ENCSR214ZAV.CREB1.GM23338 277 bp overlap
ChIP H1 ENCFF955PMP 155 bp overlap
ChIP Hep-G2 ENCSR112ALD.CREB1.Hep-G2 171 bp overlap
ChIP HepG2 ENCFF792THT 210 bp overlap
ChIP WA01 ENCSR000BSN.CREB1.WA01 280 bp overlap
CREM 1 dataset
ChIP GM12878 ENCSR839XZU.CREM.GM12878 110 bp overlap
CTBP2 2 datasets
ChIP H1 ENCFF329MAX 445 bp overlap
ChIP WA01 ENCSR000EUO.CTBP2.WA01 277 bp overlap
CTCF 5 datasets
ChIP C4-2B ENCFF821XVN 125 bp overlap
ChIP RWPE2 ENCFF911IEE 63 bp overlap
ChIP T-47D_D538G GSE148277.CTCF.T-47D_D538G 101 bp overlap
ChIP lymphoblast_mut GSE155324.CTCF.lymphoblast_mut 108 bp overlap
ChIP peripheral-blood-neutrophil_US-1 GSE126755.CTCF.peripheral-blood-neutrophil_US-1 134 bp overlap
CTNNB1 2 datasets
ChIP hESC_YAP-_activinA_15h GSE99202.CTNNB1.hESC_YAP-_activinA_15h 334 bp overlap
ChIP hESC_activinA_15h GSE99202.CTNNB1.hESC_activinA_15h 410 bp overlap
E2F6 2 datasets
ChIP WA01 ENCSR000BSI.E2F6.WA01 290 bp overlap
ChIP WA01 ENCSR000BSI.E2F6.WA01 130 bp overlap
EP300 3 datasets
ChIP GM12878 ENCSR000BHB.EP300.GM12878 81 bp overlap
ChIP H1 ENCFF927IYK 297 bp overlap
ChIP WA01 ENCSR000BKK.EP300.WA01 278 bp overlap
ETV6 1 dataset
ChIP WTC11 ENCFF812SCD 424 bp overlap
EZH2 4 datasets
ChIP LNCaP GSE39459.EZH2.LNCaP 92 bp overlap
ChIP LNCaP-abl GSE39459.EZH2.LNCaP-abl 203 bp overlap
ChIP neural progenitor cell ENCFF472NFV 337 bp overlap
ChIP peripheral-blood-mononuclear-cell_mut GSE115772.EZH2.peripheral-blood-mononuclear-cell_mut 136 bp overlap
FERD3L 1 dataset
Motif DE_12h DE_12h-FERD3L_MA1485.1 14 bp overlap
FOXA2 2 datasets
ChIP DE DE-FOXA2-1 336 bp overlap
ChIP DE DE-FOXA2-2 334 bp overlap
FOXK1 1 dataset
ChIP WTC11 ENCFF875IGU 360 bp overlap
FOXP1 1 dataset
ChIP H9 GSE31006.FOXP1.H9 273 bp overlap
FOXP4 1 dataset
ChIP WTC11 ENCFF708TAF 377 bp overlap
GATA4 2 datasets
ChIP DE DE-GATA4-1 317 bp overlap
ChIP DE DE-GATA4-2 343 bp overlap
GATA6 9 datasets
ChIP DE DE-GATA6-1 359 bp overlap
ChIP DE DE-GATA6-2 426 bp overlap
ChIP DE_D1 S09-DE-d1-GATA6-exp1 471 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 487 bp overlap
ChIP DE_D1 S41-DE-d1-GATA6-exp2 435 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 247 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 487 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 487 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 288 bp overlap
HAND2 1 dataset
Motif DE_12h DE_12h-HAND2_MA1638.2 6 bp overlap
HDAC2 1 dataset
ChIP WA01 ENCSR000AVB.HDAC2.WA01 253 bp overlap
IRF4 4 datasets
ChIP GM12878 ENCSR000BGY.IRF4.GM12878 200 bp overlap
ChIP OCI-Ly10 GSE142493.IRF4.OCI-Ly10 271 bp overlap
ChIP OCI-Ly3 GSE142493.IRF4.OCI-Ly3 260 bp overlap
ChIP OCI-Ly3_SHCTR GSE56857.IRF4.OCI-Ly3_SHCTR 255 bp overlap
JUN 6 datasets
ChIP DE_D1 S08-DE-d1-JUN-exp1 429 bp overlap
ChIP DE_D1 S40-DE-d1-JUN-exp2 487 bp overlap
ChIP DE_D2 S02-DE-d2-JUN-exp1 487 bp overlap
ChIP ESC S34-ESC-d0-JUN-exp2 487 bp overlap
ChIP HUES-8 GSE109524.JUN.HUES-8 371 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.JUN.HUES-8_DE_ctrl 487 bp overlap
JUNB 1 dataset
ChIP GM12878 ENCSR897MMC.JUNB.GM12878 201 bp overlap
JUND 2 datasets
ChIP H1 ENCFF468JZD 231 bp overlap
ChIP WA01 ENCSR000BKP.JUND.WA01 138 bp overlap
LEF1 1 dataset
ChIP hESC_WNT3A GSE64758.LEF1.hESC_WNT3A 272 bp overlap
MAX 2 datasets
ChIP WA01 ENCSR000EUP.MAX.WA01 127 bp overlap
ChIP WTC11 ENCFF223QFY 434 bp overlap
MED1 1 dataset
ChIP hESC_PRIMED GSE69646.MED1.hESC_PRIMED 362 bp overlap
MORC2 1 dataset
ChIP H9 GSE95374.MORC2.H9 320 bp overlap
MTF1 1 dataset
Motif DE_12h DE_12h-MTF1_MA0863.1 14 bp overlap
Msgn1 1 dataset
Motif DE_12h DE_12h-Msgn1_MA1524.3 10 bp overlap
NANOG 11 datasets
ChIP GM23338 ENCFF065NZG 308 bp overlap
ChIP GM23338 ENCSR061DGF.NANOG.GM23338 279 bp overlap
ChIP H1 ENCFF747ZPQ 186 bp overlap
ChIP HUES-8 GSE109524.NANOG.HUES-8 487 bp overlap
ChIP WA01 ERP004238.NANOG.WA01 404 bp overlap
ChIP WA01 ENCSR000BMT.NANOG.WA01 357 bp overlap
ChIP WA01_3IL ERP004238.NANOG.WA01_3IL 487 bp overlap
ChIP WA09 GSE105028.NANOG.WA09 487 bp overlap
ChIP WA09_heat-shock GSE105028.NANOG.WA09_heat-shock 467 bp overlap
ChIP hESC GSE20650.NANOG.hESC 332 bp overlap
ChIP hESC GSE18292.NANOG.hESC 254 bp overlap
NEUROD1 5 datasets
ChIP D283-Med GSE92582.NEUROD1.D283-Med 338 bp overlap
ChIP D283-Med_shNEUROD1-1579 GSE92582.NEUROD1.D283-Med_shNEUROD1-1579 240 bp overlap
ChIP D341-Med_shGFP GSE92582.NEUROD1.D341-Med_shGFP 263 bp overlap
ChIP D341-Med_shNEUROD1-1579 GSE92582.NEUROD1.D341-Med_shNEUROD1-1579 183 bp overlap
Motif DE_12h DE_12h-NEUROD1_MA1109.2 8 bp overlap
NEUROG2 2 datasets
Motif DE_12h DE_12h-NEUROG2_MA0669.1 10 bp overlap
Motif DE_12h DE_12h-NEUROG2_MA1642.2 7 bp overlap
NIPBL 4 datasets
ChIP hESC GSE64758.NIPBL.hESC 230 bp overlap
ChIP hESC_WNT3A GSE64758.NIPBL.hESC_WNT3A 282 bp overlap
ChIP hESC_WNT3A_ACTIVIN GSE64758.NIPBL.hESC_WNT3A_ACTIVIN 372 bp overlap
ChIP hESC_activin GSE64758.NIPBL.hESC_activin 276 bp overlap
NR2F1 2 datasets
ChIP GM12878 ENCFF273VKX 319 bp overlap
ChIP GM12878 ENCSR514VYD.NR2F1.GM12878 164 bp overlap
OLIG1 1 dataset
Motif DE_12h DE_12h-OLIG1_MA0826.1 10 bp overlap
OLIG2 1 dataset
Motif DE_12h DE_12h-OLIG2_MA0678.1 10 bp overlap
OLIG3 1 dataset
Motif DE_12h DE_12h-OLIG3_MA0827.1 10 bp overlap
ONECUT2 2 datasets
ChIP A-549 GSE102599.ONECUT2.A-549 72 bp overlap
Motif DE_12h DE_12h-ONECUT2_MA0756.3 8 bp overlap
OTX2 1 dataset
ChIP WTC11 ENCFF634NAO 245 bp overlap
PAX5 3 datasets
ChIP GM12878 ENCFF503GOV 227 bp overlap
ChIP GM12878 ENCSR000BHD.PAX5.GM12878 95 bp overlap
ChIP GM12878 ENCSR000BHJ.PAX5.GM12878 202 bp overlap
PCGF1 1 dataset
ChIP WA01 GSE104690.PCGF1.WA01 137 bp overlap
POU1F1 1 dataset
Motif DE_12h DE_12h-POU1F1_MA0784.3 14 bp overlap
POU2F1 1 dataset
Motif DE_12h DE_12h-POU2F1_MA0785.2 9 bp overlap
POU2F2 1 dataset
Motif DE_12h DE_12h-POU2F2_MA0507.3 13 bp overlap
POU2F3 1 dataset
Motif DE_12h DE_12h-POU2F3_MA0627.3 9 bp overlap
POU3F1 1 dataset
Motif DE_12h DE_12h-POU3F1_MA0786.2 10 bp overlap
POU3F2 1 dataset
Motif DE_12h DE_12h-POU3F2_MA0787.1 12 bp overlap
POU3F3 1 dataset
Motif DE_12h DE_12h-POU3F3_MA0788.1 13 bp overlap
POU3F4 1 dataset
Motif DE_12h DE_12h-POU3F4_MA0789.1 9 bp overlap
POU5F1 14 datasets
ChIP BG03 GSE21614.POU5F1.BG03 272 bp overlap
ChIP DE_D1 DED1-OCT4_Batch_II 396 bp overlap
ChIP GM23338 ENCFF333SNB 311 bp overlap
ChIP H1 ENCFF698ZAP 205 bp overlap
ChIP HUES-8 GSE109524.POU5F1.HUES-8 487 bp overlap
ChIP NCCIT_SNF5 GSE36134.POU5F1.NCCIT_SNF5 487 bp overlap
ChIP WA09 GSE105028.POU5F1.WA09 249 bp overlap
ChIP WA09_heat-shock GSE105028.POU5F1.WA09_heat-shock 331 bp overlap
ChIP hESC ENCSR264RJX.POU5F1.hESC 115 bp overlap
ChIP hESC_NAIVE GSE69646.POU5F1.hESC_NAIVE 462 bp overlap
ChIP hESC_PRIMED GSE69646.POU5F1.hESC_PRIMED 487 bp overlap
ChIP hiPSC GSE56567.POU5F1.hiPSC 275 bp overlap
ChIP hiPSC_3s2 GSE81899.POU5F1.hiPSC_3s2 180 bp overlap
ChIP neuron_bipolar_doxy_4d ENCSR362VCG.POU5F1.neuron_bipolar_doxy_4d 127 bp overlap
POU5F1_M 1 dataset
ChIP DE_D1 DED1-OCT4-M_Batch_II 273 bp overlap
Ptf1A 1 dataset
Motif DE_12h DE_12h-Ptf1A_MA1618.2 9 bp overlap
RAD21 2 datasets
ChIP HEK293_siE1A GSE130135.RAD21.HEK293_siE1A 54 bp overlap
ChIP HEK293_siYAP GSE130135.RAD21.HEK293_siYAP 159 bp overlap
RBBP5 2 datasets
ChIP H1 ENCFF905HFL 434 bp overlap
ChIP WA01 ENCSR000AQC.RBBP5.WA01 365 bp overlap
RBPJ 1 dataset
ChIP NHEK GSE29498.RBPJ.NHEK 75 bp overlap
REST 1 dataset
ChIP HeLa-S3 ENCSR000BMN.REST.HeLa-S3 134 bp overlap
SAP30 1 dataset
ChIP WA01 ENCSR000ATR.SAP30.WA01 187 bp overlap
SCRT1 1 dataset
Motif DE_12h DE_12h-SCRT1_MA0743.3 10 bp overlap
SCRT2 1 dataset
Motif DE_12h DE_12h-SCRT2_MA0744.3 10 bp overlap
SIN3A 1 dataset
ChIP WA01 ENCSR000EBO.SIN3A.WA01 120 bp overlap
SMAD2 3 datasets
ChIP hESC GSE29422.SMAD2.hESC 183 bp overlap
ChIP hESC_YAP-_activinA_15h GSE99202.SMAD2.hESC_YAP-_activinA_15h 391 bp overlap
ChIP hESC_activinA_15h GSE99202.SMAD2.hESC_activinA_15h 383 bp overlap
SMAD2-3 4 datasets
ChIP HUES-8 GSE109524.SMAD2-3.HUES-8 372 bp overlap
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 391 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.SMAD2-3.HUES-8_DE_JNKi 487 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.SMAD2-3.HUES-8_DE_ctrl 487 bp overlap
SMAD2_3 5 datasets
ChIP DE_D1 S07-DE-d1-SMAD2_3-exp1 487 bp overlap
ChIP DE_D1 S12-DE-d1-SMAD2_3-exp1 487 bp overlap
ChIP DE_D1 S39-DE-d1-SMAD2_3-exp2 485 bp overlap
ChIP DE_D2 S53-DE-d2-SMAD2_3-exp2 304 bp overlap
ChIP ESC S33-ESC-d0-SMAD2_3-exp2 465 bp overlap
SMAD3 5 datasets
ChIP BG03 GSE21614.SMAD3.BG03 227 bp overlap
ChIP BG03_DIFF_0H GSE36578.SMAD3.BG03_DIFF_0H 242 bp overlap
ChIP BG03_DIFF_2H GSE36578.SMAD3.BG03_DIFF_2H 212 bp overlap
ChIP BG03_DIFF_48H GSE36578.SMAD3.BG03_DIFF_48H 136 bp overlap
ChIP hESC GSE29422.SMAD3.hESC 281 bp overlap
SMAD4 1 dataset
ChIP hESC GSE29422.SMAD4.hESC 175 bp overlap
SMARCA4 2 datasets
ChIP hiPSC GSE124903.SMARCA4.hiPSC 271 bp overlap
ChIP hiPSC_WT GSE124903.SMARCA4.hiPSC_WT 365 bp overlap
SMARCB1 1 dataset
ChIP hiPSC_WT GSE124903.SMARCB1.hiPSC_WT 409 bp overlap
SMARCC1 3 datasets
ChIP DE_D1 S15-DE-d1-BAF155-exp1 379 bp overlap
ChIP hiPSC GSE124903.SMARCC1.hiPSC 314 bp overlap
ChIP hiPSC_WT GSE124903.SMARCC1.hiPSC_WT 396 bp overlap
SOX2 3 datasets
ChIP hESC GSE69479.SOX2.hESC 209 bp overlap
ChIP hESC GSE18292.SOX2.hESC 120 bp overlap
ChIP hiPSC GSE56567.SOX2.hiPSC 305 bp overlap
SP1 2 datasets
ChIP H1 ENCFF263FUH 321 bp overlap
ChIP WA01 ENCSR000BIR.SP1.WA01 206 bp overlap
SPI1 3 datasets
ChIP GM12878 ENCFF134LCP 144 bp overlap
ChIP GM12891 ENCFF563IUT 174 bp overlap
ChIP GM12891 ENCSR000BIJ.SPI1.GM12891 230 bp overlap
SPIB 3 datasets
ChIP OCI-Ly10 GSE56857.SPIB.OCI-Ly10 198 bp overlap
ChIP OCI-Ly3 GSE56857.SPIB.OCI-Ly3 299 bp overlap
ChIP OCI-Ly3_SHCTR GSE56857.SPIB.OCI-Ly3_SHCTR 267 bp overlap
STAT3 2 datasets
ChIP TMD8 GSE106844.STAT3.TMD8 240 bp overlap
ChIP TMD8_DMSO GSE123398.STAT3.TMD8_DMSO 171 bp overlap
TCF12 6 datasets
ChIP GM12878 ENCFF433DMU 254 bp overlap
ChIP GM12878 ENCFF506WWB 153 bp overlap
ChIP GM12878 ENCSR725VFL.TCF12.GM12878 205 bp overlap
ChIP GM12878 ENCSR000BGZ.TCF12.GM12878 224 bp overlap
ChIP H1 ENCFF203EBH 249 bp overlap
ChIP WA01 ENCSR000BIT.TCF12.WA01 315 bp overlap
TCF21 1 dataset
Motif DE_12h DE_12h-TCF21_MA1568.2 10 bp overlap
TCF3 3 datasets
ChIP GM12878 ENCFF658WIO 83 bp overlap
ChIP GM12878 ENCSR000BQT.TCF3.GM12878 286 bp overlap
ChIP NPC GSE154479.TCF3.NPC 484 bp overlap
TCF4 2 datasets
ChIP CAL-1 GSE76147.TCF4.CAL-1 283 bp overlap
ChIP GEN2-2 GSE76147.TCF4.GEN2-2 263 bp overlap
TFAP2C 1 dataset
ChIP UCLA1-hESCs_TFAP2C--_2d_TFAP2C_Induc GSE101074.TFAP2C.UCLA1-hESCs_TFAP2C--_2d_TFAP2C_Induc 487 bp overlap
TFAP4 1 dataset
Motif DE_12h DE_12h-TFAP4_MA1570.1 10 bp overlap
TP53 1 dataset
ChIP H9_ectoderm GSE142050.TP53.H9_ectoderm 456 bp overlap
Tcf21 1 dataset
Motif DE_12h DE_12h-Tcf21_MA0832.2 10 bp overlap
YY1 1 dataset
ChIP WA01 ENCSR000BKD.YY1.WA01 152 bp overlap
ZBTB21 1 dataset
ChIP WTC11 ENCFF677ZYY 365 bp overlap
ZEB1 1 dataset
ChIP GM12878 ENCSR000BND.ZEB1.GM12878 125 bp overlap
ZNF207 1 dataset
ChIP WA09 GSE118632.ZNF207.WA09 250 bp overlap
ZNF462 2 datasets
ChIP GM23338 ENCFF896CCA 216 bp overlap
ChIP GM23338 ENCSR334UWP.ZNF462.GM23338 271 bp overlap