chr4 : 56,055,093 56,055,833
740 bp 115 TFs 3 linked genes
This 740 bp open chromatin element is linked to CRACD, CEP135, and EXOC1 and is bound by 115 transcription factors.
Linked Genes
3 genes
Distance
Link type
Gene Expression Dist. to TSS Distance Link type
CRACD 6.0 kb Proximal Proximity
CEP135 106.5 kb Distal Multiome
EXOC1 201.7 kb Distal Multiome
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr4:56,050,093 – 56,060,833
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
115 transcription factors
Source
Cell type
AR 1 dataset
ChIP LNCaP_1F5_SIFOXA1 GSE30623.AR.LNCaP_1F5_SIFOXA1 143 bp overlap
ARID3A 1 dataset
ChIP HepG2 ENCFF341DES 272 bp overlap
CEBPA 2 datasets
ChIP Kasumi-1_CEBPA-ER GSE102697.CEBPA.Kasumi-1_CEBPA-ER 158 bp overlap
ChIP Kasumi-1_CEBPA-ER_E2 GSE102697.CEBPA.Kasumi-1_CEBPA-ER_E2 343 bp overlap
CHD4 1 dataset
ChIP SCMC GSE155861.CHD4.SCMC 160 bp overlap
CHD7 1 dataset
ChIP hiPSC_derived_lt-NES GSE108506.CHD7.hiPSC_derived_lt-NES 210 bp overlap
Cebpa 4 datasets
ChIP BLaER1 ENCFF093OYK 437 bp overlap
ChIP BLaER1 ENCFF335XTP 257 bp overlap
ChIP BLaER1 ENCFF346MCV 136 bp overlap
ChIP BLaER1 ENCFF460KDD 397 bp overlap
E2F6 3 datasets
Motif DE_36h DE_36h-E2F6_MA0471.3 8 bp overlap
Motif DE_48h DE_48h-E2F6_MA0471.3 8 bp overlap
Motif DE_60h DE_60h-E2F6_MA0471.3 8 bp overlap
EBF1 2 datasets
Motif DE_36h DE_36h-EBF1_MA0154.5 11 bp overlap
Motif DE_60h DE_60h-EBF1_MA0154.5 11 bp overlap
EGR1 1 dataset
ChIP fibroblast_DOX-24h GSE134924.EGR1.fibroblast_DOX-24h 200 bp overlap
EOMES 1 dataset
ChIP hESC GSE26097.EOMES.hESC 463 bp overlap
ETS1 1 dataset
ChIP CTV-1_DSG GSE128835.ETS1.CTV-1_DSG 172 bp overlap
EWSR1-FLI1 4 datasets
Motif DE_36h DE_36h-EWSR1-FLI1_MA0149.1 18 bp overlap
Motif DE_48h DE_48h-EWSR1-FLI1_MA0149.1 18 bp overlap
Motif DE_60h DE_60h-EWSR1-FLI1_MA0149.1 18 bp overlap
Motif DE_72h DE_72h-EWSR1-FLI1_MA0149.1 18 bp overlap
Ebf4 2 datasets
Motif DE_36h DE_36h-Ebf4_MA2122.1 11 bp overlap
Motif DE_60h DE_60h-Ebf4_MA2122.1 11 bp overlap
Elf5 4 datasets
Motif DE_36h DE_36h-Elf5_MA0136.4 8 bp overlap
Motif DE_48h DE_48h-Elf5_MA0136.4 8 bp overlap
Motif DE_60h DE_60h-Elf5_MA0136.4 8 bp overlap
Motif DE_72h DE_72h-Elf5_MA0136.4 8 bp overlap
FIGLA 2 datasets
Motif DE_36h DE_36h-FIGLA_MA0820.2 6 bp overlap
Motif DE_60h DE_60h-FIGLA_MA0820.2 6 bp overlap
FOXA1 1 dataset
ChIP pancreatic-progenitor_PP1 GSE104840.FOXA1.pancreatic-progenitor_PP1 239 bp overlap
FOXA2 3 datasets
ChIP DE DE-FOXA2-1 704 bp overlap
ChIP DE DE-FOXA2-2 607 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.FOXA2.pancreatic-progenitor_PP1 339 bp overlap
GATA1 5 datasets
ChIP K-562_dCas9-LSD1 GSE132212.GATA1.K-562_dCas9-LSD1 145 bp overlap
ChIP K-562_enCRISPRi-LK GSE132212.GATA1.K-562_enCRISPRi-LK 197 bp overlap
ChIP erythroblast ENCFF867JAR 522 bp overlap
ChIP erythroblast ENCSR000EXP.GATA1.erythroblast 350 bp overlap
ChIP erythroid-progenitor GSE124163.GATA1.erythroid-progenitor 156 bp overlap
GATA2 6 datasets
Motif DE_36h DE_36h-GATA2_MA0036.4 7 bp overlap
Motif DE_36h DE_36h-GATA2_MA0036.4 7 bp overlap
Motif DE_48h DE_48h-GATA2_MA0036.4 7 bp overlap
Motif DE_60h DE_60h-GATA2_MA0036.4 7 bp overlap
Motif DE_72h DE_72h-GATA2_MA0036.4 7 bp overlap
ChIP TF1 GSE73207.GATA2.TF1 166 bp overlap
GATA3 1 dataset
ChIP MCF-7_E2 GSE81510.GATA3.MCF-7_E2 210 bp overlap
GATA4 10 datasets
ChIP DE DE-GATA4-1 735 bp overlap
ChIP DE DE-GATA4-2 720 bp overlap
Motif DE_36h DE_36h-GATA4_MA0482.3 8 bp overlap
Motif DE_36h DE_36h-GATA4_MA0482.3 8 bp overlap
Motif DE_48h DE_48h-GATA4_MA0482.3 8 bp overlap
Motif DE_60h DE_60h-GATA4_MA0482.3 8 bp overlap
Motif DE_72h DE_72h-GATA4_MA0482.3 8 bp overlap
ChIP foregut GSE117136.GATA4.foregut 345 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA4.pancreatic-progenitor 532 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA4.pancreatic-progenitor_PP1 740 bp overlap
GATA5 5 datasets
Motif DE_36h DE_36h-GATA5_MA0766.3 8 bp overlap
Motif DE_36h DE_36h-GATA5_MA0766.3 8 bp overlap
Motif DE_48h DE_48h-GATA5_MA0766.3 8 bp overlap
Motif DE_60h DE_60h-GATA5_MA0766.3 8 bp overlap
Motif DE_72h DE_72h-GATA5_MA0766.3 8 bp overlap
GATA6 22 datasets
ChIP AGS GSE51705.GATA6.AGS 193 bp overlap
ChIP DE DE-GATA6-1 620 bp overlap
ChIP DE DE-GATA6-2 740 bp overlap
Motif DE_36h DE_36h-GATA6_MA1104.3 8 bp overlap
Motif DE_36h DE_36h-GATA6_MA1104.3 8 bp overlap
Motif DE_48h DE_48h-GATA6_MA1104.3 8 bp overlap
Motif DE_60h DE_60h-GATA6_MA1104.3 8 bp overlap
Motif DE_72h DE_72h-GATA6_MA1104.3 8 bp overlap
ChIP DE_D1 S09-DE-d1-GATA6-exp1 489 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 432 bp overlap
ChIP DE_D1 S41-DE-d1-GATA6-exp2 640 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 503 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 620 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 673 bp overlap
ChIP HUG1N GSE51936.GATA6.HUG1N 164 bp overlap
ChIP YCC-3 GSE51705.GATA6.YCC-3 226 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 466 bp overlap
ChIP foregut GSE117136.GATA6.foregut 392 bp overlap
ChIP foregut_KO GSE117136.GATA6.foregut_KO 397 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA6.pancreatic-progenitor 373 bp overlap
ChIP pancreatic-progenitor_KO GSE117136.GATA6.pancreatic-progenitor_KO 289 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA6.pancreatic-progenitor_PP1 505 bp overlap
GRHL1 1 dataset
Motif DE_36h DE_36h-GRHL1_MA0647.2 10 bp overlap
GRHL2 1 dataset
Motif DE_36h DE_36h-GRHL2_MA1105.3 8 bp overlap
Gata3 5 datasets
Motif DE_36h DE_36h-Gata3_MA0037.5 8 bp overlap
Motif DE_36h DE_36h-Gata3_MA0037.5 8 bp overlap
Motif DE_48h DE_48h-Gata3_MA0037.5 8 bp overlap
Motif DE_60h DE_60h-Gata3_MA0037.5 8 bp overlap
Motif DE_72h DE_72h-Gata3_MA0037.5 8 bp overlap
HAND2 1 dataset
ChIP SK-N-BE2-C GSE94822.HAND2.SK-N-BE2-C 109 bp overlap
HNF1A 1 dataset
Motif DE_36h DE_36h-HNF1A_MA0046.3 13 bp overlap
HNF1B 1 dataset
Motif DE_36h DE_36h-HNF1B_MA0153.2 13 bp overlap
HNF4A 4 datasets
ChIP Caco-2_DIFF GSE23436.HNF4A.Caco-2_DIFF 104 bp overlap
ChIP Hep-G2 ENCSR000BLF.HNF4A.Hep-G2 173 bp overlap
ChIP Hep-G2 ENCSR469FBY.HNF4A.Hep-G2 159 bp overlap
ChIP gastric-epithelial-cell_OE GSE114018.HNF4A.gastric-epithelial-cell_OE 69 bp overlap
HOXB13 1 dataset
ChIP G-401 GSE65381.HOXB13.G-401 187 bp overlap
Hmga1 1 dataset
Motif DE_36h DE_36h-Hmga1_MA2124.1 8 bp overlap
IKZF2 4 datasets
Motif DE_36h DE_36h-IKZF2_MA2326.1 6 bp overlap
Motif DE_48h DE_48h-IKZF2_MA2326.1 6 bp overlap
Motif DE_60h DE_60h-IKZF2_MA2326.1 6 bp overlap
Motif DE_72h DE_72h-IKZF2_MA2326.1 6 bp overlap
KDM1A 1 dataset
ChIP pancreatic-progenitor_PP2-KDM1A-inh GSE104840.KDM1A.pancreatic-progenitor_PP2-KDM1A-inh 137 bp overlap
KMT2A 4 datasets
ChIP HEK293T_N-term_C104 GSE90762.KMT2A.HEK293T_N-term_C104 404 bp overlap
ChIP HEK293T_N-term_C49 GSE90762.KMT2A.HEK293T_N-term_C49 313 bp overlap
ChIP HEK293T_N-term_shCtrl_TASP1-KO GSE90762.KMT2A.HEK293T_N-term_shCtrl_TASP1-KO 441 bp overlap
ChIP Hep-G2 ENCSR692GFR.KMT2A.Hep-G2 175 bp overlap
LIN54 1 dataset
Motif DE_36h DE_36h-LIN54_MA0619.2 7 bp overlap
Lhx3 1 dataset
Motif DE_36h DE_36h-Lhx3_MA0135.2 12 bp overlap
MAZ 1 dataset
ChIP HEK293 GSE76494.MAZ.HEK293 210 bp overlap
MEF2B 1 dataset
ChIP tonsil GSE110682.MEF2B.tonsil 58 bp overlap
MIXL1 1 dataset
ChIP HepG2 ENCFF817YFO 313 bp overlap
MYC 1 dataset
ChIP Jurkat GSE83777.MYC.Jurkat 192 bp overlap
Mecom 1 dataset
Motif DE_36h DE_36h-Mecom_MA0029.2 11 bp overlap
NEUROD1 4 datasets
ChIP D283-Med GSE92582.NEUROD1.D283-Med 293 bp overlap
ChIP D283-Med_shNEUROD1-1579 GSE92582.NEUROD1.D283-Med_shNEUROD1-1579 185 bp overlap
ChIP D341-Med_shGFP GSE92582.NEUROD1.D341-Med_shGFP 106 bp overlap
ChIP D341-Med_shNEUROD1-1579 GSE92582.NEUROD1.D341-Med_shNEUROD1-1579 168 bp overlap
NFYB 1 dataset
Motif DE_36h DE_36h-NFYB_MA0502.3 9 bp overlap
NKX2-2 1 dataset
Motif DE_36h DE_36h-NKX2-2_MA1645.2 8 bp overlap
NOTCH1 1 dataset
ChIP GSC8-11_dasatinib GSE74557.NOTCH1.GSC8-11_dasatinib 63 bp overlap
NR2F2 1 dataset
ChIP Hep-G2 ENCSR000BVM.NR2F2.Hep-G2 125 bp overlap
NR2F6 2 datasets
Motif DE_36h DE_36h-NR2F6_MA1539.1 15 bp overlap
ChIP Hep-G2 ENCSR518WPL.NR2F6.Hep-G2 175 bp overlap
ONECUT1 3 datasets
ChIP H9 ERP004206.ONECUT1.H9 142 bp overlap
ChIP Hep-G2 ENCSR956OSX.ONECUT1.Hep-G2 107 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.ONECUT1.pancreatic-progenitor_PP1 119 bp overlap
PATZ1 1 dataset
ChIP HEK293 ENCSR966ULI.PATZ1.HEK293 399 bp overlap
PDX1 3 datasets
ChIP H9 ERP004206.PDX1.H9 163 bp overlap
ChIP hiPSC GSE125768.PDX1.hiPSC 309 bp overlap
ChIP hiPSC_derived_PPs GSE106949.PDX1.hiPSC_derived_PPs 192 bp overlap
PHIP 1 dataset
ChIP HEK293_ab833 GSE101646.PHIP.HEK293_ab833 292 bp overlap
PHOX2A 1 dataset
Motif DE_36h DE_36h-PHOX2A_MA0713.1 11 bp overlap
PHOX2B 1 dataset
Motif DE_36h DE_36h-PHOX2B_MA0681.3 12 bp overlap
PITX3 1 dataset
ChIP SH-SY5Y GSE93275.PITX3.SH-SY5Y 191 bp overlap
POU2F1 1 dataset
Motif DE_36h DE_36h-POU2F1_MA0785.2 9 bp overlap
POU3F1 1 dataset
Motif DE_36h DE_36h-POU3F1_MA0786.2 10 bp overlap
POU3F2 1 dataset
Motif DE_36h DE_36h-POU3F2_MA0787.1 12 bp overlap
POU3F3 1 dataset
Motif DE_36h DE_36h-POU3F3_MA0788.1 13 bp overlap
POU4F2 1 dataset
ChIP HNPC_DIF GSE74814.POU4F2.HNPC_DIF 92 bp overlap
PRDM10 1 dataset
ChIP HEK293 ENCFF145WQQ 263 bp overlap
PRDM15 1 dataset
ChIP WTC11 ENCFF108TMF 342 bp overlap
PRDM6 1 dataset
ChIP HEK293 ENCFF283AJL 107 bp overlap
PROP1 1 dataset
Motif DE_36h DE_36h-PROP1_MA0715.1 11 bp overlap
PSIP1 1 dataset
ChIP T-REx-293 GSE117155.PSIP1.T-REx-293 210 bp overlap
Plagl1 1 dataset
Motif DE_36h DE_36h-Plagl1_MA1615.2 8 bp overlap
Prdm4 1 dataset
Motif DE_36h DE_36h-Prdm4_MA1647.3 11 bp overlap
RARA 1 dataset
ChIP hiPSC_D5 GSE132532.RARA.hiPSC_D5 248 bp overlap
RBPJ 1 dataset
ChIP GSC8-11_dasatinib GSE74557.RBPJ.GSC8-11_dasatinib 69 bp overlap
RELA 1 dataset
ChIP SGBS_TNF GSE64233.RELA.SGBS_TNF 467 bp overlap
SCRT2 1 dataset
Motif DE_36h DE_36h-SCRT2_MA0744.3 10 bp overlap
SIX2 1 dataset
ChIP HEK GSE73865.SIX2.HEK 170 bp overlap
SMAD2 1 dataset
ChIP endoderm GSE29422.SMAD2.endoderm 148 bp overlap
SMAD2-3 2 datasets
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 566 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.SMAD2-3.HUES-8_DE_JNKi 405 bp overlap
SMAD2_3 3 datasets
ChIP DE_D1 S12-DE-d1-SMAD2_3-exp1 325 bp overlap
ChIP DE_D2 S01-DE-d2-SMAD2_3-exp1 511 bp overlap
ChIP DE_D2 S53-DE-d2-SMAD2_3-exp2 552 bp overlap
SMAD3 1 dataset
ChIP endoderm GSE29422.SMAD3.endoderm 225 bp overlap
SMAD4 1 dataset
ChIP endoderm GSE29422.SMAD4.endoderm 222 bp overlap
SMARCA4 9 datasets
ChIP CTV-1_DSG GSE128835.SMARCA4.CTV-1_DSG 293 bp overlap
ChIP CTV-1_FLAG GSE128835.SMARCA4.CTV-1_FLAG 244 bp overlap
ChIP CTV-1_delQ GSE128835.SMARCA4.CTV-1_delQ 182 bp overlap
ChIP NPC_K755R-siAP GSE122631.SMARCA4.NPC_K755R-siAP 198 bp overlap
ChIP NPC_K755R-siCT GSE122631.SMARCA4.NPC_K755R-siCT 328 bp overlap
ChIP NPC_siCT GSE122631.SMARCA4.NPC_siCT 334 bp overlap
ChIP NPC_siSOX GSE122631.SMARCA4.NPC_siSOX 253 bp overlap
ChIP NPC_siSOX GSE122631.SMARCA4.NPC_siSOX 183 bp overlap
ChIP NSC GSE125033.SMARCA4.NSC 451 bp overlap
SMARCB1 1 dataset
ChIP hiPSC_WT GSE124903.SMARCB1.hiPSC_WT 170 bp overlap
SMARCC1 1 dataset
ChIP hiPSC_WT GSE124903.SMARCC1.hiPSC_WT 212 bp overlap
SOX13 1 dataset
ChIP HepG2 ENCFF062VSQ 267 bp overlap
SOX17_M 1 dataset
ChIP DE_D2 DED2-SOX17-M_Batch_II 454 bp overlap
SOX2 1 dataset
ChIP HNSC GSE69479.SOX2.HNSC 185 bp overlap
SP1 2 datasets
Motif DE_36h DE_36h-SP1_MA0079.5 9 bp overlap
Motif DE_60h DE_60h-SP1_MA0079.5 9 bp overlap
SP2 3 datasets
Motif DE_36h DE_36h-SP2_MA0516.3 9 bp overlap
Motif DE_60h DE_60h-SP2_MA0516.3 9 bp overlap
ChIP K-562 ENCSR000BNL.SP2.K-562 113 bp overlap
SP5 2 datasets
Motif DE_36h DE_36h-SP5_MA1965.2 6 bp overlap
Motif DE_60h DE_60h-SP5_MA1965.2 6 bp overlap
SREBF1 1 dataset
Motif DE_36h DE_36h-SREBF1_MA0595.1 10 bp overlap
SREBF2 1 dataset
Motif DE_36h DE_36h-SREBF2_MA0596.1 10 bp overlap
SRF 2 datasets
ChIP Hep-G2 ENCSR000BLV.SRF.Hep-G2 156 bp overlap
ChIP HepG2 ENCFF625QHW 185 bp overlap
STAT3 1 dataset
ChIP MCF-7_jc5836 GSE126004.STAT3.MCF-7_jc5836 251 bp overlap
SUZ12 1 dataset
ChIP CRL-7250_shSS18 GSE108025.SUZ12.CRL-7250_shSS18 296 bp overlap
TFDP1 3 datasets
Motif DE_36h DE_36h-TFDP1_MA1122.2 8 bp overlap
Motif DE_48h DE_48h-TFDP1_MA1122.2 8 bp overlap
Motif DE_60h DE_60h-TFDP1_MA1122.2 8 bp overlap
THRB 1 dataset
ChIP Hep-G2 ENCSR430JGJ.THRB.Hep-G2 190 bp overlap
TRPS1 5 datasets
Motif DE_36h DE_36h-TRPS1_MA1970.2 8 bp overlap
Motif DE_36h DE_36h-TRPS1_MA1970.2 8 bp overlap
Motif DE_48h DE_48h-TRPS1_MA1970.2 8 bp overlap
Motif DE_60h DE_60h-TRPS1_MA1970.2 8 bp overlap
Motif DE_72h DE_72h-TRPS1_MA1970.2 8 bp overlap
TWIST1 2 datasets
ChIP BE2C GSE80151.TWIST1.BE2C 113 bp overlap
ChIP SK-N-BE2-C GSE80151.TWIST1.SK-N-BE2-C 113 bp overlap
YY1 2 datasets
ChIP Hep-G2_NC GSE120104.YY1.Hep-G2_NC 314 bp overlap
ChIP Hep-G2_RBM25-KD GSE120104.YY1.Hep-G2_RBM25-KD 489 bp overlap
ZBTB26 1 dataset
Motif DE_36h DE_36h-ZBTB26_MA1579.2 8 bp overlap
ZEB1 3 datasets
Motif DE_36h DE_36h-ZEB1_MA0103.4 6 bp overlap
Motif DE_60h DE_60h-ZEB1_MA0103.4 6 bp overlap
ChIP neuron_bipolar_doxy_4d ENCSR418KUS.ZEB1.neuron_bipolar_doxy_4d 193 bp overlap
ZNF207 1 dataset
ChIP WA09 GSE118632.ZNF207.WA09 245 bp overlap
ZNF467 1 dataset
ChIP HEK293 GSE76494.ZNF467.HEK293 148 bp overlap
ZNF652 2 datasets
ChIP Hep-G2 ENCSR502GAX.ZNF652.Hep-G2 665 bp overlap
ChIP HepG2 ENCFF331VPZ 195 bp overlap
ZNF770 2 datasets
ChIP HEK293 ENCSR242BGR.ZNF770.HEK293 302 bp overlap
ChIP HEK293 GSE76494.ZNF770.HEK293 203 bp overlap