chr4 : 87,974,968 87,975,292
324 bp 100 TFs 1 linked gene
This 324 bp open chromatin element is linked to SPP1 and is bound by 100 transcription factors.
Linked Genes
1 gene
Gene Expression Dist. to TSS Distance Link type
SPP1 417 bp At TSS Proximity
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr4:87,969,968 – 87,980,292
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
100 transcription factors
Source
Cell type
AR 1 dataset
ChIP LNCaP_androgen-N_hypoxia-Y GSE114732.AR.LNCaP_androgen-N_hypoxia-Y 139 bp overlap
ARID1A 3 datasets
ChIP RMG-I GSE104545.ARID1A.RMG-I 324 bp overlap
ChIP RMG-I GSE120058.ARID1A.RMG-I 324 bp overlap
ChIP endometrial-epithelial-cells GSE106660.ARID1A.endometrial-epithelial-cells 168 bp overlap
ARID5B 1 dataset
ChIP HepG2 ENCFF964FWK 254 bp overlap
ASH2L 1 dataset
ChIP Hep-G2 ENCSR508LMH.ASH2L.Hep-G2 199 bp overlap
ATF3 1 dataset
ChIP liver ENCSR205FOW.ATF3.liver 142 bp overlap
ATRX 1 dataset
ChIP metastatic-neuroblastoma_SKNMM GSE100148.ATRX.metastatic-neuroblastoma_SKNMM 259 bp overlap
BACH1 5 datasets
ChIP A-549 ENCSR043EHG.BACH1.A-549 213 bp overlap
ChIP H1 ENCFF282VDB 216 bp overlap
ChIP Hep-G2 ENCSR699TNT.BACH1.Hep-G2 185 bp overlap
ChIP SW1990 GSE124406.BACH1.SW1990 186 bp overlap
ChIP WA01 ENCSR000EBQ.BACH1.WA01 170 bp overlap
BACH2 1 dataset
ChIP B-cell_IL2 GSE102460.BACH2.B-cell_IL2 183 bp overlap
BRD4 3 datasets
ChIP LNCaP-C4-2_OE GSE88871.BRD4.LNCaP-C4-2_OE 85 bp overlap
ChIP SUM229PE_pos_DMSO_24h GSE87418.BRD4.SUM229PE_pos_DMSO_24h 200 bp overlap
ChIP hESC GSE33281.BRD4.hESC 61 bp overlap
CBX2 1 dataset
ChIP K-562 ENCSR000ATU.CBX2.K-562 171 bp overlap
CEBPA 1 dataset
ChIP liver ERP002306.CEBPA.liver 133 bp overlap
CEBPB 6 datasets
ChIP A-549 ENCSR000BUB.CEBPB.A-549 131 bp overlap
ChIP A549 ENCFF235AIY 165 bp overlap
ChIP HCT116 ENCFF097OLY 114 bp overlap
ChIP Hep-G2 ENCSR000EEX.CEBPB.Hep-G2 113 bp overlap
ChIP Ishikawa ENCFF010USJ 154 bp overlap
ChIP Ishikawa ENCSR000BTT.CEBPB.Ishikawa 128 bp overlap
CEBPG 1 dataset
ChIP HepG2 ENCFF503XBC 177 bp overlap
CTCF 7 datasets
ChIP HT-1080 GSE135580.CTCF.HT-1080 123 bp overlap
ChIP anterior-temporal-cortex_adult GSE116825.CTCF.anterior-temporal-cortex_adult 152 bp overlap
ChIP brain ENCFF163BBN 126 bp overlap
ChIP chondrocyte ENCFF134ORZ 307 bp overlap
ChIP dorsolateral prefrontal cortex ENCFF359BHR 123 bp overlap
ChIP hepatocyte ERP000395.CTCF.hepatocyte 101 bp overlap
ChIP neural progenitor cell ENCFF581WPG 69 bp overlap
Cebpa 1 dataset
ChIP BLaER1 ENCFF896HSY 111 bp overlap
DAXX 2 datasets
ChIP PC-3 GSE68647.DAXX.PC-3 168 bp overlap
ChIP PC-3_DAXX GSE68647.DAXX.PC-3_DAXX 164 bp overlap
DMAP1 1 dataset
ChIP Hep-G2 ENCSR670YPQ.DMAP1.Hep-G2 128 bp overlap
DRAP1 1 dataset
ChIP Hep-G2 ENCSR765MKZ.DRAP1.Hep-G2 104 bp overlap
ELF3 2 datasets
ChIP PDAC_KOKLF5 GSE64557.ELF3.PDAC_KOKLF5 324 bp overlap
ChIP PDAC_SHCTR GSE64557.ELF3.PDAC_SHCTR 54 bp overlap
EMSY 1 dataset
ChIP K562 ENCFF511ZZZ 306 bp overlap
EP300 2 datasets
ChIP A-549 ENCSR000BPW.EP300.A-549 116 bp overlap
ChIP Hep-G2 ENCSR000BLW.EP300.Hep-G2 113 bp overlap
ESR1 14 datasets
ChIP Ishikawa ENCSR000BIY.ESR1.Ishikawa 146 bp overlap
ChIP Ishikawa ENCSR000BKL.ESR1.Ishikawa 83 bp overlap
ChIP Ishikawa_Dex_E2 GSE109891.ESR1.Ishikawa_Dex_E2 98 bp overlap
ChIP Ishikawa_E2 GSE132426.ESR1.Ishikawa_E2 166 bp overlap
ChIP Ishikawa_E2 GSE109891.ESR1.Ishikawa_E2 100 bp overlap
ChIP Ishikawa_E2_D538G_clone2 GSE132426.ESR1.Ishikawa_E2_D538G_clone2 222 bp overlap
ChIP Ishikawa_ETV4-KO1_Mili GSE129803.ESR1.Ishikawa_ETV4-KO1_Mili 195 bp overlap
ChIP Ishikawa_ETV4-KO2_Mili GSE129803.ESR1.Ishikawa_ETV4-KO2_Mili 173 bp overlap
ChIP Ishikawa_ETV4-OE GSE129803.ESR1.Ishikawa_ETV4-OE 234 bp overlap
ChIP Ishikawa_M1-66 GSE147141.ESR1.Ishikawa_M1-66 142 bp overlap
ChIP Ishikawa_siETV4-72h GSE129803.ESR1.Ishikawa_siETV4-72h 186 bp overlap
ChIP Ishikawa_siNTC-72h GSE129803.ESR1.Ishikawa_siNTC-72h 162 bp overlap
ChIP endometrioid-adenocarcinoma_tumor_3 GSE94031.ESR1.endometrioid-adenocarcinoma_tumor_3 162 bp overlap
ChIP primary-endometrium-cancer_E1_DSG GSE114737.ESR1.primary-endometrium-cancer_E1_DSG 324 bp overlap
EZH2 3 datasets
ChIP DND41 ENCSR000ASW.EZH2.DND41 193 bp overlap
ChIP DND41 ENCSR000ASW.EZH2.DND41 75 bp overlap
ChIP Jurkat_KO GSE147198.EZH2.Jurkat_KO 65 bp overlap
FEZF2 1 dataset
Motif DE_12h DE_12h-FEZF2_MA2341.1 8 bp overlap
FOSL2 1 dataset
ChIP HepG2 ENCFF548CXY 240 bp overlap
FOXA1 2 datasets
ChIP PDAC_KOKLF5 GSE64557.FOXA1.PDAC_KOKLF5 324 bp overlap
ChIP PDAC_SHCTR GSE64557.FOXA1.PDAC_SHCTR 227 bp overlap
HDAC2 1 dataset
ChIP Hep-G2 ENCSR000BMC.HDAC2.Hep-G2 118 bp overlap
HNF1A 1 dataset
ChIP HEE_1 GSE76376.HNF1A.HEE_1 157 bp overlap
HNF1B 1 dataset
ChIP PDAC GSE64557.HNF1B.PDAC 72 bp overlap
HNF4A 2 datasets
ChIP liver ENCFF354NRH 273 bp overlap
ChIP liver ENCFF449HPV 283 bp overlap
JUN 1 dataset
ChIP HAEC_oxpapc_4h GSE89970.JUN.HAEC_oxpapc_4h 130 bp overlap
JUND 2 datasets
ChIP liver ENCFF007WWT 290 bp overlap
ChIP liver ENCSR837GTK.JUND.liver 118 bp overlap
KDM1A 3 datasets
ChIP HepG2 ENCFF240UWG 324 bp overlap
ChIP HepG2 ENCFF240UWG 303 bp overlap
ChIP HepG2 ENCFF240UWG 97 bp overlap
KLF6 1 dataset
ChIP Hep-G2 ENCSR757EKM.KLF6.Hep-G2 139 bp overlap
MAF::NFE2 2 datasets
Motif DE_12h DE_12h-MAFNFE2_MA0501.2 11 bp overlap
Motif DE_24h DE_24h-MAFNFE2_MA0501.2 11 bp overlap
MAFB 1 dataset
ChIP islet ERP004003.MAFB.islet 181 bp overlap
MAFF 2 datasets
Motif DE_12h DE_12h-MAFF_MA0495.4 11 bp overlap
ChIP HepG2 ENCFF452YUT 207 bp overlap
MAFG 2 datasets
ChIP K-562 ENCSR818DQV.MAFG.K-562 176 bp overlap
ChIP K562 ENCFF455EEO 266 bp overlap
MAFG::NFE2L1 2 datasets
Motif DE_12h DE_12h-MAFGNFE2L1_MA0089.3 11 bp overlap
Motif DE_24h DE_24h-MAFGNFE2L1_MA0089.3 11 bp overlap
MAFK 9 datasets
ChIP A549 ENCFF371EPR 147 bp overlap
Motif DE_12h DE_12h-MAFK_MA0496.4 10 bp overlap
Motif DE_24h DE_24h-MAFK_MA0496.4 10 bp overlap
ChIP H1 ENCFF854XWE 200 bp overlap
ChIP Hep-G2 ENCSR000EDZ.MAFK.Hep-G2 151 bp overlap
ChIP HepG2 ENCFF743ZOF 185 bp overlap
ChIP HepG2 ENCFF767LDG 163 bp overlap
ChIP OCI-Ly7 GSE47784.MAFK.OCI-Ly7 142 bp overlap
ChIP WA01 ENCSR000EBS.MAFK.WA01 157 bp overlap
MAX 1 dataset
ChIP Ishikawa ENCSR000BTY.MAX.Ishikawa 115 bp overlap
MED26 2 datasets
ChIP HCT-116_aux GSE121355.MED26.HCT-116_aux 324 bp overlap
ChIP HEK293T GSE121024.MED26.HEK293T 148 bp overlap
MEIS1 2 datasets
Motif DE_12h DE_12h-MEIS1_MA0498.3 5 bp overlap
Motif DE_24h DE_24h-MEIS1_MA0498.3 5 bp overlap
MYCN 2 datasets
ChIP SHEP-21N GSE80151.MYCN.SHEP-21N 180 bp overlap
ChIP SHEP-21N_24h GSE80151.MYCN.SHEP-21N_24h 225 bp overlap
MYPOP 1 dataset
ChIP HepG2 ENCFF176TQL 132 bp overlap
NFE2 7 datasets
ChIP HepG2 ENCFF403RMK 201 bp overlap
ChIP K-562 ENCSR552YGL.NFE2.K-562 240 bp overlap
ChIP K-562 ENCSR000FCC.NFE2.K-562 158 bp overlap
ChIP K562 ENCFF047YKA 276 bp overlap
ChIP K562 ENCFF163BSI 198 bp overlap
ChIP ProEs GSE59087.NFE2.ProEs 96 bp overlap
ChIP erythroid_R3R4 GSE43625.NFE2.erythroid_R3R4 61 bp overlap
NFE2L2 8 datasets
ChIP A-549 GSE113497.NFE2L2.A-549 324 bp overlap
ChIP A-549 ENCSR584GHV.NFE2L2.A-549 183 bp overlap
ChIP A-549 GSE141497.NFE2L2.A-549 145 bp overlap
ChIP A549 ENCFF474YMB 152 bp overlap
ChIP HeLa-S3 ENCSR707IUN.NFE2L2.HeLa-S3 116 bp overlap
ChIP Hep-G2 ENCSR488EES.NFE2L2.Hep-G2 189 bp overlap
ChIP HepG2 ENCFF178DRC 114 bp overlap
ChIP IMR-90 ENCSR197WGI.NFE2L2.IMR-90 128 bp overlap
NFIA 2 datasets
ChIP Hep-G2 GSE97661.NFIA.Hep-G2 110 bp overlap
ChIP HepG2 ENCFF815HWK 128 bp overlap
NFIC 2 datasets
ChIP Ishikawa ENCFF029AAD 213 bp overlap
ChIP Ishikawa ENCSR000BUT.NFIC.Ishikawa 131 bp overlap
NFIL3 1 dataset
ChIP HepG2 ENCFF686VLI 156 bp overlap
NFIX 1 dataset
Motif DE_12h DE_12h-NFIX_MA1528.2 14 bp overlap
NIPBL 1 dataset
ChIP HCT-116_RAD21-mAC_500uM_auxin GSE104888.NIPBL.HCT-116_RAD21-mAC_500uM_auxin 324 bp overlap
NR2F1 1 dataset
ChIP HepG2 ENCFF518ZRY 292 bp overlap
NR2F2 4 datasets
ChIP liver ENCFF427MRU 243 bp overlap
ChIP liver ENCFF565JGD 286 bp overlap
ChIP liver ENCSR168SMX.NR2F2.liver 259 bp overlap
ChIP liver ENCSR338MMB.NR2F2.liver 206 bp overlap
NR2F6 1 dataset
ChIP Hep-G2 ENCSR518WPL.NR2F6.Hep-G2 186 bp overlap
NR3C1 2 datasets
ChIP Ishikawa ENCSR000BJC.NR3C1.Ishikawa 130 bp overlap
ChIP Ishikawa_Dex GSE109891.NR3C1.Ishikawa_Dex 111 bp overlap
NR5A1 1 dataset
ChIP Hep-G2 ENCSR310OZS.NR5A1.Hep-G2 110 bp overlap
NRF1 1 dataset
ChIP HeLa-S3 GSE108856.NRF1.HeLa-S3 312 bp overlap
Nfe2l2 2 datasets
Motif DE_12h DE_12h-Nfe2l2_MA0150.3 11 bp overlap
Motif DE_24h DE_24h-Nfe2l2_MA0150.3 11 bp overlap
OSR2 1 dataset
ChIP HEK293 GSE76494.OSR2.HEK293 135 bp overlap
PHIP 2 datasets
ChIP HCT-116_MLL1-KO__ab833 GSE101646.PHIP.HCT-116_MLL1-KO__ab833 324 bp overlap
ChIP HCT-116_ab834 GSE101646.PHIP.HCT-116_ab834 324 bp overlap
PPARG 1 dataset
ChIP Hep-G2 ENCSR130VQL.PPARG.Hep-G2 324 bp overlap
RAD21 3 datasets
ChIP SLK_CTCF-KD GSE138105.RAD21.SLK_CTCF-KD 307 bp overlap
ChIP liver ENCFF485PAC 230 bp overlap
ChIP liver ENCFF522JHE 93 bp overlap
REST 3 datasets
Motif DE_12h DE_12h-REST_MA0138.3 20 bp overlap
Motif DE_24h DE_24h-REST_MA0138.3 20 bp overlap
ChIP WA01 ENCSR000BHM.REST.WA01 120 bp overlap
RUNX1 3 datasets
ChIP HL-60 GSE107553.RUNX1.HL-60 54 bp overlap
ChIP SKH1_RUNX1-EVI1_KD GSE87283.RUNX1.SKH1_RUNX1-EVI1_KD 57 bp overlap
ChIP epididymis_HEE GSE109061.RUNX1.epididymis_HEE 136 bp overlap
RXRA 3 datasets
ChIP liver ENCFF077DAP 253 bp overlap
ChIP liver ENCFF077DAP 324 bp overlap
ChIP liver ENCFF807CIA 276 bp overlap
SAP130 1 dataset
ChIP HepG2 ENCFF892EHZ 310 bp overlap
SIN3A 1 dataset
ChIP HCT116 ENCFF203YBB 246 bp overlap
SMAD2 1 dataset
Motif DE_24h DE_24h-SMAD2_MA1964.2 6 bp overlap
SMARCA2 1 dataset
ChIP NPC_R1159Q_AM GSE122631.SMARCA2.NPC_R1159Q_AM 324 bp overlap
SMARCA4 15 datasets
ChIP 501-mel_SHCTR GSE61965.SMARCA4.501-mel_SHCTR 93 bp overlap
ChIP A-549_AG15677 GSE132290.SMARCA4.A-549_AG15677 71 bp overlap
ChIP A-549_AG15678 GSE132290.SMARCA4.A-549_AG15678 74 bp overlap
ChIP A-549_AG15685 GSE132290.SMARCA4.A-549_AG15685 193 bp overlap
ChIP A-549_AG15720 GSE132290.SMARCA4.A-549_AG15720 129 bp overlap
ChIP A-549_AG15721 GSE132290.SMARCA4.A-549_AG15721 102 bp overlap
ChIP A-549_AG15722 GSE132290.SMARCA4.A-549_AG15722 101 bp overlap
ChIP A-549_AG15723 GSE132290.SMARCA4.A-549_AG15723 86 bp overlap
ChIP A-549_AG15724 GSE132290.SMARCA4.A-549_AG15724 66 bp overlap
ChIP TTC-1240 GSE124903.SMARCA4.TTC-1240 324 bp overlap
ChIP TTC-1240_R377H GSE124903.SMARCA4.TTC-1240_R377H 324 bp overlap
ChIP TTC-1240_SMARCB1-FL GSE124903.SMARCA4.TTC-1240_SMARCB1-FL 324 bp overlap
ChIP TTC-1240_delC GSE124903.SMARCA4.TTC-1240_delC 173 bp overlap
ChIP TTC-1240_empty GSE124903.SMARCA4.TTC-1240_empty 324 bp overlap
ChIP endometrial-epithelial-cells_KO GSE106660.SMARCA4.endometrial-epithelial-cells_KO 229 bp overlap
SMARCC1 2 datasets
ChIP TTC-1240_R377H GSE124903.SMARCC1.TTC-1240_R377H 239 bp overlap
ChIP TTC-1240_SMARCB1-FL GSE124903.SMARCC1.TTC-1240_SMARCB1-FL 227 bp overlap
SMC3 1 dataset
ChIP peripheral-blood-neutrophil_Ecoli-1 GSE126755.SMC3.peripheral-blood-neutrophil_Ecoli-1 199 bp overlap
SNAI2 1 dataset
ChIP PC-9_2DF GSE131687.SNAI2.PC-9_2DF 324 bp overlap
SREBP2 3 datasets
ChIP monocyte GSE129202.SREBP2.monocyte 117 bp overlap
ChIP monocyte GSE129202.SREBP2.monocyte 70 bp overlap
ChIP monocyte_TNF GSE129202.SREBP2.monocyte_TNF 324 bp overlap
STAG1 1 dataset
ChIP Hep-G2 ENCSR167MTG.STAG1.Hep-G2 166 bp overlap
TAL1 1 dataset
ChIP PRIMA2 GSE33850.TAL1.PRIMA2 95 bp overlap
TCF3 2 datasets
ChIP Kasumi-1 GSE43834.TCF3.Kasumi-1 164 bp overlap
ChIP Kasumi-1 GSE43834.TCF3.Kasumi-1 106 bp overlap
TEAD1 1 dataset
ChIP MSTO GSE68170.TEAD1.MSTO 180 bp overlap
TEAD3 1 dataset
ChIP HepG2 ENCFF054UUL 226 bp overlap
TEAD4 4 datasets
ChIP BJ_fibroblast_CD13-neg GSE114367.TEAD4.BJ_fibroblast_CD13-neg 173 bp overlap
ChIP Ishikawa ENCFF772OTG 219 bp overlap
ChIP Ishikawa ENCSR000BSW.TEAD4.Ishikawa 134 bp overlap
ChIP PC-9_1DF_DMSO GSE131687.TEAD4.PC-9_1DF_DMSO 245 bp overlap
TFE3 1 dataset
ChIP Hep-G2 ENCSR589SNT.TFE3.Hep-G2 123 bp overlap
UBTF 1 dataset
ChIP HepG2 ENCFF424RNN 78 bp overlap
USF1 1 dataset
ChIP Ishikawa ENCSR000BSX.USF1.Ishikawa 96 bp overlap
VDR 2 datasets
ChIP LX2_CALCIPOTRIOL_TGFB1 GSE38103.VDR.LX2_CALCIPOTRIOL_TGFB1 130 bp overlap
ChIP kidney-cortex GSE129585.VDR.kidney-cortex 324 bp overlap
ZBTB24 1 dataset
ChIP HCT-116 GSE111683.ZBTB24.HCT-116 189 bp overlap
ZEB1 1 dataset
ChIP Hep-G2 ENCSR000BVN.ZEB1.Hep-G2 124 bp overlap
ZEB2 1 dataset
ChIP K-562 ENCSR004GKA.ZEB2.K-562 139 bp overlap
ZFP36 1 dataset
ChIP K-562 ENCSR776CYN.ZFP36.K-562 127 bp overlap
ZFP64 1 dataset
ChIP Hep-G2 ENCSR487CPI.ZFP64.Hep-G2 115 bp overlap
ZGPAT 1 dataset
ChIP Hep-G2 ENCSR704IGU.ZGPAT.Hep-G2 324 bp overlap
ZMYM3 2 datasets
ChIP Hep-G2_AC_JH39-2-2B9 GSE97661.ZMYM3.Hep-G2_AC_JH39-2-2B9 54 bp overlap
ChIP Hep-G2_Ab_JH39-2-2F10 GSE97661.ZMYM3.Hep-G2_Ab_JH39-2-2F10 72 bp overlap
ZNF12 1 dataset
ChIP Hep-G2 ENCSR711KBM.ZNF12.Hep-G2 115 bp overlap
ZNF664 2 datasets
ChIP HEK293 ENCFF343XSW 172 bp overlap
ChIP HEK293 ENCSR714LZQ.ZNF664.HEK293 93 bp overlap