chr12 : 52,947,826 52,948,366
540 bp 114 TFs 2 linked genes
This 540 bp open chromatin element is linked to KRT18 and KRT8 and is bound by 114 transcription factors.
Linked Genes
2 genes
Distance
Gene Expression Dist. to TSS Distance Link type
KRT18 750 bp At TSS Proximity
KRT8 1.5 kb Proximal Proximity
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr12:52,942,826 – 52,953,366
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
114 transcription factors
Source
Cell type
AFF4 1 dataset
ChIP MCF-7 GSE144036.AFF4.MCF-7 141 bp overlap
AR 1 dataset
ChIP LNCaP_FOXA1 ERP003503.AR.LNCaP_FOXA1 143 bp overlap
ARID1A 1 dataset
ChIP MCF-7_4-hydroxytamoxifen GSE123284.ARID1A.MCF-7_4-hydroxytamoxifen 352 bp overlap
ASH2L 1 dataset
ChIP WA01 ENCSR850KIP.ASH2L.WA01 343 bp overlap
BCL6 1 dataset
ChIP CD4 GSE59933.BCL6.CD4 143 bp overlap
BCOR 3 datasets
ChIP WA01 GSE104690.BCOR.WA01 333 bp overlap
ChIP WA01_RNF2-R GSE104690.BCOR.WA01_RNF2-R 219 bp overlap
ChIP WA01_RNF2-R_Doxycyclin GSE104690.BCOR.WA01_RNF2-R_Doxycyclin 540 bp overlap
BRD1 2 datasets
ChIP HUES-64 GSE104059.BRD1.HUES-64 349 bp overlap
ChIP RKO GSE47190.BRD1.RKO 156 bp overlap
BRD4 7 datasets
ChIP COLO-205 GSE73319.BRD4.COLO-205 540 bp overlap
ChIP HCT-15 GSE73319.BRD4.HCT-15 540 bp overlap
ChIP MCF-7_ARID1A-KO_4-OHT GSE123284.BRD4.MCF-7_ARID1A-KO_4-OHT 540 bp overlap
ChIP MCF-7_parental GSE123284.BRD4.MCF-7_parental 540 bp overlap
ChIP MCF-7_parental_4-hydroxytamoxifen GSE123284.BRD4.MCF-7_parental_4-hydroxytamoxifen 540 bp overlap
ChIP PC-3_GDC-R_SAHA GSE137207.BRD4.PC-3_GDC-R_SAHA 280 bp overlap
ChIP SW480 GSE73319.BRD4.SW480 540 bp overlap
CAMTA2 1 dataset
ChIP HepG2 ENCFF305ZLM 241 bp overlap
CBFB 1 dataset
ChIP Hep-G2 ENCSR479RZV.CBFB.Hep-G2 92 bp overlap
CHD1 3 datasets
ChIP HeLa-S3 ENCSR392EAA.CHD1.HeLa-S3 147 bp overlap
ChIP MCF-7 ENCSR360JOC.CHD1.MCF-7 540 bp overlap
ChIP WA01 ENCSR000AQK.CHD1.WA01 507 bp overlap
CHD4 1 dataset
ChIP A-549 ENCSR550SCU.CHD4.A-549 416 bp overlap
CHD7 1 dataset
ChIP WA01 ENCSR000AVA.CHD7.WA01 193 bp overlap
CREB1 1 dataset
ChIP WA01 ENCSR000BSN.CREB1.WA01 120 bp overlap
CREBBP 2 datasets
ChIP PC-3 GSE147455.CREBBP.PC-3 116 bp overlap
ChIP PC-3_GDC-resist GSE147455.CREBBP.PC-3_GDC-resist 255 bp overlap
CRY1 1 dataset
ChIP U2OS GSE44236.CRY1.U2OS 251 bp overlap
DDX5 1 dataset
ChIP NTERA2 GSE58641.DDX5.NTERA2 143 bp overlap
E2F1 1 dataset
ChIP MDA-MB-231 GSE95303.E2F1.MDA-MB-231 231 bp overlap
E2F4 1 dataset
ChIP K-562 ENCSR000EWL.E2F4.K-562 128 bp overlap
EGR1 1 dataset
ChIP Hep-G2 ENCSR026GSW.EGR1.Hep-G2 155 bp overlap
EMX1 1 dataset
ChIP WTC11 ENCFF692RZJ 540 bp overlap
EP300 1 dataset
ChIP HeLa-S3 ENCSR680OFU.EP300.HeLa-S3 189 bp overlap
ESR1 7 datasets
ChIP MCF-7_DHT GSE99626.ESR1.MCF-7_DHT 428 bp overlap
ChIP T-47D GSE148277.ESR1.T-47D 232 bp overlap
ChIP T-47D_DMSO GSE148277.ESR1.T-47D_DMSO 344 bp overlap
ChIP T-47D_Y537S GSE148277.ESR1.T-47D_Y537S 412 bp overlap
ChIP T-47D_Y537S_E2 GSE148277.ESR1.T-47D_Y537S_E2 540 bp overlap
ChIP breast_tumor_Male_20 GSE104399.ESR1.breast_tumor_Male_20 172 bp overlap
ChIP breast_tumor_Male_21 GSE104399.ESR1.breast_tumor_Male_21 330 bp overlap
ESRRA 1 dataset
ChIP SK-BR-3_HRG GSE81651.ESRRA.SK-BR-3_HRG 282 bp overlap
ETS1 3 datasets
ChIP HEY-A8 GSE101832.ETS1.HEY-A8 422 bp overlap
ChIP HUVEC-C_VEGF_12H GSE41166.ETS1.HUVEC-C_VEGF_12H 190 bp overlap
ChIP cardiomyocyte_D9 GSE129986.ETS1.cardiomyocyte_D9 165 bp overlap
EZH2 1 dataset
ChIP THP-1 GSE135024.EZH2.THP-1 192 bp overlap
FOS 1 dataset
ChIP HCT-116_A7A GSE152144.FOS.HCT-116_A7A 420 bp overlap
FOSB 1 dataset
ChIP A549 ENCFF029EEU 341 bp overlap
FOXA1 1 dataset
ChIP T-47D_shRNF2_8h GSE137579.FOXA1.T-47D_shRNF2_8h 106 bp overlap
GATA6 3 datasets
ChIP DE_D1 S09-DE-d1-GATA6-exp1 270 bp overlap
ChIP DE_D1 S41-DE-d1-GATA6-exp2 310 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 334 bp overlap
GTF3C2 1 dataset
ChIP H9 GSE94418.GTF3C2.H9 144 bp overlap
HDAC2 1 dataset
ChIP PC-3 GSE147455.HDAC2.PC-3 329 bp overlap
HDGF 1 dataset
ChIP K-562 ENCSR563YDA.HDGF.K-562 355 bp overlap
HOXB13 2 datasets
ChIP A-549 ENCSR967ZMR.HOXB13.A-549 255 bp overlap
ChIP A549 ENCFF870NOA 351 bp overlap
JUN 9 datasets
ChIP DE_D1 S08-DE-d1-JUN-exp1 349 bp overlap
ChIP DE_D1 S13-DE-d1-JUN-exp1 284 bp overlap
ChIP DE_D1 S40-DE-d1-JUN-exp2 540 bp overlap
ChIP DE_D2 S02-DE-d2-JUN-exp1 261 bp overlap
ChIP ESC S34-ESC-d0-JUN-exp2 493 bp overlap
ChIP HUES-8 GSE109524.JUN.HUES-8 472 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.JUN.HUES-8_DE_ctrl 480 bp overlap
ChIP MDA-BoM-1833 GSE112444.JUN.MDA-BoM-1833 305 bp overlap
ChIP MDA-MB-231 GSE112444.JUN.MDA-MB-231 262 bp overlap
KAT7 1 dataset
ChIP K562 ENCFF175ZTN 476 bp overlap
KDM1A 1 dataset
ChIP NCI-H526_DMSO GSE66297.KDM1A.NCI-H526_DMSO 141 bp overlap
KMT2A 1 dataset
ChIP Hep-G2 ENCSR692GFR.KMT2A.Hep-G2 231 bp overlap
MBD3 1 dataset
ChIP MCF-7 GSE44737.MBD3.MCF-7 516 bp overlap
MED1 4 datasets
ChIP HUVEC-C_siCTL GSE124892.MED1.HUVEC-C_siCTL 224 bp overlap
ChIP HUVEC-C_siCTL GSE124892.MED1.HUVEC-C_siCTL 227 bp overlap
ChIP HUVEC-C_siERG GSE124892.MED1.HUVEC-C_siERG 314 bp overlap
ChIP cardiomyocyte_5 GSE85628.MED1.cardiomyocyte_5 202 bp overlap
MSX2 2 datasets
ChIP MCF-7 ENCFF179YRV 297 bp overlap
ChIP MCF-7 ENCFF179YRV 244 bp overlap
MYBL2 1 dataset
ChIP A-673 GSE119971.MYBL2.A-673 129 bp overlap
NANOG 5 datasets
ChIP GM23338 ENCSR061DGF.NANOG.GM23338 126 bp overlap
ChIP HUES-8 GSE109524.NANOG.HUES-8 427 bp overlap
ChIP WA01 ERP004238.NANOG.WA01 247 bp overlap
ChIP WA01_3IL ERP004238.NANOG.WA01_3IL 435 bp overlap
ChIP hESC GSE18292.NANOG.hESC 93 bp overlap
NCAPH2 2 datasets
ChIP RMG-I_ARID1A-KO GSE120058.NCAPH2.RMG-I_ARID1A-KO 190 bp overlap
ChIP RMG-I_ARID1A-KO GSE120058.NCAPH2.RMG-I_ARID1A-KO 158 bp overlap
NKX2-1 1 dataset
ChIP NCI-H3122 GSE39998.NKX2-1.NCI-H3122 200 bp overlap
NR3C1 2 datasets
ChIP K562 ENCFF877YZJ 91 bp overlap
ChIP T47D-A1-2_Dex GSE112491.NR3C1.T47D-A1-2_Dex 87 bp overlap
OTX1 1 dataset
ChIP MCF-7 ENCFF645GYL 317 bp overlap
PAF1 1 dataset
ChIP HCT-116_ab20662 GSE97527.PAF1.HCT-116_ab20662 264 bp overlap
PCBP1 2 datasets
ChIP Hep-G2 ENCSR872EVQ.PCBP1.Hep-G2 189 bp overlap
ChIP Hep-G2 GSE120104.PCBP1.Hep-G2 189 bp overlap
PCBP2 2 datasets
ChIP Hep-G2 ENCSR945NSF.PCBP2.Hep-G2 234 bp overlap
ChIP Hep-G2 GSE120104.PCBP2.Hep-G2 207 bp overlap
PGR 4 datasets
ChIP MCF-7_PROG GSE68355.PGR.MCF-7_PROG 224 bp overlap
ChIP MCF-7_R5020 GSE68355.PGR.MCF-7_R5020 212 bp overlap
ChIP T-47D GSE126859.PGR.T-47D 168 bp overlap
ChIP T-47D_R5020-A1 GSE126859.PGR.T-47D_R5020-A1 194 bp overlap
PHIP 4 datasets
ChIP HCT-116_MLL1-KO__ab833 GSE101646.PHIP.HCT-116_MLL1-KO__ab833 540 bp overlap
ChIP HCT-116_MLL4-SET-KO_ab833 GSE101646.PHIP.HCT-116_MLL4-SET-KO_ab833 249 bp overlap
ChIP HCT-116_ab833 GSE101646.PHIP.HCT-116_ab833 540 bp overlap
ChIP HCT-116_ab834 GSE101646.PHIP.HCT-116_ab834 540 bp overlap
POLR2A 8 datasets
ChIP GM23338 ENCFF450WCS 456 bp overlap
ChIP H1 ENCFF566JSR 310 bp overlap
ChIP H1 ENCFF833NJP 316 bp overlap
ChIP HepG2 ENCFF736SLT 322 bp overlap
ChIP MCF-7 ENCFF309IKZ 255 bp overlap
ChIP MCF-7 ENCFF411WCU 280 bp overlap
ChIP Panc1 ENCFF290KAB 391 bp overlap
ChIP Panc1 ENCFF290KAB 187 bp overlap
POU2F1 1 dataset
ChIP T-47D GSE148277.POU2F1.T-47D 341 bp overlap
POU2F3 1 dataset
ChIP NCI-H1048 GSE115123.POU2F3.NCI-H1048 186 bp overlap
POU5F1 5 datasets
ChIP DE_D1 DED1-OCT4_Batch_II 540 bp overlap
ChIP GM23338 ENCFF333SNB 311 bp overlap
ChIP HUES-8 GSE109524.POU5F1.HUES-8 430 bp overlap
ChIP hESC ENCSR264RJX.POU5F1.hESC 158 bp overlap
ChIP hiPSC GSE56567.POU5F1.hiPSC 150 bp overlap
POU5F1_M 1 dataset
ChIP DE_D1 DED1-OCT4-M_Batch_II 504 bp overlap
PRPF4 1 dataset
ChIP Hep-G2 GSE120104.PRPF4.Hep-G2 152 bp overlap
RAD21 1 dataset
ChIP HCT-116 GSE131606.RAD21.HCT-116 398 bp overlap
RARA 1 dataset
ChIP U-937_ATRA-treated_RAR GSE98006.RARA.U-937_ATRA-treated_RAR 110 bp overlap
RBFOX2 2 datasets
ChIP K-562 GSE120104.RBFOX2.K-562 430 bp overlap
ChIP K-562 ENCSR822LBD.RBFOX2.K-562 384 bp overlap
RBM39 3 datasets
ChIP Hep-G2 ENCSR339JTP.RBM39.Hep-G2 164 bp overlap
ChIP Hep-G2 GSE120104.RBM39.Hep-G2 158 bp overlap
ChIP K-562 ENCSR764OXF.RBM39.K-562 53 bp overlap
SIN3A 2 datasets
ChIP A-549 ENCSR513XQX.SIN3A.A-549 323 bp overlap
ChIP WA01 ENCSR000EBO.SIN3A.WA01 63 bp overlap
SMAD2-3 2 datasets
ChIP HUES-8_DE_JNKi GSE109524.SMAD2-3.HUES-8_DE_JNKi 284 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.SMAD2-3.HUES-8_DE_ctrl 455 bp overlap
SMAD3 2 datasets
ChIP HCC1954_TGFb GSE104760.SMAD3.HCC1954_TGFb 94 bp overlap
ChIP NCI-H441_SIITTF1 GSE51509.SMAD3.NCI-H441_SIITTF1 168 bp overlap
SMAD4 1 dataset
ChIP K562 ENCFF316DFN 371 bp overlap
SMAD5 2 datasets
ChIP K-562 ENCSR000FCD.SMAD5.K-562 122 bp overlap
ChIP K-562 ENCSR000FCD.SMAD5.K-562 132 bp overlap
SMAD7 2 datasets
ChIP HepG2 ENCFF850FXR 424 bp overlap
ChIP HepG2 ENCFF850FXR 540 bp overlap
SMARCA4 5 datasets
ChIP MCF-7 GSE140185.SMARCA4.MCF-7 274 bp overlap
ChIP MCF-7_ARID1A-KO_clone14 GSE123284.SMARCA4.MCF-7_ARID1A-KO_clone14 298 bp overlap
ChIP endometrial-epithelial-cells_KO GSE106660.SMARCA4.endometrial-epithelial-cells_KO 540 bp overlap
ChIP hiPSC GSE124903.SMARCA4.hiPSC 225 bp overlap
ChIP hiPSC_WT GSE124903.SMARCA4.hiPSC_WT 392 bp overlap
SMARCB1 3 datasets
ChIP HeLa-S3 ENCFF733PLR 367 bp overlap
ChIP hiPSC GSE124903.SMARCB1.hiPSC 190 bp overlap
ChIP hiPSC_WT GSE124903.SMARCB1.hiPSC_WT 169 bp overlap
SMARCC1 5 datasets
ChIP DE_D1 S10-DE-d1-BAF155-exp1 257 bp overlap
ChIP DE_D1 S15-DE-d1-BAF155-exp1 442 bp overlap
ChIP ESC S25-ESC-d0-BAF155-exp1 534 bp overlap
ChIP hiPSC GSE124903.SMARCC1.hiPSC 249 bp overlap
ChIP hiPSC_WT GSE124903.SMARCC1.hiPSC_WT 181 bp overlap
SMC1 1 dataset
ChIP DKO GSE131606.SMC1.DKO 264 bp overlap
SMC3 1 dataset
ChIP A549 ENCFF747SCJ 231 bp overlap
SOX2 1 dataset
ChIP hiPSC GSE56567.SOX2.hiPSC 193 bp overlap
SP1 1 dataset
ChIP Hep-G2 ENCSR460YAM.SP1.Hep-G2 113 bp overlap
SP4 1 dataset
ChIP H1 ENCFF473YOB 157 bp overlap
SRSF3 2 datasets
ChIP K-562 GSE120104.SRSF3.K-562 333 bp overlap
ChIP K-562 ENCSR268QIQ.SRSF3.K-562 333 bp overlap
SSRP1 2 datasets
ChIP HepG2 ENCFF540BLL 440 bp overlap
ChIP HepG2 ENCFF540BLL 432 bp overlap
STAT6 1 dataset
ChIP HepG2 ENCFF370LZV 216 bp overlap
SUPT5H 1 dataset
ChIP DLD-1_NELFCD-AID GSE144786.SUPT5H.DLD-1_NELFCD-AID 540 bp overlap
SUPT6H 1 dataset
ChIP HCT-116 GSE130509.SUPT6H.HCT-116 345 bp overlap
T 2 datasets
ChIP H9_ENDODERM GSE60606.T.H9_ENDODERM 185 bp overlap
ChIP H9_MESODERM GSE60606.T.H9_MESODERM 201 bp overlap
TAF1 1 dataset
ChIP H1 ENCFF478SZO 140 bp overlap
TARDBP 3 datasets
ChIP K-562 GSE120104.TARDBP.K-562 258 bp overlap
ChIP K-562 ENCSR429XTR.TARDBP.K-562 195 bp overlap
ChIP K562 ENCFF021QCU 345 bp overlap
TBP 5 datasets
ChIP WA01 ENCSR000ECB.TBP.WA01 149 bp overlap
ChIP hESC GSE122298.TBP.hESC 363 bp overlap
ChIP hESC_10h GSE122298.TBP.hESC_10h 305 bp overlap
ChIP hESC_2h GSE122298.TBP.hESC_2h 313 bp overlap
ChIP hESC_8h GSE122298.TBP.hESC_8h 110 bp overlap
TBX5 5 datasets
ChIP G296S GSE85628.TBX5.G296S 171 bp overlap
ChIP G296S_2 GSE85628.TBX5.G296S_2 171 bp overlap
ChIP G296S_4 GSE85628.TBX5.G296S_4 406 bp overlap
ChIP cardiomyocyte GSE85628.TBX5.cardiomyocyte 316 bp overlap
ChIP cardiomyocyte_1 GSE85628.TBX5.cardiomyocyte_1 316 bp overlap
TCF12 1 dataset
ChIP A-549 ENCSR000BQQ.TCF12.A-549 221 bp overlap
TEAD3 1 dataset
ChIP HepG2 ENCFF054UUL 214 bp overlap
TFAP2C 2 datasets
ChIP UCLA1-hESCs_TFAP2C--_1d_TFAP2C_Induc GSE101074.TFAP2C.UCLA1-hESCs_TFAP2C--_1d_TFAP2C_Induc 499 bp overlap
ChIP UCLA1-hESCs_TFAP2C--_2d_TFAP2C_Induc GSE101074.TFAP2C.UCLA1-hESCs_TFAP2C--_2d_TFAP2C_Induc 254 bp overlap
TSC22D1 1 dataset
ChIP MCF-7 ENCFF552PAC 251 bp overlap
TSC22D2 1 dataset
ChIP MCF-7 ENCFF588BUN 297 bp overlap
ZBTB43 2 datasets
ChIP WTC11 ENCFF058JUB 485 bp overlap
ChIP WTC11 ENCFF058JUB 293 bp overlap
ZC3H4 2 datasets
ChIP K562 ENCFF343JOP 387 bp overlap
ChIP K562 ENCFF343JOP 179 bp overlap
ZFP3 1 dataset
ChIP SK-N-SH_all-trans-retinoic-acid_6-uM_48-hour ENCSR581GUY.ZFP3.SK-N-SH_all-trans-retinoic-acid_6-uM_48-hour 255 bp overlap
ZFX 1 dataset
ChIP MCF-7 GSE102616.ZFX.MCF-7 295 bp overlap
ZKSCAN1 2 datasets
ChIP Hep-G2 ENCSR157CAU.ZKSCAN1.Hep-G2 298 bp overlap
ChIP K-562 ENCSR882ERE.ZKSCAN1.K-562 208 bp overlap
ZNF257 1 dataset
ChIP HEK293T GSE78099.ZNF257.HEK293T 208 bp overlap
ZNF302 1 dataset
ChIP A549 ENCFF333REI 271 bp overlap
ZNF362 2 datasets
ChIP HEK293 ENCSR715QNO.ZNF362.HEK293 129 bp overlap
ChIP HepG2 ENCFF256AZN 149 bp overlap
ZNF776 1 dataset
ChIP HepG2 ENCFF009LSZ 282 bp overlap