chr10 : 32,154,889 32,155,312
423 bp 92 TFs 0 linked genes
This 423 bp open chromatin element has no linked target genes and is bound by 92 transcription factors.
Linked Genes
No linked genes
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr10:32,149,889 – 32,160,312
This element Other elements ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
92 transcription factors
Source
Cell type
AR 1 dataset
ChIP LNCaP GSE80256.AR.LNCaP 318 bp overlap
ARID1A 4 datasets
ChIP 12Z GSE129781.ARID1A.12Z 149 bp overlap
ChIP MCF-7 GSE123284.ARID1A.MCF-7 296 bp overlap
ChIP MCF-7_DCDT GSE123284.ARID1A.MCF-7_DCDT 314 bp overlap
ChIP MCF-7_JQ1 GSE123284.ARID1A.MCF-7_JQ1 296 bp overlap
BRD2 1 dataset
ChIP HCC1806_NTsiRNA GSE116879.BRD2.HCC1806_NTsiRNA 199 bp overlap
BRD4 5 datasets
ChIP HEK293_sgNT GSE129407.BRD4.HEK293_sgNT 307 bp overlap
ChIP LNCaP-C4-2_F133V GSE88871.BRD4.LNCaP-C4-2_F133V 98 bp overlap
ChIP MCF-7_parental GSE123284.BRD4.MCF-7_parental 423 bp overlap
ChIP SUM159PT_100nMtrametinib_1h GSE87418.BRD4.SUM159PT_100nMtrametinib_1h 111 bp overlap
ChIP keratinocyte GSE140992.BRD4.keratinocyte 423 bp overlap
CARM1 1 dataset
ChIP MCF-7_E2 GSE124448.CARM1.MCF-7_E2 381 bp overlap
CBX1 1 dataset
ChIP Hep-G2 ENCSR819WZE.CBX1.Hep-G2 156 bp overlap
CEBPA 2 datasets
ChIP Hep-G2 ERP000209.CEBPA.Hep-G2 137 bp overlap
ChIP HepG2 ENCFF175DFS 291 bp overlap
CEBPB 1 dataset
ChIP IMR-90 ENCFF468UGY 251 bp overlap
CREB1 1 dataset
ChIP LNCaP-abl_SHFOXA1 GSE63034.CREB1.LNCaP-abl_SHFOXA1 64 bp overlap
CREBBP 4 datasets
ChIP PC-3 GSE147455.CREBBP.PC-3 292 bp overlap
ChIP PC-3_GDC-resist GSE147455.CREBBP.PC-3_GDC-resist 422 bp overlap
ChIP keratinocyte GSE140991.CREBBP.keratinocyte 316 bp overlap
ChIP keratinocyte_KLF3i GSE140991.CREBBP.keratinocyte_KLF3i 252 bp overlap
CTBP1 1 dataset
ChIP MCF-7 ENCSR636EYA.CTBP1.MCF-7 145 bp overlap
CTCF 30 datasets
ChIP AG10803 ENCFF549AQK 257 bp overlap
ChIP aorta_thoracic ENCSR668BTN.CTCF.aorta_thoracic 368 bp overlap
ChIP ascending-aorta ENCSR846JKO.CTCF.ascending-aorta 279 bp overlap
ChIP astrocyte_spinal_cord ENCSR000DSU.CTCF.astrocyte_spinal_cord 145 bp overlap
ChIP bonchial_epithelial ENCSR000DXI.CTCF.bonchial_epithelial 113 bp overlap
ChIP brain microvascular endothelial cell ENCFF526VHE 281 bp overlap
ChIP cardiac fibroblast ENCFF326EDY 265 bp overlap
ChIP cardiac_muscle ENCSR000DTI.CTCF.cardiac_muscle 222 bp overlap
ChIP choroid plexus epithelial cell ENCFF407YNR 277 bp overlap
ChIP choroid-plexus_epithelial ENCSR000DTL.CTCF.choroid-plexus_epithelial 180 bp overlap
ChIP endothelial_brain-microvascular ENCSR000DTA.CTCF.endothelial_brain-microvascular 168 bp overlap
ChIP epithelial cell of prostate ENCFF086GTI 413 bp overlap
ChIP epithelial_mammary ENCSR000ALV.CTCF.epithelial_mammary 169 bp overlap
ChIP esophagus_squamous-epithelium ENCSR450FRI.CTCF.esophagus_squamous-epithelium 294 bp overlap
ChIP fibroblast_AORTIC_ADVENTITIA ENCSR000DPY.CTCF.fibroblast_AORTIC_ADVENTITIA 101 bp overlap
ChIP fibroblast_CARDIAC ENCSR000DTF.CTCF.fibroblast_CARDIAC 224 bp overlap
ChIP fibroblast_PULMONARY_ARTERY ENCSR000DUX.CTCF.fibroblast_PULMONARY_ARTERY 148 bp overlap
ChIP fibroblast_SKIN_LEG ENCSR000DPG.CTCF.fibroblast_SKIN_LEG 150 bp overlap
ChIP keratinocyte ENCSR000ALJ.CTCF.keratinocyte 175 bp overlap
ChIP keratinocyte ENCSR000DWX.CTCF.keratinocyte 172 bp overlap
ChIP keratinocyte GSE154221.CTCF.keratinocyte 248 bp overlap
ChIP keratinocyte ENCSR000DNC.CTCF.keratinocyte 133 bp overlap
ChIP mammary epithelial cell ENCFF164SPU 265 bp overlap
ChIP mammary epithelial cell ENCFF873ERE 271 bp overlap
ChIP osteoblast ENCFF491ZJZ 369 bp overlap
ChIP osteoblast ENCSR000APF.CTCF.osteoblast 377 bp overlap
ChIP primary-epidermal-keratinocyte_diff_d0 GSE84657.CTCF.primary-epidermal-keratinocyte_diff_d0 158 bp overlap
ChIP primary-epidermal-keratinocyte_diff_d6 GSE84657.CTCF.primary-epidermal-keratinocyte_diff_d6 148 bp overlap
ChIP prostate_epithelial ENCSR196HOM.CTCF.prostate_epithelial 302 bp overlap
ChIP thoracic aorta ENCFF012WJQ 380 bp overlap
Cebpa 1 dataset
ChIP BLaER1 ENCFF335XTP 251 bp overlap
DPF2 2 datasets
ChIP MCF-7 ENCFF712EXQ 97 bp overlap
ChIP MCF-7 ENCSR234VCE.DPF2.MCF-7 156 bp overlap
DRAP1 1 dataset
ChIP Hep-G2 ENCSR765MKZ.DRAP1.Hep-G2 321 bp overlap
EGR1 1 dataset
ChIP fibroblast_DOX-24h GSE134924.EGR1.fibroblast_DOX-24h 202 bp overlap
ELF3 1 dataset
ChIP PDAC_SHCTR GSE64557.ELF3.PDAC_SHCTR 423 bp overlap
EOMES 1 dataset
ChIP hESC GSE26097.EOMES.hESC 205 bp overlap
EP300 2 datasets
ChIP Hep-G2 ENCSR000BLW.EP300.Hep-G2 206 bp overlap
ChIP PC-3 GSE147455.EP300.PC-3 310 bp overlap
ERG 3 datasets
ChIP aortic-endothelial-cell_D14 GSE139377.ERG.aortic-endothelial-cell_D14 380 bp overlap
ChIP aortic-endothelial-cell_D24 GSE139377.ERG.aortic-endothelial-cell_D24 171 bp overlap
ChIP aortic-endothelial-cell_D49 GSE139377.ERG.aortic-endothelial-cell_D49 71 bp overlap
ETV4 1 dataset
ChIP Hep-G2 ENCSR714YZG.ETV4.Hep-G2 155 bp overlap
FOS 3 datasets
ChIP CFPAC-1 GSE119930.FOS.CFPAC-1 420 bp overlap
ChIP IMR-90 ENCSR124AIG.FOS.IMR-90 159 bp overlap
ChIP MCF-7 ENCSR569XNP.FOS.MCF-7 177 bp overlap
FOSL2 1 dataset
ChIP MCF-7 ENCSR000BUI.FOSL2.MCF-7 134 bp overlap
FOXA1 6 datasets
ChIP HepG2 ENCFF207NVJ 205 bp overlap
ChIP HepG2 ENCFF740VZW 202 bp overlap
ChIP LNCaP_SHFOXA1_R1881 GSE37345.FOXA1.LNCaP_SHFOXA1_R1881 97 bp overlap
ChIP MCF-7-TAMR-1_4OH-Tam GSE75201.FOXA1.MCF-7-TAMR-1_4OH-Tam 75 bp overlap
ChIP MCF-7_vehicle_ab2 GSE112969.FOXA1.MCF-7_vehicle_ab2 115 bp overlap
ChIP primary-breast-cancer_B1_DSG GSE114737.FOXA1.primary-breast-cancer_B1_DSG 250 bp overlap
FOXA2 4 datasets
ChIP CFPAC-1_EV GSE119930.FOXA2.CFPAC-1_EV 112 bp overlap
ChIP CFPAC-1_HNF1B-KO GSE119930.FOXA2.CFPAC-1_HNF1B-KO 73 bp overlap
ChIP DE DE-FOXA2-1 388 bp overlap
ChIP DE DE-FOXA2-2 344 bp overlap
FOXK1 1 dataset
ChIP Hep-G2 ENCSR603BJQ.FOXK1.Hep-G2 123 bp overlap
FOXL2 3 datasets
ChIP KGN_1512 GSE138496.FOXL2.KGN_1512 242 bp overlap
ChIP KGN_1513 GSE138496.FOXL2.KGN_1513 256 bp overlap
ChIP SVOG-3e_DOX-WT GSE126171.FOXL2.SVOG-3e_DOX-WT 194 bp overlap
FOXP4 1 dataset
ChIP Hep-G2 ENCSR232LLP.FOXP4.Hep-G2 360 bp overlap
GATA2 4 datasets
ChIP Hep-G2 ENCSR897LDT.GATA2.Hep-G2 318 bp overlap
ChIP HepG2 ENCFF905PYM 371 bp overlap
ChIP dermal-fibroblast_alone GSE51025.GATA2.dermal-fibroblast_alone 123 bp overlap
ChIP endothelial cell of umbilical vein ENCFF148NLK 243 bp overlap
GATA4 1 dataset
ChIP DE DE-GATA4-2 406 bp overlap
GATA6 2 datasets
ChIP DE DE-GATA6-2 377 bp overlap
ChIP foregut_KO GSE117136.GATA6.foregut_KO 244 bp overlap
GLIS3 1 dataset
ChIP H9_plus GSE109562.GLIS3.H9_plus 353 bp overlap
GRHL2 3 datasets
ChIP MCF-7 GSE109820.GRHL2.MCF-7 67 bp overlap
ChIP MCF-7-WS8_VEH GSE113092.GRHL2.MCF-7-WS8_VEH 135 bp overlap
ChIP MCF-7_E2_45min GSE109820.GRHL2.MCF-7_E2_45min 112 bp overlap
HDAC2 4 datasets
ChIP Hep-G2 ENCSR000BMC.HDAC2.Hep-G2 346 bp overlap
ChIP PC-3 GSE147455.HDAC2.PC-3 399 bp overlap
ChIP PC-3_GDC-resist GSE147455.HDAC2.PC-3_GDC-resist 195 bp overlap
ChIP PC-3_GDC-resist GSE147455.HDAC2.PC-3_GDC-resist 94 bp overlap
HNF1B 1 dataset
ChIP Hep-G2 ENCSR127XTZ.HNF1B.Hep-G2 282 bp overlap
HNF4A 2 datasets
ChIP Hep-G2 ENCSR469FBY.HNF4A.Hep-G2 247 bp overlap
ChIP HepG2 ENCFF146SSF 356 bp overlap
HOXA3 2 datasets
ChIP Hep-G2 ENCSR999JKC.HOXA3.Hep-G2 296 bp overlap
ChIP HepG2 ENCFF374TCI 423 bp overlap
ISL2 1 dataset
ChIP Hep-G2 ENCSR492IHH.ISL2.Hep-G2 170 bp overlap
JUN 5 datasets
ChIP HAEC_IL1b_4h GSE89970.JUN.HAEC_IL1b_4h 200 bp overlap
ChIP HAEC_TNFa_4h GSE89970.JUN.HAEC_TNFa_4h 304 bp overlap
ChIP MCF-7_DMSO GSE157385.JUN.MCF-7_DMSO 313 bp overlap
ChIP MCF-7_E2 GSE102410.JUN.MCF-7_E2 380 bp overlap
ChIP MCF-7_abemaciclib GSE157385.JUN.MCF-7_abemaciclib 328 bp overlap
JUNB 2 datasets
ChIP CFPAC-1 GSE119930.JUNB.CFPAC-1 423 bp overlap
ChIP HAEC GSE89970.JUNB.HAEC 216 bp overlap
JUND 1 dataset
ChIP HCT116 ENCFF748ZQX 87 bp overlap
KDM4C 1 dataset
ChIP KYSE-150 GSE53938.KDM4C.KYSE-150 199 bp overlap
KLF4 1 dataset
ChIP PDAC GSE64557.KLF4.PDAC 350 bp overlap
KMT2C 1 dataset
ChIP BIN-67_lentivirus-SMARCA4 GSE117734.KMT2C.BIN-67_lentivirus-SMARCA4 174 bp overlap
MAX 1 dataset
ChIP HepG2 ENCFF507HCX 423 bp overlap
MAZ 1 dataset
ChIP IMR-90 ENCSR000EFF.MAZ.IMR-90 109 bp overlap
MED1 12 datasets
ChIP HCT-116_thaps GSE121798.MED1.HCT-116_thaps 264 bp overlap
ChIP Hep-G2 ENCSR959XNY.MED1.Hep-G2 379 bp overlap
ChIP HepG2 ENCFF495TSS 381 bp overlap
ChIP U-87MG GSE36354.MED1.U-87MG 282 bp overlap
ChIP VCaP_Darolutamide GSE148358.MED1.VCaP_Darolutamide 206 bp overlap
ChIP hMSC-TERT4_4h GSE104537.MED1.hMSC-TERT4_4h 342 bp overlap
ChIP hMSC-TERT4_D14 GSE104537.MED1.hMSC-TERT4_D14 155 bp overlap
ChIP hMSC-TERT4_D3 GSE104537.MED1.hMSC-TERT4_D3 127 bp overlap
ChIP hMSC-TERT4_adipocyte-4H GSE113253.MED1.hMSC-TERT4_adipocyte-4H 250 bp overlap
ChIP hMSC-TERT4_osteoblast-D1 GSE113253.MED1.hMSC-TERT4_osteoblast-D1 63 bp overlap
ChIP hMSC-TERT4_osteoblast-D3 GSE113253.MED1.hMSC-TERT4_osteoblast-D3 203 bp overlap
ChIP hMSC-TERT4_osteoblast-D7 GSE113253.MED1.hMSC-TERT4_osteoblast-D7 313 bp overlap
MYB 1 dataset
ChIP DU528 GSE94000.MYB.DU528 221 bp overlap
MYOG 1 dataset
ChIP RH4_Entinostat-6H GSE116344.MYOG.RH4_Entinostat-6H 285 bp overlap
NCAPH2 2 datasets
ChIP IMR-90_Triptolide-5min GSE118494.NCAPH2.IMR-90_Triptolide-5min 232 bp overlap
ChIP IMR-90_alpha-amanitin-30min GSE118494.NCAPH2.IMR-90_alpha-amanitin-30min 317 bp overlap
NCOA3 1 dataset
ChIP MCF-7_E2 ERP000901.NCOA3.MCF-7_E2 160 bp overlap
NFYC 1 dataset
ChIP Hep-G2 ENCSR569ARC.NFYC.Hep-G2 62 bp overlap
PATZ1 1 dataset
ChIP HepG2 ENCFF723PFC 358 bp overlap
PHF5A 1 dataset
ChIP Hep-G2 ENCSR191TLD.PHF5A.Hep-G2 234 bp overlap
POU4F2 1 dataset
ChIP HNPC_DIF GSE74814.POU4F2.HNPC_DIF 126 bp overlap
RAD21 1 dataset
ChIP IMR-90 ENCSR000EFJ.RAD21.IMR-90 114 bp overlap
RCOR1 1 dataset
ChIP IMR-90 ENCSR000EFG.RCOR1.IMR-90 158 bp overlap
RELA 13 datasets
ChIP HAEC_IL1b_4h GSE89970.RELA.HAEC_IL1b_4h 211 bp overlap
ChIP HAEC_TNFa_4h GSE89970.RELA.HAEC_TNFa_4h 78 bp overlap
ChIP aortic-endothelial-cell_IL1B_D14 GSE139377.RELA.aortic-endothelial-cell_IL1B_D14 293 bp overlap
ChIP aortic-endothelial-cell_IL1B_D21 GSE139377.RELA.aortic-endothelial-cell_IL1B_D21 203 bp overlap
ChIP aortic-endothelial-cell_IL1B_D25 GSE139377.RELA.aortic-endothelial-cell_IL1B_D25 174 bp overlap
ChIP aortic-endothelial-cell_IL1B_D38 GSE139377.RELA.aortic-endothelial-cell_IL1B_D38 265 bp overlap
ChIP aortic-endothelial-cell_IL1B_D4 GSE139377.RELA.aortic-endothelial-cell_IL1B_D4 316 bp overlap
ChIP aortic-endothelial-cell_IL1B_D40 GSE139377.RELA.aortic-endothelial-cell_IL1B_D40 218 bp overlap
ChIP aortic-endothelial-cell_IL1B_D41 GSE139377.RELA.aortic-endothelial-cell_IL1B_D41 79 bp overlap
ChIP aortic-endothelial-cell_IL1B_D45 GSE139377.RELA.aortic-endothelial-cell_IL1B_D45 258 bp overlap
ChIP aortic-endothelial-cell_IL1B_D47 GSE139377.RELA.aortic-endothelial-cell_IL1B_D47 190 bp overlap
ChIP aortic-endothelial-cell_IL1B_D7 GSE139377.RELA.aortic-endothelial-cell_IL1B_D7 191 bp overlap
ChIP aortic-endothelial-cell_IL1B_D9 GSE139377.RELA.aortic-endothelial-cell_IL1B_D9 325 bp overlap
SALL1 1 dataset
ChIP HepG2 ENCFF426MCK 398 bp overlap
SETX 1 dataset
ChIP A-549_Influenza_PR8_NS1 GSE52936.SETX.A-549_Influenza_PR8_NS1 116 bp overlap
SMAD3 2 datasets
ChIP HCC1954 GSE104760.SMAD3.HCC1954 154 bp overlap
ChIP HCC1954_TGFb GSE104760.SMAD3.HCC1954_TGFb 176 bp overlap
SMARCA2 1 dataset
ChIP NPC_R1159Q_AM GSE122631.SMARCA2.NPC_R1159Q_AM 130 bp overlap
SMARCA4 7 datasets
ChIP A-549_AG15725 GSE132290.SMARCA4.A-549_AG15725 63 bp overlap
ChIP BIN-67_lentivirus-SMARCA4 GSE117734.SMARCA4.BIN-67_lentivirus-SMARCA4 98 bp overlap
ChIP MCF-7 GSE123284.SMARCA4.MCF-7 219 bp overlap
ChIP MCF-7_ARID1A-KO_4-OHT_clone14 GSE123284.SMARCA4.MCF-7_ARID1A-KO_4-OHT_clone14 423 bp overlap
ChIP MCF-7_ARID1A-KO_clone14 GSE123284.SMARCA4.MCF-7_ARID1A-KO_clone14 195 bp overlap
ChIP MCF-7_parental GSE123284.SMARCA4.MCF-7_parental 345 bp overlap
ChIP endometrial-epithelial-cells_KO GSE106660.SMARCA4.endometrial-epithelial-cells_KO 345 bp overlap
SMARCB1 7 datasets
ChIP MCF-7 GSE123284.SMARCB1.MCF-7 83 bp overlap
ChIP MCF-7_4-hydroxytamoxifen GSE123284.SMARCB1.MCF-7_4-hydroxytamoxifen 186 bp overlap
ChIP MCF-7_DCDT GSE123284.SMARCB1.MCF-7_DCDT 257 bp overlap
ChIP MCF-7_Fulvestrant GSE123284.SMARCB1.MCF-7_Fulvestrant 104 bp overlap
ChIP MCF-7_JQ1 GSE123284.SMARCB1.MCF-7_JQ1 159 bp overlap
ChIP MCF-7_estrogen GSE123284.SMARCB1.MCF-7_estrogen 295 bp overlap
ChIP proliferating-human-fibroblast_ASO_Ctrl GSE131711.SMARCB1.proliferating-human-fibroblast_ASO_Ctrl 423 bp overlap
SMC1 1 dataset
ChIP HCAEC GSE101921.SMC1.HCAEC 242 bp overlap
SNAI2 1 dataset
ChIP PC-9_2DF GSE131687.SNAI2.PC-9_2DF 411 bp overlap
SOX6 1 dataset
ChIP HepG2 ENCFF767OCK 316 bp overlap
SPI1 3 datasets
ChIP CD34_FETAL GSE70660.SPI1.CD34_FETAL 245 bp overlap
ChIP K-562 GSE70482.SPI1.K-562 151 bp overlap
ChIP NCI-H929 GSE56857.SPI1.NCI-H929 105 bp overlap
SS18 1 dataset
ChIP BIN-67_lentivirus-SMARCA4 GSE117734.SS18.BIN-67_lentivirus-SMARCA4 115 bp overlap
STAG1 2 datasets
ChIP Hep-G2 ENCSR167MTG.STAG1.Hep-G2 118 bp overlap
ChIP Hep-G2 ERP000209.STAG1.Hep-G2 146 bp overlap
STAT3 2 datasets
ChIP MCF-7_jc5840 GSE126004.STAT3.MCF-7_jc5840 95 bp overlap
ChIP MCF-7_jc5848 GSE126004.STAT3.MCF-7_jc5848 63 bp overlap
TEAD3 2 datasets
ChIP HepG2 ENCFF054UUL 371 bp overlap
ChIP HepG2 ENCFF054UUL 178 bp overlap
TEAD4 4 datasets
ChIP PC-9_1DF GSE131687.TEAD4.PC-9_1DF 86 bp overlap
ChIP PC-9_1DF_DMSO GSE131687.TEAD4.PC-9_1DF_DMSO 131 bp overlap
ChIP PC-9_2DF GSE131687.TEAD4.PC-9_2DF 238 bp overlap
ChIP hMSC-TERT4 GSE140782.TEAD4.hMSC-TERT4 223 bp overlap
TFEB 1 dataset
ChIP HUVEC-C GSE88894.TFEB.HUVEC-C 165 bp overlap
THRA 1 dataset
ChIP HepG2 ENCFF025KMX 385 bp overlap
THRB 2 datasets
ChIP Hep-G2 ENCSR430JGJ.THRB.Hep-G2 393 bp overlap
ChIP HepG2 ENCFF476INC 416 bp overlap
XRCC5 2 datasets
ChIP K-562 ENCSR506KWJ.XRCC5.K-562 174 bp overlap
ChIP K-562 GSE120104.XRCC5.K-562 155 bp overlap
YY1 1 dataset
ChIP HCT-116 ENCSR000BNX.YY1.HCT-116 102 bp overlap
YY1AP1 3 datasets
ChIP PC-9_1DF GSE131687.YY1AP1.PC-9_1DF 226 bp overlap
ChIP PC-9_1DF_DMSO GSE131687.YY1AP1.PC-9_1DF_DMSO 104 bp overlap
ChIP PC-9_2DF_DMSO GSE131687.YY1AP1.PC-9_2DF_DMSO 98 bp overlap
ZBTB7B 2 datasets
ChIP Hep-G2 ENCSR322ULL.ZBTB7B.Hep-G2 416 bp overlap
ChIP HepG2 ENCFF763OCV 423 bp overlap
ZNF217 2 datasets
ChIP Hep-G2 ENCSR055FQB.ZNF217.Hep-G2 177 bp overlap
ChIP MCF-7 ENCSR465XQW.ZNF217.MCF-7 67 bp overlap
ZNF219 1 dataset
ChIP HepG2 ENCFF266JIR 362 bp overlap
ZNF26 2 datasets
ChIP HEK293 ENCSR028EGI.ZNF26.HEK293 131 bp overlap
ChIP HEK293T GSE78099.ZNF26.HEK293T 199 bp overlap
ZNF750 1 dataset
ChIP keratinocyte_diff GSE57702.ZNF750.keratinocyte_diff 116 bp overlap