chr4 : 155,761,883 155,762,290
407 bp 162 TFs 1 linked gene
This 407 bp open chromatin element is linked to GUCY1B1 and is bound by 162 transcription factors.
Linked Genes
1 gene
Gene Expression Dist. to TSS Distance Link type
GUCY1B1 2.9 kb Proximal Proximity
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr4:155,756,883 – 155,767,290
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
162 transcription factors
Source
Cell type
ALX3 1 dataset
Motif DE_12h DE_12h-ALX3_MA0634.2 6 bp overlap
ATF7 1 dataset
ChIP GM12878 ENCSR014YCR.ATF7.GM12878 248 bp overlap
Arid3a 2 datasets
Motif DE_12h DE_12h-Arid3a_MA0151.1 6 bp overlap
Motif DE_12h DE_12h-Arid3a_MA0151.1 6 bp overlap
BARX1 1 dataset
Motif DE_12h DE_12h-BARX1_MA0875.2 6 bp overlap
BCOR 1 dataset
ChIP WA01_RNF2-R GSE104690.BCOR.WA01_RNF2-R 224 bp overlap
BSX 1 dataset
Motif DE_12h DE_12h-BSX_MA0876.2 6 bp overlap
Bcl11B 1 dataset
Motif DE_12h DE_12h-Bcl11B_MA1989.2 9 bp overlap
CEBPB 1 dataset
ChIP Ishikawa ENCSR000BTT.CEBPB.Ishikawa 159 bp overlap
DLX1 1 dataset
Motif DE_12h DE_12h-DLX1_MA0879.3 6 bp overlap
DLX6 1 dataset
Motif DE_12h DE_12h-DLX6_MA0882.2 6 bp overlap
DRGX 1 dataset
Motif DE_12h DE_12h-DRGX_MA1481.2 6 bp overlap
Dlx3 1 dataset
Motif DE_12h DE_12h-Dlx3_MA0880.2 6 bp overlap
Dlx4 1 dataset
Motif DE_12h DE_12h-Dlx4_MA0881.2 6 bp overlap
E2F8 1 dataset
ChIP GM12878 ENCSR793HVL.E2F8.GM12878 257 bp overlap
EMX1 1 dataset
Motif DE_12h DE_12h-EMX1_MA0612.3 6 bp overlap
EMX2 1 dataset
Motif DE_12h DE_12h-EMX2_MA0886.2 6 bp overlap
EN1 1 dataset
Motif DE_12h DE_12h-EN1_MA0027.3 6 bp overlap
EP300 2 datasets
ChIP Ishikawa ENCFF364ZWT 333 bp overlap
ChIP Ishikawa ENCSR000BUE.EP300.Ishikawa 407 bp overlap
ESR1 19 datasets
ChIP Ishikawa ENCSR000BIY.ESR1.Ishikawa 361 bp overlap
ChIP Ishikawa ENCSR000BQR.ESR1.Ishikawa 287 bp overlap
ChIP Ishikawa ENCSR000BKL.ESR1.Ishikawa 253 bp overlap
ChIP Ishikawa_C1-5 GSE147141.ESR1.Ishikawa_C1-5 398 bp overlap
ChIP Ishikawa_C2-3 GSE147141.ESR1.Ishikawa_C2-3 317 bp overlap
ChIP Ishikawa_DMSO_D538G_clone3 GSE132426.ESR1.Ishikawa_DMSO_D538G_clone3 341 bp overlap
ChIP Ishikawa_Dex_E2 GSE109891.ESR1.Ishikawa_Dex_E2 162 bp overlap
ChIP Ishikawa_E2 GSE132426.ESR1.Ishikawa_E2 343 bp overlap
ChIP Ishikawa_E2 GSE109891.ESR1.Ishikawa_E2 247 bp overlap
ChIP Ishikawa_E2_D538G GSE132426.ESR1.Ishikawa_E2_D538G 407 bp overlap
ChIP Ishikawa_E2_D538G_clone2 GSE132426.ESR1.Ishikawa_E2_D538G_clone2 407 bp overlap
ChIP Ishikawa_E2_clone2 GSE132426.ESR1.Ishikawa_E2_clone2 229 bp overlap
ChIP Ishikawa_ETV4-KO1 GSE129803.ESR1.Ishikawa_ETV4-KO1 261 bp overlap
ChIP Ishikawa_ETV4-KO1_Mili GSE129803.ESR1.Ishikawa_ETV4-KO1_Mili 337 bp overlap
ChIP Ishikawa_ETV4-KO2_Mili GSE129803.ESR1.Ishikawa_ETV4-KO2_Mili 357 bp overlap
ChIP Ishikawa_ETV4-KO2_Rescue GSE129803.ESR1.Ishikawa_ETV4-KO2_Rescue 352 bp overlap
ChIP Ishikawa_M1-66 GSE147141.ESR1.Ishikawa_M1-66 353 bp overlap
ChIP Ishikawa_M1-9-1 GSE147141.ESR1.Ishikawa_M1-9-1 392 bp overlap
ChIP Ishikawa_M2-5 GSE147141.ESR1.Ishikawa_M2-5 343 bp overlap
EVX1 1 dataset
Motif DE_12h DE_12h-EVX1_MA0887.2 6 bp overlap
EVX2 1 dataset
Motif DE_12h DE_12h-EVX2_MA0888.2 6 bp overlap
FOXA1 5 datasets
ChIP CFPAC-1 GSE119930.FOXA1.CFPAC-1 284 bp overlap
ChIP Ishikawa ENCSR000BKW.FOXA1.Ishikawa 316 bp overlap
ChIP PDAC GSE64557.FOXA1.PDAC 221 bp overlap
ChIP PDAC_KOKLF5 GSE64557.FOXA1.PDAC_KOKLF5 360 bp overlap
ChIP ZR-75-1_estrogen_ab2 GSE112969.FOXA1.ZR-75-1_estrogen_ab2 214 bp overlap
FOXA2 5 datasets
ChIP CFPAC-1_HNF1B-KO GSE119930.FOXA2.CFPAC-1_HNF1B-KO 209 bp overlap
ChIP CFPAC-1_HOXB8-OE GSE119930.FOXA2.CFPAC-1_HOXB8-OE 201 bp overlap
ChIP DE DE-FOXA2-1 303 bp overlap
ChIP DE DE-FOXA2-2 259 bp overlap
ChIP PANC-1_HNF1B-OE GSE119930.FOXA2.PANC-1_HNF1B-OE 221 bp overlap
FOXM1 2 datasets
ChIP Ishikawa ENCFF578VDD 393 bp overlap
ChIP Ishikawa ENCSR000BUS.FOXM1.Ishikawa 257 bp overlap
GATA6 2 datasets
ChIP DE_D1 S41-DE-d1-GATA6-exp2 267 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 284 bp overlap
GBX2 1 dataset
Motif DE_12h DE_12h-GBX2_MA0890.2 6 bp overlap
GLIS1 2 datasets
ChIP HEK293 ENCFF299RSE 389 bp overlap
ChIP HEK293 ENCSR482BBZ.GLIS1.HEK293 407 bp overlap
GLIS3 1 dataset
ChIP H9_plus GSE109562.GLIS3.H9_plus 352 bp overlap
GSX1 1 dataset
Motif DE_12h DE_12h-GSX1_MA0892.2 6 bp overlap
GSX2 1 dataset
Motif DE_12h DE_12h-GSX2_MA0893.3 7 bp overlap
HESX1 1 dataset
Motif DE_12h DE_12h-HESX1_MA0894.2 6 bp overlap
HIC2 1 dataset
Motif DE_12h DE_12h-HIC2_MA0738.2 6 bp overlap
HOXA1 1 dataset
Motif DE_12h DE_12h-HOXA1_MA1495.2 6 bp overlap
HOXA2 1 dataset
Motif DE_12h DE_12h-HOXA2_MA0900.3 6 bp overlap
HOXA3 1 dataset
Motif DE_12h DE_12h-HOXA3_MA2119.1 7 bp overlap
HOXA5 1 dataset
Motif DE_12h DE_12h-HOXA5_MA0158.2 8 bp overlap
HOXA6 1 dataset
Motif DE_12h DE_12h-HOXA6_MA1497.2 7 bp overlap
HOXA7 1 dataset
Motif DE_12h DE_12h-HOXA7_MA1498.3 6 bp overlap
HOXB1 1 dataset
Motif DE_12h DE_12h-HOXB1_MA2093.1 7 bp overlap
HOXB2 1 dataset
Motif DE_12h DE_12h-HOXB2_MA0902.3 6 bp overlap
HOXB3 1 dataset
Motif DE_12h DE_12h-HOXB3_MA0903.2 6 bp overlap
HOXB5 1 dataset
Motif DE_12h DE_12h-HOXB5_MA0904.3 6 bp overlap
HOXB6 1 dataset
Motif DE_12h DE_12h-HOXB6_MA1500.2 7 bp overlap
HOXB7 1 dataset
Motif DE_12h DE_12h-HOXB7_MA1501.2 7 bp overlap
HOXB8 1 dataset
Motif DE_12h DE_12h-HOXB8_MA1502.2 7 bp overlap
HOXC8 1 dataset
Motif DE_12h DE_12h-HOXC8_MA1505.2 6 bp overlap
HOXD8 1 dataset
Motif DE_12h DE_12h-HOXD8_MA0910.3 7 bp overlap
ISX 1 dataset
Motif DE_12h DE_12h-ISX_MA0654.2 6 bp overlap
Irf1 1 dataset
Motif DE_12h DE_12h-Irf1_MA0050.4 11 bp overlap
JUN 1 dataset
ChIP HUES-8_DE_ctrl GSE109524.JUN.HUES-8_DE_ctrl 337 bp overlap
KLF1 1 dataset
Motif DE_12h DE_12h-KLF1_MA0493.3 8 bp overlap
KLF11 1 dataset
Motif DE_12h DE_12h-KLF11_MA1512.2 10 bp overlap
KLF2 1 dataset
Motif DE_12h DE_12h-KLF2_MA1515.2 8 bp overlap
KLF3 1 dataset
Motif DE_12h DE_12h-KLF3_MA1516.2 10 bp overlap
KLF4 1 dataset
Motif DE_12h DE_12h-KLF4_MA0039.5 8 bp overlap
KLF5 1 dataset
Motif DE_12h DE_12h-KLF5_MA0599.1 10 bp overlap
KLF6 1 dataset
Motif DE_12h DE_12h-KLF6_MA1517.2 9 bp overlap
KLF9 1 dataset
Motif DE_12h DE_12h-KLF9_MA1107.3 11 bp overlap
LBX2 1 dataset
Motif DE_12h DE_12h-LBX2_MA0699.2 6 bp overlap
LHX2 1 dataset
Motif DE_12h DE_12h-LHX2_MA0700.3 6 bp overlap
LHX5 1 dataset
Motif DE_12h DE_12h-LHX5_MA1519.2 7 bp overlap
Lhx4 1 dataset
Motif DE_12h DE_12h-Lhx4_MA0704.2 6 bp overlap
Lhx8 1 dataset
Motif DE_12h DE_12h-Lhx8_MA0705.2 6 bp overlap
MAX 1 dataset
ChIP Ishikawa ENCSR000BTY.MAX.Ishikawa 177 bp overlap
MAZ 1 dataset
Motif DE_12h DE_12h-MAZ_MA1522.2 8 bp overlap
MEOX1 1 dataset
Motif DE_12h DE_12h-MEOX1_MA0661.2 7 bp overlap
MEOX2 1 dataset
Motif DE_12h DE_12h-MEOX2_MA0706.2 7 bp overlap
MIXL1 1 dataset
Motif DE_12h DE_12h-MIXL1_MA0662.2 6 bp overlap
MLLT1 1 dataset
ChIP GM12878 ENCSR552XSN.MLLT1.GM12878 340 bp overlap
MNX1 1 dataset
Motif DE_12h DE_12h-MNX1_MA0707.3 6 bp overlap
MSX1 1 dataset
Motif DE_12h DE_12h-MSX1_MA0666.3 6 bp overlap
MSX2 1 dataset
Motif DE_12h DE_12h-MSX2_MA0708.3 6 bp overlap
Msx3 1 dataset
Motif DE_12h DE_12h-Msx3_MA0709.2 6 bp overlap
NANOG 2 datasets
ChIP WA01_3IL ERP004238.NANOG.WA01_3IL 352 bp overlap
ChIP hESC GSE18292.NANOG.hESC 98 bp overlap
NFATC3 2 datasets
Motif DE_12h DE_12h-NFATC3_MA0625.3 6 bp overlap
Motif DE_12h DE_12h-NFATC3_MA0625.3 6 bp overlap
NFIC 2 datasets
ChIP Ishikawa ENCFF029AAD 345 bp overlap
ChIP Ishikawa ENCSR000BUT.NFIC.Ishikawa 374 bp overlap
NFYB 1 dataset
Motif DE_12h DE_12h-NFYB_MA0502.3 9 bp overlap
NKX6-2 1 dataset
Motif DE_12h DE_12h-NKX6-2_MA0675.2 6 bp overlap
NR1D1 1 dataset
Motif DE_12h DE_12h-NR1D1_MA1531.2 14 bp overlap
NR1D2 1 dataset
Motif DE_12h DE_12h-NR1D2_MA1532.2 15 bp overlap
NR1I2 1 dataset
Motif DE_12h DE_12h-NR1I2_MA1533.2 15 bp overlap
NR1I3 1 dataset
Motif DE_12h DE_12h-NR1I3_MA1534.2 8 bp overlap
NR3C1 3 datasets
ChIP Ishikawa ENCSR000BJC.NR3C1.Ishikawa 339 bp overlap
ChIP Ishikawa ENCSR000BLB.NR3C1.Ishikawa 123 bp overlap
ChIP Ishikawa_Dex GSE109891.NR3C1.Ishikawa_Dex 356 bp overlap
Nfatc1 2 datasets
Motif DE_12h DE_12h-Nfatc1_MA0624.3 6 bp overlap
Motif DE_12h DE_12h-Nfatc1_MA0624.3 6 bp overlap
Nobox 1 dataset
Motif DE_12h DE_12h-Nobox_MA0125.2 6 bp overlap
Olig2 1 dataset
Motif DE_12h DE_12h-Olig2_MA1997.2 6 bp overlap
PDX1 1 dataset
Motif DE_12h DE_12h-PDX1_MA0132.3 6 bp overlap
POU4F2 1 dataset
Motif DE_12h DE_12h-POU4F2_MA0683.2 15 bp overlap
POU6F1 1 dataset
Motif DE_12h DE_12h-POU6F1_MA0628.2 6 bp overlap
PRDM9 1 dataset
Motif DE_12h DE_12h-PRDM9_MA1723.2 20 bp overlap
PRRX1 1 dataset
Motif DE_12h DE_12h-PRRX1_MA0716.2 6 bp overlap
Prdm15 1 dataset
Motif DE_12h DE_12h-Prdm15_MA1616.2 11 bp overlap
RAD21 1 dataset
ChIP Ishikawa ENCSR000BTU.RAD21.Ishikawa 163 bp overlap
RAD51 1 dataset
ChIP GM12878 ENCSR482TWQ.RAD51.GM12878 163 bp overlap
RARA 1 dataset
ChIP hiPSC_D4 GSE109172.RARA.hiPSC_D4 230 bp overlap
RAX 1 dataset
Motif DE_12h DE_12h-RAX_MA0718.2 6 bp overlap
RAX2 1 dataset
Motif DE_12h DE_12h-RAX2_MA0717.2 6 bp overlap
RELB 1 dataset
ChIP GM12878 ENCSR387QUV.RELB.GM12878 271 bp overlap
RREB1 1 dataset
Motif DE_12h DE_12h-RREB1_MA0073.2 19 bp overlap
RUNX2 1 dataset
Motif DE_12h DE_12h-RUNX2_MA0511.2 9 bp overlap
Rarg 1 dataset
Motif DE_12h DE_12h-Rarg_MA0860.1 17 bp overlap
Runx1 1 dataset
Motif DE_12h DE_12h-Runx1_MA0002.3 9 bp overlap
SHOX 1 dataset
Motif DE_12h DE_12h-SHOX_MA0630.2 6 bp overlap
SMAD2 1 dataset
Motif DE_12h DE_12h-SMAD2_MA1964.2 6 bp overlap
SMAD2-3 1 dataset
ChIP HUES-8_DE_ctrl GSE109524.SMAD2-3.HUES-8_DE_ctrl 362 bp overlap
SMARCA4 2 datasets
ChIP BIN-67_lentivirus-SMARCA4-T910M GSE117734.SMARCA4.BIN-67_lentivirus-SMARCA4-T910M 282 bp overlap
ChIP hiPSC GSE124903.SMARCA4.hiPSC 396 bp overlap
SMARCC1 3 datasets
ChIP BIN-67_lentivirus-SMARCA4-T910M GSE117734.SMARCC1.BIN-67_lentivirus-SMARCA4-T910M 202 bp overlap
ChIP HS-SY-2 GSE108025.SMARCC1.HS-SY-2 311 bp overlap
ChIP hiPSC GSE124903.SMARCC1.hiPSC 407 bp overlap
SP1 2 datasets
Motif DE_12h DE_12h-SP1_MA0079.5 9 bp overlap
Motif DE_12h DE_12h-SP1_MA0079.5 9 bp overlap
SP3 1 dataset
Motif DE_12h DE_12h-SP3_MA0746.3 11 bp overlap
SP4 2 datasets
Motif DE_12h DE_12h-SP4_MA0685.2 9 bp overlap
Motif DE_12h DE_12h-SP4_MA0685.2 9 bp overlap
SP5 2 datasets
Motif DE_12h DE_12h-SP5_MA1965.2 6 bp overlap
Motif DE_12h DE_12h-SP5_MA1965.2 6 bp overlap
SS18 1 dataset
ChIP BIN-67_lentivirus-SMARCA4-T910M GSE117734.SS18.BIN-67_lentivirus-SMARCA4-T910M 243 bp overlap
STAT1 1 dataset
Motif DE_12h DE_12h-STAT1_MA0137.4 9 bp overlap
STAT3 2 datasets
Motif DE_12h DE_12h-STAT3_MA0144.3 9 bp overlap
ChIP MDA-MB-157_EtOH GSE85579.STAT3.MDA-MB-157_EtOH 313 bp overlap
Shox2 1 dataset
Motif DE_12h DE_12h-Shox2_MA0720.2 6 bp overlap
Sox1 1 dataset
Motif DE_12h DE_12h-Sox1_MA0870.1 15 bp overlap
Stat4 1 dataset
Motif DE_12h DE_12h-Stat4_MA0518.2 10 bp overlap
Stat5b 1 dataset
Motif DE_12h DE_12h-Stat5b_MA1625.2 9 bp overlap
Stat6 1 dataset
Motif DE_12h DE_12h-Stat6_MA0520.2 10 bp overlap
T 2 datasets
ChIP H9_ENDODERM GSE60606.T.H9_ENDODERM 232 bp overlap
ChIP H9_MESODERM GSE60606.T.H9_MESODERM 304 bp overlap
TCF12 2 datasets
ChIP Ishikawa ENCFF467DDW 391 bp overlap
ChIP Ishikawa ENCSR000BUV.TCF12.Ishikawa 407 bp overlap
TEAD4 2 datasets
ChIP Ishikawa ENCFF772OTG 298 bp overlap
ChIP Ishikawa ENCSR000BSW.TEAD4.Ishikawa 384 bp overlap
TFAP4 2 datasets
Motif DE_12h DE_12h-TFAP4_MA0691.1 10 bp overlap
ChIP DLD-1 GSE46935.TFAP4.DLD-1 251 bp overlap
TLX2 1 dataset
Motif DE_12h DE_12h-TLX2_MA1577.2 6 bp overlap
Tcf12 1 dataset
Motif DE_12h DE_12h-Tcf12_MA0521.3 6 bp overlap
Twist2 1 dataset
Motif DE_12h DE_12h-Twist2_MA0633.3 6 bp overlap
UNCX 1 dataset
Motif DE_12h DE_12h-UNCX_MA0721.2 6 bp overlap
VAX2 1 dataset
Motif DE_12h DE_12h-VAX2_MA0723.3 6 bp overlap
VDR 1 dataset
ChIP kidney-cortex GSE129585.VDR.kidney-cortex 402 bp overlap
Vdr 1 dataset
Motif DE_12h DE_12h-Vdr_MA0693.4 7 bp overlap
YY1 1 dataset
ChIP Ishikawa ENCSR000BSY.YY1.Ishikawa 292 bp overlap
ZBTB42 1 dataset
ChIP HEK293 GSE76494.ZBTB42.HEK293 291 bp overlap
ZFP14 2 datasets
Motif DE_12h DE_12h-ZFP14_MA1972.1 15 bp overlap
Motif DE_12h DE_12h-ZFP14_MA1972.1 15 bp overlap
ZIC2 1 dataset
ChIP HEK293 ENCFF033NQQ 393 bp overlap
ZNF140 1 dataset
Motif DE_12h DE_12h-ZNF140_MA1589.2 19 bp overlap
ZNF148 2 datasets
Motif DE_12h DE_12h-ZNF148_MA1653.2 10 bp overlap
Motif DE_12h DE_12h-ZNF148_MA1653.2 10 bp overlap
ZNF213 1 dataset
Motif DE_12h DE_12h-ZNF213_MA2121.1 12 bp overlap
ZNF263 2 datasets
Motif DE_12h DE_12h-ZNF263_MA0528.3 7 bp overlap
Motif DE_12h DE_12h-ZNF263_MA0528.3 7 bp overlap
ZNF281 1 dataset
Motif DE_12h DE_12h-ZNF281_MA1630.3 10 bp overlap
ZNF416 1 dataset
Motif DE_12h DE_12h-ZNF416_MA1979.2 10 bp overlap
ZNF549 1 dataset
Motif DE_12h DE_12h-ZNF549_MA1728.2 8 bp overlap
ZNF701 1 dataset
Motif DE_12h DE_12h-ZNF701_MA1987.2 17 bp overlap