chr5 : 174,425,721 174,426,822
1,101 bp 121 TFs 1 linked gene
This 1.1 kb open chromatin element is linked to MSX2 and is bound by 121 transcription factors.
Linked Genes
1 gene
Gene Expression Dist. to TSS Distance Link type
MSX2 298.5 kb Distal Multiome+HiCAR
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr5:174,420,721 – 174,431,822
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
121 transcription factors
Source
Cell type
Atoh1 1 dataset
Motif DE_72h DE_72h-Atoh1_MA1467.3 7 bp overlap
BRD2 1 dataset
ChIP HCC1806_BAZ2B_JQ1 GSE116879.BRD2.HCC1806_BAZ2B_JQ1 52 bp overlap
BRD4 8 datasets
ChIP COLO-205 GSE73319.BRD4.COLO-205 259 bp overlap
ChIP COLO-205 GSE73319.BRD4.COLO-205 147 bp overlap
ChIP HT29 GSE73319.BRD4.HT29 137 bp overlap
ChIP Kelly_sensitive GSE103030.BRD4.Kelly_sensitive 239 bp overlap
ChIP RH4_shCHD4 GSE140115.BRD4.RH4_shCHD4 92 bp overlap
ChIP SW480 GSE73319.BRD4.SW480 759 bp overlap
ChIP SW480 GSE73319.BRD4.SW480 75 bp overlap
ChIP keratinocyte GSE140992.BRD4.keratinocyte 301 bp overlap
CHD4 1 dataset
ChIP SCMC GSE155861.CHD4.SCMC 297 bp overlap
CHD8 2 datasets
ChIP T-47D_ETOH_45 GSE62428.CHD8.T-47D_ETOH_45 160 bp overlap
ChIP T-47D_ETOH_5 GSE62428.CHD8.T-47D_ETOH_5 286 bp overlap
CTCF 2 datasets
ChIP RH4 GSE83726.CTCF.RH4 218 bp overlap
ChIP hiPSC_IIIB12 GSE106870.CTCF.hiPSC_IIIB12 130 bp overlap
EBF1 2 datasets
Motif DE_60h DE_60h-EBF1_MA0154.5 11 bp overlap
Motif DE_72h DE_72h-EBF1_MA0154.5 11 bp overlap
EBF3 2 datasets
Motif DE_60h DE_60h-EBF3_MA1637.2 9 bp overlap
Motif DE_72h DE_72h-EBF3_MA1637.2 9 bp overlap
ELF3 1 dataset
ChIP PDAC_SHCTR GSE64557.ELF3.PDAC_SHCTR 50 bp overlap
EOMES 2 datasets
ChIP hESC GSE26097.EOMES.hESC 267 bp overlap
ChIP hESC GSE26097.EOMES.hESC 171 bp overlap
ESR1 3 datasets
ChIP MCF-7_E2_45m GSE67295.ESR1.MCF-7_E2_45m 188 bp overlap
ChIP T-47D_Y537S GSE148277.ESR1.T-47D_Y537S 168 bp overlap
ChIP T-47D_Y537S_E2 GSE148277.ESR1.T-47D_Y537S_E2 204 bp overlap
ESRRA 3 datasets
ChIP BT-474 GSE81651.ESRRA.BT-474 795 bp overlap
ChIP BT-474_EGF GSE81651.ESRRA.BT-474_EGF 362 bp overlap
ChIP BT-474_HRG GSE81651.ESRRA.BT-474_HRG 501 bp overlap
ESRRG 1 dataset
ChIP cardiomyocyte GSE113760.ESRRG.cardiomyocyte 529 bp overlap
EZH2 7 datasets
ChIP A-1847 GSE95643.EZH2.A-1847 707 bp overlap
ChIP DND41 ENCSR000ASW.EZH2.DND41 267 bp overlap
ChIP Karpas-422_CPI360-D8 GSE134136.EZH2.Karpas-422_CPI360-D8 242 bp overlap
ChIP PC-9 ENCSR793USK.EZH2.PC-9 193 bp overlap
ChIP RH30_DMSO GSE85169.EZH2.RH30_DMSO 287 bp overlap
ChIP THP-1 GSE135024.EZH2.THP-1 559 bp overlap
ChIP THP-1 GSE135024.EZH2.THP-1 135 bp overlap
Ebf2 2 datasets
Motif DE_60h DE_60h-Ebf2_MA1604.2 9 bp overlap
Motif DE_72h DE_72h-Ebf2_MA1604.2 9 bp overlap
Ebf4 2 datasets
Motif DE_60h DE_60h-Ebf4_MA2122.1 11 bp overlap
Motif DE_72h DE_72h-Ebf4_MA2122.1 11 bp overlap
FEZF1 3 datasets
ChIP HEK293 ENCSR827NWO.FEZF1.HEK293 70 bp overlap
ChIP HEK293 ENCSR827NWO.FEZF1.HEK293 211 bp overlap
ChIP HEK293 ENCSR827NWO.FEZF1.HEK293 237 bp overlap
FLI1 5 datasets
ChIP A-673 GSE99959.FLI1.A-673 394 bp overlap
ChIP A-673_1_KRAB_GFP GSE106914.FLI1.A-673_1_KRAB_GFP 468 bp overlap
ChIP A-673_2_KRAB_eSOX2_1 GSE106914.FLI1.A-673_2_KRAB_eSOX2_1 393 bp overlap
ChIP A-673_D14 GSE129155.FLI1.A-673_D14 545 bp overlap
ChIP A-673_D17 GSE129155.FLI1.A-673_D17 463 bp overlap
FOXA2 2 datasets
ChIP DE DE-FOXA2-1 852 bp overlap
ChIP DE DE-FOXA2-2 827 bp overlap
GATA1 2 datasets
Motif DE_60h DE_60h-GATA1_MA0035.5 7 bp overlap
Motif DE_72h DE_72h-GATA1_MA0035.5 7 bp overlap
GATA4 4 datasets
ChIP DE DE-GATA4-1 965 bp overlap
ChIP DE DE-GATA4-2 944 bp overlap
ChIP Hep-G2 GSE135714.GATA4.Hep-G2 669 bp overlap
ChIP foregut GSE117136.GATA4.foregut 537 bp overlap
GATA6 8 datasets
ChIP DE DE-GATA6-1 946 bp overlap
ChIP DE DE-GATA6-2 1064 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 354 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 893 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 557 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 929 bp overlap
ChIP foregut GSE117136.GATA6.foregut 758 bp overlap
ChIP foregut_KO GSE117136.GATA6.foregut_KO 420 bp overlap
GLIS1 2 datasets
ChIP HEK293 ENCFF299RSE 287 bp overlap
ChIP HEK293 ENCSR482BBZ.GLIS1.HEK293 922 bp overlap
HAND2 1 dataset
Motif DE_72h DE_72h-HAND2_MA1638.2 6 bp overlap
HDAC2 1 dataset
ChIP RH4_Entinostat-6H GSE116344.HDAC2.RH4_Entinostat-6H 495 bp overlap
HDAC3 1 dataset
ChIP RH4_DMSO-6H GSE116344.HDAC3.RH4_DMSO-6H 57 bp overlap
HOXA9 1 dataset
ChIP HEK293-FT GSE62586.HOXA9.HEK293-FT 256 bp overlap
INSM1 1 dataset
Motif DE_72h DE_72h-INSM1_MA0155.1 12 bp overlap
JUN 1 dataset
ChIP DE_D2 S02-DE-d2-JUN-exp1 558 bp overlap
KDM6B 1 dataset
ChIP CUTLL1 GSE56694.KDM6B.CUTLL1 180 bp overlap
KLF1 1 dataset
Motif DE_36h DE_36h-KLF1_MA0493.3 8 bp overlap
KLF10 1 dataset
Motif DE_36h DE_36h-KLF10_MA1511.2 9 bp overlap
KLF14 1 dataset
Motif DE_36h DE_36h-KLF14_MA0740.2 9 bp overlap
KLF2 1 dataset
Motif DE_36h DE_36h-KLF2_MA1515.2 8 bp overlap
KLF4 1 dataset
Motif DE_36h DE_36h-KLF4_MA0039.5 8 bp overlap
KLF5 1 dataset
Motif DE_36h DE_36h-KLF5_MA0599.1 10 bp overlap
MAZ 1 dataset
ChIP HEK293 GSE76494.MAZ.HEK293 256 bp overlap
MED1 3 datasets
ChIP HCT-116_shLuc_thaps GSE121798.MED1.HCT-116_shLuc_thaps 301 bp overlap
ChIP RH4 GSE83726.MED1.RH4 162 bp overlap
ChIP RH4 GSE83726.MED1.RH4 153 bp overlap
MEIS1 3 datasets
Motif DE_60h DE_60h-MEIS1_MA0498.3 5 bp overlap
Motif DE_72h DE_72h-MEIS1_MA0498.3 5 bp overlap
Motif DE_72h DE_72h-MEIS1_MA0498.3 5 bp overlap
MEIS2 2 datasets
Motif DE_60h DE_60h-MEIS2_MA0774.1 8 bp overlap
Motif DE_72h DE_72h-MEIS2_MA0774.1 8 bp overlap
MEIS3 2 datasets
Motif DE_60h DE_60h-MEIS3_MA0775.2 7 bp overlap
Motif DE_72h DE_72h-MEIS3_MA0775.2 7 bp overlap
MYB 1 dataset
ChIP DU528 GSE94000.MYB.DU528 102 bp overlap
MYCN 1 dataset
ChIP RH4 GSE83726.MYCN.RH4 535 bp overlap
MYOD1 2 datasets
ChIP RH30_DMSO GSE85169.MYOD1.RH30_DMSO 199 bp overlap
ChIP RH4 GSE83726.MYOD1.RH4 566 bp overlap
MYOG 2 datasets
ChIP RH4 GSE83726.MYOG.RH4 156 bp overlap
ChIP RH4 GSE83726.MYOG.RH4 353 bp overlap
MZF1 2 datasets
ChIP HEK293 ENCSR298QUH.MZF1.HEK293 310 bp overlap
ChIP HEK293 ENCSR298QUH.MZF1.HEK293 320 bp overlap
Mafg 2 datasets
Motif DE_36h DE_36h-Mafg_MA0659.4 12 bp overlap
Motif DE_72h DE_72h-Mafg_MA0659.4 12 bp overlap
NEUROD1 9 datasets
ChIP D283-Med GSE92582.NEUROD1.D283-Med 222 bp overlap
ChIP D283-Med GSE92582.NEUROD1.D283-Med 406 bp overlap
ChIP D283-Med_shNEUROD1-1579 GSE92582.NEUROD1.D283-Med_shNEUROD1-1579 210 bp overlap
ChIP D283-Med_shNEUROD1-1579 GSE92582.NEUROD1.D283-Med_shNEUROD1-1579 356 bp overlap
ChIP D341-Med_shGFP GSE92582.NEUROD1.D341-Med_shGFP 233 bp overlap
ChIP D341-Med_shNEUROD1-1154 GSE92582.NEUROD1.D341-Med_shNEUROD1-1154 256 bp overlap
ChIP D341-Med_shNEUROD1-1154 GSE92582.NEUROD1.D341-Med_shNEUROD1-1154 318 bp overlap
ChIP D341-Med_shNEUROD1-1579 GSE92582.NEUROD1.D341-Med_shNEUROD1-1579 294 bp overlap
ChIP D341-Med_shNEUROD1-1579 GSE92582.NEUROD1.D341-Med_shNEUROD1-1579 322 bp overlap
NFIB 1 dataset
ChIP MCF-7 ENCSR702BYX.NFIB.MCF-7 192 bp overlap
NRF1 2 datasets
ChIP HeLa-S3 GSE108856.NRF1.HeLa-S3 148 bp overlap
ChIP HeLa-S3 GSE108856.NRF1.HeLa-S3 296 bp overlap
OSR2 3 datasets
ChIP HEK293 ENCSR324LTM.OSR2.HEK293 190 bp overlap
ChIP HEK293 ENCSR324LTM.OSR2.HEK293 366 bp overlap
ChIP HEK293 GSE76494.OSR2.HEK293 150 bp overlap
PATZ1 1 dataset
Motif DE_36h DE_36h-PATZ1_MA1961.2 11 bp overlap
PHIP 5 datasets
ChIP HCT-116_MLL1-KO__ab833 GSE101646.PHIP.HCT-116_MLL1-KO__ab833 612 bp overlap
ChIP HCT-116_ab833 GSE101646.PHIP.HCT-116_ab833 421 bp overlap
ChIP HCT-116_ab834 GSE101646.PHIP.HCT-116_ab834 397 bp overlap
ChIP HEK293_ab833 GSE101646.PHIP.HEK293_ab833 324 bp overlap
ChIP HEK293_ab833 GSE101646.PHIP.HEK293_ab833 58 bp overlap
POU2F1 1 dataset
ChIP T-47D_D538G GSE148277.POU2F1.T-47D_D538G 200 bp overlap
PRDM1 1 dataset
ChIP HEK293 GSE76494.PRDM1.HEK293 152 bp overlap
PRDM6 1 dataset
ChIP HEK293 ENCSR892QHR.PRDM6.HEK293 288 bp overlap
PRDM9 1 dataset
Motif DE_36h DE_36h-PRDM9_MA1723.2 20 bp overlap
PSIP1 1 dataset
ChIP T-REx-293 GSE117155.PSIP1.T-REx-293 165 bp overlap
RAD21 1 dataset
ChIP RH4 GSE83726.RAD21.RH4 413 bp overlap
RARA 1 dataset
ChIP hiPSC_D5 GSE132532.RARA.hiPSC_D5 182 bp overlap
RBBP4 1 dataset
ChIP SCMC GSE155861.RBBP4.SCMC 360 bp overlap
REST 3 datasets
Motif DE_36h DE_36h-REST_MA0138.3 20 bp overlap
Motif DE_60h DE_60h-REST_MA0138.3 20 bp overlap
Motif DE_72h DE_72h-REST_MA0138.3 20 bp overlap
SCRT2 1 dataset
ChIP HEK293 ENCFF711QQB 204 bp overlap
SMAD2 1 dataset
ChIP endoderm GSE29422.SMAD2.endoderm 293 bp overlap
SMAD2-3 2 datasets
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 1101 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.SMAD2-3.HUES-8_DE_JNKi 346 bp overlap
SMAD2_3 2 datasets
ChIP DE_D2 S01-DE-d2-SMAD2_3-exp1 1101 bp overlap
ChIP DE_D2 S53-DE-d2-SMAD2_3-exp2 1054 bp overlap
SMAD3 1 dataset
ChIP endoderm GSE29422.SMAD3.endoderm 251 bp overlap
SMAD4 2 datasets
ChIP endoderm GSE29422.SMAD4.endoderm 285 bp overlap
ChIP endoderm GSE29422.SMAD4.endoderm 140 bp overlap
SMARCC1 7 datasets
ChIP Aska-SS GSE108025.SMARCC1.Aska-SS 235 bp overlap
ChIP SK-N-MC GSE94275.SMARCC1.SK-N-MC 280 bp overlap
ChIP SYO-1 GSE108025.SMARCC1.SYO-1 441 bp overlap
ChIP SYO-1 GSE108025.SMARCC1.SYO-1 229 bp overlap
ChIP SYO-1_shCt GSE139053.SMARCC1.SYO-1_shCt 181 bp overlap
ChIP SYO-1_shCt GSE139053.SMARCC1.SYO-1_shCt 588 bp overlap
ChIP SYO-1_shRING1A-B GSE139053.SMARCC1.SYO-1_shRING1A-B 1033 bp overlap
SOX12 2 datasets
Motif DE_36h DE_36h-SOX12_MA1561.2 10 bp overlap
Motif DE_72h DE_72h-SOX12_MA1561.2 10 bp overlap
SOX14 2 datasets
Motif DE_36h DE_36h-SOX14_MA1562.2 9 bp overlap
Motif DE_72h DE_72h-SOX14_MA1562.2 9 bp overlap
SOX17 2 datasets
ChIP DE_D2 DED2-SOX17_Batch_II 428 bp overlap
ChIP DE_D2 DED2-SOX17_Batch_II 225 bp overlap
SOX17_M 1 dataset
ChIP DE_D2 DED2-SOX17-M_Batch_II 1100 bp overlap
SOX2 1 dataset
ChIP LK2_GFP GSE137459.SOX2.LK2_GFP 360 bp overlap
SOX21 2 datasets
Motif DE_36h DE_36h-SOX21_MA0866.1 15 bp overlap
Motif DE_72h DE_72h-SOX21_MA0866.1 15 bp overlap
SP5 2 datasets
Motif DE_36h DE_36h-SP5_MA1965.2 6 bp overlap
Motif DE_72h DE_72h-SP5_MA1965.2 6 bp overlap
SP7 2 datasets
ChIP HEK293 ENCSR468IJT.SP7.HEK293 323 bp overlap
ChIP HEK293 ENCSR468IJT.SP7.HEK293 278 bp overlap
SS18 5 datasets
ChIP Aska-SS GSE108025.SS18.Aska-SS 531 bp overlap
ChIP Aska-SS GSE108025.SS18.Aska-SS 326 bp overlap
ChIP Aska-SS_BAF47KO1 GSE108025.SS18.Aska-SS_BAF47KO1 269 bp overlap
ChIP Aska-SS_BAF47KO1 GSE108025.SS18.Aska-SS_BAF47KO1 284 bp overlap
ChIP SYO-1_shRING1A-B GSE139053.SS18.SYO-1_shRING1A-B 298 bp overlap
SUPT5H 1 dataset
ChIP HEK293_PNUTSW401A GSE134198.SUPT5H.HEK293_PNUTSW401A 217 bp overlap
Sox1 2 datasets
Motif DE_36h DE_36h-Sox1_MA0870.1 15 bp overlap
Motif DE_72h DE_72h-Sox1_MA0870.1 15 bp overlap
T 1 dataset
ChIP H9_MESODERM GSE60606.T.H9_MESODERM 183 bp overlap
TAL1 1 dataset
ChIP CHRF28811 ERP008568.TAL1.CHRF28811 291 bp overlap
TBX2 1 dataset
ChIP Kelly GSE94822.TBX2.Kelly 215 bp overlap
TEAD4 2 datasets
ChIP BE2C GSE84389.TEAD4.BE2C 374 bp overlap
ChIP SK-N-BE2 GSE84389.TEAD4.SK-N-BE2 374 bp overlap
TFAP2A 1 dataset
ChIP WA09 GSE105081.TFAP2A.WA09 407 bp overlap
TP53 10 datasets
ChIP HCT-116_5FU GSE58506.TP53.HCT-116_5FU 104 bp overlap
ChIP HCT-116_Nutlin3a GSE125927.TP53.HCT-116_Nutlin3a 411 bp overlap
ChIP HCT-116_nutlin GSE86164.TP53.HCT-116_nutlin 121 bp overlap
ChIP HCT-116_siGLIS2-1-5FU GSE125927.TP53.HCT-116_siGLIS2-1-5FU 210 bp overlap
ChIP MCF-10A_Nutlin3A GSE111009.TP53.MCF-10A_Nutlin3A 246 bp overlap
ChIP MCF-7_7-5h_IR_10Gy_Nutlin GSE100099.TP53.MCF-7_7-5h_IR_10Gy_Nutlin 144 bp overlap
ChIP MCF-7_nutlin GSE86164.TP53.MCF-7_nutlin 145 bp overlap
ChIP SJSA-1_nutlin GSE86164.TP53.SJSA-1_nutlin 118 bp overlap
ChIP SaOS-2 GSE15780.TP53.SaOS-2 147 bp overlap
ChIP U2OS_NUT GSE46641.TP53.U2OS_NUT 307 bp overlap
TP63 3 datasets
ChIP HaCaT_LacZ_TGFB GSE60814.TP63.HaCaT_LacZ_TGFB 139 bp overlap
ChIP HaCaT_caRAS_TGFB GSE60814.TP63.HaCaT_caRAS_TGFB 166 bp overlap
ChIP SUIT-2 GSE115461.TP63.SUIT-2 78 bp overlap
TRIM28 3 datasets
ChIP HEK293 ENCFF582MWI 322 bp overlap
ChIP HEK293 ENCFF582MWI 582 bp overlap
ChIP HEK293 ENCSR000EUZ.TRIM28.HEK293 100 bp overlap
WT1 1 dataset
ChIP HEK293 ENCSR966PJJ.WT1.HEK293 253 bp overlap
ZBED4 1 dataset
Motif DE_36h DE_36h-ZBED4_MA2328.1 10 bp overlap
ZBTB12 2 datasets
ChIP HEK293 ENCFF963HPT 141 bp overlap
ChIP HEK293 ENCSR543KOA.ZBTB12.HEK293 314 bp overlap
ZBTB24 1 dataset
ChIP HCT-116 GSE111683.ZBTB24.HCT-116 92 bp overlap
ZBTB48 3 datasets
ChIP HEK293 ENCFF809BPK 382 bp overlap
ChIP HEK293 ENCFF809BPK 465 bp overlap
ChIP HEK293 ENCSR781EQJ.ZBTB48.HEK293 588 bp overlap
ZBTB7A 1 dataset
ChIP VCaP_FBS GSE123089.ZBTB7A.VCaP_FBS 340 bp overlap
ZFP14 2 datasets
Motif DE_36h DE_36h-ZFP14_MA1972.1 15 bp overlap
Motif DE_72h DE_72h-ZFP14_MA1972.1 15 bp overlap
ZFP64 1 dataset
ChIP HEK293 GSE76494.ZFP64.HEK293 229 bp overlap
ZNF140 1 dataset
ChIP HEK293 GSE76494.ZNF140.HEK293 169 bp overlap
ZNF143 1 dataset
ChIP MCF-7 GSE76454.ZNF143.MCF-7 226 bp overlap
ZNF16 2 datasets
Motif DE_36h DE_36h-ZNF16_MA1654.2 21 bp overlap
Motif DE_72h DE_72h-ZNF16_MA1654.2 21 bp overlap
ZNF189 2 datasets
ChIP HEK293 ENCFF638TIB 115 bp overlap
ChIP HEK293 ENCSR163RYW.ZNF189.HEK293 101 bp overlap
ZNF2 1 dataset
ChIP HEK293 ENCSR011CKE.ZNF2.HEK293 282 bp overlap
ZNF263 3 datasets
ChIP HEK293 ENCFF336CWQ 491 bp overlap
ChIP HEK293 ENCFF336CWQ 591 bp overlap
ChIP HEK293 ENCFF336CWQ 433 bp overlap
ZNF35 1 dataset
ChIP HEK293 GSE76494.ZNF35.HEK293 151 bp overlap
ZNF382 2 datasets
Motif DE_60h DE_60h-ZNF382_MA1594.1 24 bp overlap
Motif DE_72h DE_72h-ZNF382_MA1594.1 24 bp overlap
ZNF467 1 dataset
ChIP HEK293 GSE76494.ZNF467.HEK293 205 bp overlap
ZNF528 1 dataset
ChIP HEK293 GSE76494.ZNF528.HEK293 235 bp overlap
ZNF596 2 datasets
ChIP HEK293 ENCFF854MGB 321 bp overlap
ChIP HEK293 ENCSR344SBD.ZNF596.HEK293 305 bp overlap
ZNF664 2 datasets
ChIP HEK293 ENCFF343XSW 169 bp overlap
ChIP HEK293 ENCSR714LZQ.ZNF664.HEK293 376 bp overlap
ZNF680 2 datasets
Motif DE_36h DE_36h-ZNF680_MA1729.2 11 bp overlap
Motif DE_72h DE_72h-ZNF680_MA1729.2 11 bp overlap
ZNF684 2 datasets
Motif DE_36h DE_36h-ZNF684_MA1600.2 14 bp overlap
Motif DE_72h DE_72h-ZNF684_MA1600.2 14 bp overlap