chr4 : 110,639,185 110,639,765
580 bp 111 TFs 1 linked gene
This 580 bp open chromatin element is linked to PITX2 and is bound by 111 transcription factors.
Linked Genes
1 gene
Gene Expression Dist. to TSS Distance Link type
PITX2 9.9 kb Proximal Proximity
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr4:110,634,185 – 110,644,765
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
111 transcription factors
Source
Cell type
AR 2 datasets
ChIP MDA-MB-453 ERP003503.AR.MDA-MB-453 167 bp overlap
ChIP prostate_fetal-fibroblast GSE90772.AR.prostate_fetal-fibroblast 212 bp overlap
ARNT::HIF1A 2 datasets
Motif DE_12h DE_12h-ARNTHIF1A_MA0259.2 5 bp overlap
Motif ES_0h ES_0h-ARNTHIF1A_MA0259.2 5 bp overlap
ASH2L 3 datasets
ChIP H1 ENCFF399KAM 538 bp overlap
ChIP H1 ENCFF399KAM 580 bp overlap
ChIP WA01 ENCSR850KIP.ASH2L.WA01 580 bp overlap
Ascl2 1 dataset
Motif DE_12h DE_12h-Ascl2_MA0816.1 10 bp overlap
BCOR 3 datasets
ChIP WA01 GSE104690.BCOR.WA01 580 bp overlap
ChIP WA01_KDM2B_KO GSE104690.BCOR.WA01_KDM2B_KO 448 bp overlap
ChIP WA01_RNF2-R GSE104690.BCOR.WA01_RNF2-R 580 bp overlap
BHLHE22 2 datasets
Motif DE_12h DE_12h-BHLHE22_MA1635.2 6 bp overlap
Motif DE_12h DE_12h-BHLHE22_MA1635.2 6 bp overlap
BRD1 1 dataset
ChIP HUES-64 GSE104059.BRD1.HUES-64 580 bp overlap
CBX2 1 dataset
ChIP K-562 ENCSR000ATU.CBX2.K-562 333 bp overlap
CBX7 4 datasets
ChIP HEK293T_PCGF2fl GSE119618.CBX7.HEK293T_PCGF2fl 224 bp overlap
ChIP hESC GSE133412.CBX7.hESC 297 bp overlap
ChIP hESC_QKO GSE133412.CBX7.hESC_QKO 287 bp overlap
ChIP hESC_TKO GSE133412.CBX7.hESC_TKO 295 bp overlap
CBX8 1 dataset
ChIP H1 ENCFF095JHA 94 bp overlap
CHD1 2 datasets
ChIP H1 ENCFF998XEK 258 bp overlap
ChIP WA01 ENCSR000AQK.CHD1.WA01 341 bp overlap
CHD7 1 dataset
ChIP WA01 ENCSR000AVA.CHD7.WA01 228 bp overlap
CTBP2 3 datasets
ChIP H1 ENCFF329MAX 567 bp overlap
ChIP H1 ENCFF329MAX 345 bp overlap
ChIP WA01 ENCSR000EUO.CTBP2.WA01 576 bp overlap
CTCF 2 datasets
Motif DE_12h DE_12h-CTCF_MA1930.2 33 bp overlap
Motif ES_0h ES_0h-CTCF_MA1930.2 33 bp overlap
E2F6 3 datasets
Motif DE_12h DE_12h-E2F6_MA0471.3 8 bp overlap
ChIP H1 ENCFF785DWK 68 bp overlap
ChIP WA01 ENCSR000BSI.E2F6.WA01 55 bp overlap
EGR1 2 datasets
ChIP H1 ENCFF451BLH 261 bp overlap
ChIP WA01 ENCSR000BJA.EGR1.WA01 154 bp overlap
ETS1 1 dataset
ChIP cardiomyocyte_D0 GSE129986.ETS1.cardiomyocyte_D0 459 bp overlap
EZH2 29 datasets
ChIP DND41 ENCSR000ASW.EZH2.DND41 352 bp overlap
ChIP GM23338 ENCFF613YON 352 bp overlap
ChIP GM23338 ENCFF613YON 61 bp overlap
ChIP H1 ENCFF232NZA 580 bp overlap
ChIP HUVEC-C GSE109625.EZH2.HUVEC-C 362 bp overlap
ChIP HUVEC-C_VEGF_12h GSE109625.EZH2.HUVEC-C_VEGF_12h 285 bp overlap
ChIP HUVEC-C_VEGF_1h GSE109625.EZH2.HUVEC-C_VEGF_1h 365 bp overlap
ChIP HUVEC-C_VEGF_4h GSE109625.EZH2.HUVEC-C_VEGF_4h 263 bp overlap
ChIP LNCaP GSE39459.EZH2.LNCaP 355 bp overlap
ChIP LNCaP-abl GSE39459.EZH2.LNCaP-abl 370 bp overlap
ChIP THP-1 GSE135024.EZH2.THP-1 569 bp overlap
ChIP astrocyte ENCFF365JTP 471 bp overlap
ChIP astrocyte ENCFF365JTP 63 bp overlap
ChIP endothelial cell of umbilical vein ENCFF539AKL 253 bp overlap
ChIP endothelial cell of umbilical vein ENCFF539AKL 544 bp overlap
ChIP fibroblast of lung ENCFF479BAW 453 bp overlap
ChIP hepatocyte ENCFF118DKH 327 bp overlap
ChIP hepatocyte ENCFF552DZB 431 bp overlap
ChIP keratinocyte ENCFF070STK 487 bp overlap
ChIP keratinocyte ENCFF070STK 277 bp overlap
ChIP keratinocyte ENCFF070STK 76 bp overlap
ChIP mammary epithelial cell ENCFF224GAI 389 bp overlap
ChIP neural ENCSR511CUH.EZH2.neural 580 bp overlap
ChIP neural cell ENCFF610EPB 401 bp overlap
ChIP neural cell ENCFF610EPB 312 bp overlap
ChIP neural progenitor cell ENCFF018MKA 576 bp overlap
ChIP neural progenitor cell ENCFF018MKA 580 bp overlap
ChIP neural progenitor cell ENCFF018MKA 120 bp overlap
ChIP neural progenitor cell ENCFF472NFV 580 bp overlap
FEZF2 1 dataset
Motif DE_12h DE_12h-FEZF2_MA2341.1 8 bp overlap
FOXL2 1 dataset
ChIP HGrC1_EV-TGF GSE138496.FOXL2.HGrC1_EV-TGF 218 bp overlap
GATA6 4 datasets
ChIP DE_D1 S09-DE-d1-GATA6-exp1 385 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 450 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 503 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 580 bp overlap
GTF3C2 1 dataset
ChIP H9 GSE94418.GTF3C2.H9 212 bp overlap
Gfi1B 4 datasets
Motif DE_12h DE_12h-Gfi1B_MA0483.2 10 bp overlap
Motif DE_36h DE_36h-Gfi1B_MA0483.2 10 bp overlap
Motif DE_60h DE_60h-Gfi1B_MA0483.2 10 bp overlap
Motif ES_0h ES_0h-Gfi1B_MA0483.2 10 bp overlap
HCFC1R1 1 dataset
ChIP cartilage GSE100311.HCFC1R1.cartilage 280 bp overlap
HDAC2 1 dataset
ChIP WA01 ENCSR000BNR.HDAC2.WA01 130 bp overlap
IRF7 1 dataset
Motif DE_12h DE_12h-IRF7_MA0772.2 13 bp overlap
JARID2 7 datasets
ChIP AMIPS6 GSE48516.JARID2.AMIPS6 287 bp overlap
ChIP EDOMIPS2 GSE48516.JARID2.EDOMIPS2 507 bp overlap
ChIP MRC-5_IPS25_MEG3 GSE48515.JARID2.MRC-5_IPS25_MEG3 580 bp overlap
ChIP UTEIPS11 GSE48516.JARID2.UTEIPS11 480 bp overlap
ChIP UTEIPS4 GSE48516.JARID2.UTEIPS4 514 bp overlap
ChIP hESC GSE133412.JARID2.hESC 262 bp overlap
ChIP hESC_TKO GSE133412.JARID2.hESC_TKO 276 bp overlap
JUN 6 datasets
ChIP DE_D1 S08-DE-d1-JUN-exp1 539 bp overlap
ChIP DE_D1 S40-DE-d1-JUN-exp2 321 bp overlap
ChIP DE_D2 S02-DE-d2-JUN-exp1 503 bp overlap
ChIP ESC S34-ESC-d0-JUN-exp2 553 bp overlap
ChIP HUES-8 GSE109524.JUN.HUES-8 580 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.JUN.HUES-8_DE_ctrl 580 bp overlap
KDM1A 1 dataset
ChIP pancreatic-progenitor_PP1 GSE104840.KDM1A.pancreatic-progenitor_PP1 289 bp overlap
KDM4A 2 datasets
ChIP H1 ENCFF078LED 580 bp overlap
ChIP WA01 ENCSR000AVC.KDM4A.WA01 283 bp overlap
KDM6B 1 dataset
ChIP CUTLL1 GSE56694.KDM6B.CUTLL1 180 bp overlap
MAX 1 dataset
ChIP H1 ENCFF914VQY 357 bp overlap
MSC 1 dataset
Motif DE_12h DE_12h-MSC_MA0665.1 10 bp overlap
MYC 1 dataset
ChIP NB69 GSE138295.MYC.NB69 222 bp overlap
MYOD1 1 dataset
ChIP IMR-90_GM GSE128527.MYOD1.IMR-90_GM 228 bp overlap
MYOG 1 dataset
Motif DE_12h DE_12h-MYOG_MA0500.3 8 bp overlap
NANOG 1 dataset
ChIP WA09 GSE105028.NANOG.WA09 211 bp overlap
NHLH1 1 dataset
Motif DE_12h DE_12h-NHLH1_MA0048.3 9 bp overlap
NIPBL 1 dataset
ChIP WA09 GSE105028.NIPBL.WA09 305 bp overlap
Neurod2 2 datasets
Motif DE_12h DE_12h-Neurod2_MA1993.2 6 bp overlap
Motif DE_12h DE_12h-Neurod2_MA1993.2 6 bp overlap
OSR2 2 datasets
Motif DE_12h DE_12h-OSR2_MA1646.2 8 bp overlap
Motif ES_0h ES_0h-OSR2_MA1646.2 8 bp overlap
Olig2 2 datasets
Motif DE_12h DE_12h-Olig2_MA1997.2 6 bp overlap
Motif DE_12h DE_12h-Olig2_MA1997.2 6 bp overlap
PATZ1 2 datasets
Motif DE_12h DE_12h-PATZ1_MA1961.2 11 bp overlap
Motif ES_0h ES_0h-PATZ1_MA1961.2 11 bp overlap
PAX6 1 dataset
ChIP retina_pigment GSE60024.PAX6.retina_pigment 166 bp overlap
PCGF2 2 datasets
ChIP HEK293T_PCGF2fl GSE119618.PCGF2.HEK293T_PCGF2fl 191 bp overlap
ChIP NT2-D1 GSE101538.PCGF2.NT2-D1 580 bp overlap
PHF19 1 dataset
ChIP DU145_SH4 GSE135623.PHF19.DU145_SH4 223 bp overlap
POLR2A 1 dataset
ChIP neural cell ENCFF604SPB 98 bp overlap
POU1F1 1 dataset
Motif DE_12h DE_12h-POU1F1_MA0784.3 14 bp overlap
POU2F1 1 dataset
Motif DE_12h DE_12h-POU2F1_MA0785.2 9 bp overlap
POU2F2 1 dataset
Motif DE_12h DE_12h-POU2F2_MA0507.3 13 bp overlap
POU2F3 1 dataset
Motif DE_12h DE_12h-POU2F3_MA0627.3 9 bp overlap
POU3F1 1 dataset
Motif DE_12h DE_12h-POU3F1_MA0786.2 10 bp overlap
POU3F2 1 dataset
Motif DE_12h DE_12h-POU3F2_MA0787.1 12 bp overlap
POU3F3 1 dataset
Motif DE_12h DE_12h-POU3F3_MA0788.1 13 bp overlap
POU3F4 1 dataset
Motif DE_12h DE_12h-POU3F4_MA0789.1 9 bp overlap
POU5F1 4 datasets
ChIP BG03 GSE21614.POU5F1.BG03 165 bp overlap
Motif DE_12h DE_12h-POU5F1_MA1115.2 7 bp overlap
ChIP DE_D1 DED1-OCT4_Batch_II 580 bp overlap
ChIP WA01_3IL ERP004238.POU5F1.WA01_3IL 461 bp overlap
POU5F1_M 1 dataset
ChIP DE_D1 DED1-OCT4-M_Batch_II 552 bp overlap
RAD21 1 dataset
ChIP WA09 GSE105028.RAD21.WA09 273 bp overlap
RBBP5 1 dataset
ChIP WA01 ENCSR000AQC.RBBP5.WA01 422 bp overlap
RBPJ 1 dataset
Motif DE_12h DE_12h-RBPJ_MA1116.2 6 bp overlap
RING1 1 dataset
ChIP SYO-1_shRING1A-B GSE139053.RING1.SYO-1_shRING1A-B 573 bp overlap
RNF2 5 datasets
ChIP H1 ENCFF239FFS 580 bp overlap
ChIP H1 ENCFF239FFS 580 bp overlap
ChIP HEK293T_PCGF135fl_OHT GSE119618.RNF2.HEK293T_PCGF135fl_OHT 159 bp overlap
ChIP HEK293T_PCGF2fl GSE119618.RNF2.HEK293T_PCGF2fl 188 bp overlap
ChIP NCCIT GSE71675.RNF2.NCCIT 366 bp overlap
RUNX1 1 dataset
ChIP Jurkat GSE85524.RUNX1.Jurkat 246 bp overlap
SALL3 1 dataset
ChIP hiPSC GSE104863.SALL3.hiPSC 487 bp overlap
SAP30 1 dataset
ChIP WA01 ENCSR000ATR.SAP30.WA01 219 bp overlap
SIN3A 1 dataset
ChIP H1 ENCFF042ZSL 356 bp overlap
SIX1 1 dataset
Motif ES_0h ES_0h-SIX1_MA1118.2 9 bp overlap
SIX2 1 dataset
Motif ES_0h ES_0h-SIX2_MA1119.2 11 bp overlap
SMAD2 3 datasets
Motif DE_12h DE_12h-SMAD2_MA1964.2 6 bp overlap
Motif ES_0h ES_0h-SMAD2_MA1964.2 6 bp overlap
ChIP hESC_YAP-_activinA_15h GSE99202.SMAD2.hESC_YAP-_activinA_15h 370 bp overlap
SMAD2-3 4 datasets
ChIP HUES-8 GSE109524.SMAD2-3.HUES-8 557 bp overlap
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 483 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.SMAD2-3.HUES-8_DE_JNKi 332 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.SMAD2-3.HUES-8_DE_ctrl 580 bp overlap
SMAD2_3 3 datasets
ChIP DE_D1 S07-DE-d1-SMAD2_3-exp1 278 bp overlap
ChIP DE_D1 S39-DE-d1-SMAD2_3-exp2 272 bp overlap
ChIP ESC S33-ESC-d0-SMAD2_3-exp2 557 bp overlap
SMAD3 1 dataset
ChIP BG03 GSE21614.SMAD3.BG03 142 bp overlap
SMARCC1 1 dataset
ChIP SYO-1_shRING1A-B GSE139053.SMARCC1.SYO-1_shRING1A-B 100 bp overlap
SMC1 1 dataset
ChIP IMR-90_G GSE118494.SMC1.IMR-90_G 139 bp overlap
SOX17_M 1 dataset
ChIP DE_D2 DED2-SOX17-M_Batch_II 232 bp overlap
SP1 3 datasets
Motif DE_12h DE_12h-SP1_MA0079.5 9 bp overlap
Motif ES_0h ES_0h-SP1_MA0079.5 9 bp overlap
ChIP WA01 ENCSR000BIR.SP1.WA01 384 bp overlap
SP2 2 datasets
Motif DE_12h DE_12h-SP2_MA0516.3 9 bp overlap
Motif ES_0h ES_0h-SP2_MA0516.3 9 bp overlap
SP5 2 datasets
Motif DE_12h DE_12h-SP5_MA1965.2 6 bp overlap
Motif ES_0h ES_0h-SP5_MA1965.2 6 bp overlap
SPIC 2 datasets
Motif DE_12h DE_12h-SPIC_MA0687.2 13 bp overlap
Motif ES_0h ES_0h-SPIC_MA0687.2 13 bp overlap
SS18 2 datasets
ChIP Aska-SS GSE108025.SS18.Aska-SS 387 bp overlap
ChIP SYO-1 GSE108025.SS18.SYO-1 364 bp overlap
SUZ12 16 datasets
ChIP GM12878 ENCSR091BOQ.SUZ12.GM12878 133 bp overlap
ChIP GM12878 ENCSR091BOQ.SUZ12.GM12878 262 bp overlap
ChIP H1 ENCFF881NFR 580 bp overlap
ChIP HEK293T_PCGF1356fl GSE119618.SUZ12.HEK293T_PCGF1356fl 114 bp overlap
ChIP HEK293T_PCGF1356fl_OHT GSE119618.SUZ12.HEK293T_PCGF1356fl_OHT 93 bp overlap
ChIP HEK293T_PCGF135fl_OHT GSE119618.SUZ12.HEK293T_PCGF135fl_OHT 244 bp overlap
ChIP HEK293T_PCGF2fl GSE119618.SUZ12.HEK293T_PCGF2fl 273 bp overlap
ChIP HEK293T_PCGF2fl_OHT GSE119618.SUZ12.HEK293T_PCGF2fl_OHT 236 bp overlap
ChIP K-562 ENCSR000AUC.SUZ12.K-562 187 bp overlap
ChIP NCCIT GSE71675.SUZ12.NCCIT 580 bp overlap
ChIP NT2/D1 ENCFF574SXS 580 bp overlap
ChIP NT2/D1 ENCFF574SXS 449 bp overlap
ChIP NT2/D1 ENCFF574SXS 242 bp overlap
ChIP WA01 ENCSR000ATS.SUZ12.WA01 580 bp overlap
ChIP WA01 ENCSR000EUQ.SUZ12.WA01 240 bp overlap
ChIP hESC_TKO GSE133412.SUZ12.hESC_TKO 225 bp overlap
Six4 1 dataset
Motif ES_0h ES_0h-Six4_MA2001.2 7 bp overlap
Stat2 2 datasets
Motif DE_12h DE_12h-Stat2_MA1623.2 10 bp overlap
Motif DE_12h DE_12h-Stat2_MA1623.2 10 bp overlap
TCF12 2 datasets
ChIP WA01 ENCSR000BIT.TCF12.WA01 54 bp overlap
ChIP WA01 ENCSR000BIT.TCF12.WA01 395 bp overlap
TCFL5 1 dataset
Motif DE_12h DE_12h-TCFL5_MA0632.3 8 bp overlap
TFAP4 1 dataset
Motif DE_12h DE_12h-TFAP4_MA0691.1 10 bp overlap
TP53 1 dataset
ChIP H9_ectoderm GSE142050.TP53.H9_ectoderm 377 bp overlap
Tcf12 2 datasets
Motif DE_12h DE_12h-Tcf12_MA0521.3 6 bp overlap
Motif DE_12h DE_12h-Tcf12_MA0521.3 6 bp overlap
Tcf21 1 dataset
Motif DE_12h DE_12h-Tcf21_MA0832.2 10 bp overlap
Twist2 2 datasets
Motif DE_12h DE_12h-Twist2_MA0633.3 6 bp overlap
Motif DE_12h DE_12h-Twist2_MA0633.3 6 bp overlap
VEZF1 2 datasets
Motif DE_12h DE_12h-VEZF1_MA1578.2 6 bp overlap
Motif ES_0h ES_0h-VEZF1_MA1578.2 6 bp overlap
YY1 1 dataset
ChIP WA01 ENCSR000BKD.YY1.WA01 188 bp overlap
ZFP14 3 datasets
Motif DE_12h DE_12h-ZFP14_MA1972.1 15 bp overlap
Motif DE_12h DE_12h-ZFP14_MA1972.1 15 bp overlap
Motif ES_0h ES_0h-ZFP14_MA1972.1 15 bp overlap
ZFP42 1 dataset
Motif DE_12h DE_12h-ZFP42_MA1651.2 13 bp overlap
ZNF189 1 dataset
Motif DE_12h DE_12h-ZNF189_MA1725.2 9 bp overlap
ZNF274 4 datasets
Motif DE_12h DE_12h-ZNF274_MA1592.2 12 bp overlap
Motif DE_36h DE_36h-ZNF274_MA1592.2 12 bp overlap
Motif DE_60h DE_60h-ZNF274_MA1592.2 12 bp overlap
Motif ES_0h ES_0h-ZNF274_MA1592.2 12 bp overlap
ZNF460 2 datasets
Motif DE_12h DE_12h-ZNF460_MA1596.1 16 bp overlap
Motif ES_0h ES_0h-ZNF460_MA1596.1 16 bp overlap
ZNF530 6 datasets
Motif DE_12h DE_12h-ZNF530_MA1981.2 14 bp overlap
Motif DE_12h DE_12h-ZNF530_MA1981.2 14 bp overlap
Motif DE_36h DE_36h-ZNF530_MA1981.2 14 bp overlap
Motif DE_60h DE_60h-ZNF530_MA1981.2 14 bp overlap
Motif ES_0h ES_0h-ZNF530_MA1981.2 14 bp overlap
Motif ES_0h ES_0h-ZNF530_MA1981.2 14 bp overlap
ZNF534 1 dataset
ChIP HEK293T GSE78099.ZNF534.HEK293T 402 bp overlap
ZNF558 1 dataset
Motif DE_12h DE_12h-ZNF558_MA2335.1 29 bp overlap
ZNF701 2 datasets
Motif DE_12h DE_12h-ZNF701_MA1987.2 17 bp overlap
Motif DE_12h DE_12h-ZNF701_MA1987.2 17 bp overlap
ZNF75A 1 dataset
Motif DE_12h DE_12h-ZNF75A_MA2097.1 12 bp overlap
ZNF75D 1 dataset
Motif DE_12h DE_12h-ZNF75D_MA1601.2 12 bp overlap
ZNF93 2 datasets
Motif DE_12h DE_12h-ZNF93_MA1721.2 14 bp overlap
Motif ES_0h ES_0h-ZNF93_MA1721.2 14 bp overlap