chr4 : 70,842,890 70,843,551
661 bp 102 TFs 1 linked gene
This 661 bp open chromatin element is linked to GRSF1 and is bound by 102 transcription factors.
Linked Genes
1 gene
Gene Expression Dist. to TSS Distance Link type
GRSF1 9.7 kb Proximal Proximity
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr4:70,837,890 – 70,848,551
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
102 transcription factors
Source
Cell type
AFF4 1 dataset
ChIP CD4_Th1_DMSO GSE62482.AFF4.CD4_Th1_DMSO 109 bp overlap
AR 1 dataset
ChIP THP-1_R1881 GSE131381.AR.THP-1_R1881 198 bp overlap
ATF7 2 datasets
ChIP GM12878 ENCFF037PYH 517 bp overlap
ChIP GM12878 ENCSR014YCR.ATF7.GM12878 328 bp overlap
BATF 1 dataset
ChIP GM12878 ENCFF954REE 231 bp overlap
BCL11A 3 datasets
ChIP CD34_Day7_30min GSE104676.BCL11A.CD34_Day7_30min 112 bp overlap
ChIP GM12878 ENCFF717YPR 271 bp overlap
ChIP GM12878 ENCSR000BHA.BCL11A.GM12878 196 bp overlap
BCL6 1 dataset
ChIP OCI-Ly1 GSE29282.BCL6.OCI-Ly1 155 bp overlap
BORCS8-MEF2B,MEF2B 1 dataset
ChIP GM12878 ENCFF427QAI 581 bp overlap
BRD4 5 datasets
ChIP CLL_patient1_4h_CpG GSE109411.BRD4.CLL_patient1_4h_CpG 227 bp overlap
ChIP CLL_patient2_4h_CpG GSE109411.BRD4.CLL_patient2_4h_CpG 661 bp overlap
ChIP CLL_patient2_4h_CpG_BET GSE109411.BRD4.CLL_patient2_4h_CpG_BET 200 bp overlap
ChIP CLL_patient3_4h_CpG GSE109411.BRD4.CLL_patient3_4h_CpG 150 bp overlap
ChIP CLL_patient4_4h_CpG GSE109411.BRD4.CLL_patient4_4h_CpG 203 bp overlap
CBFA2T3 1 dataset
ChIP K562 ENCFF673OEZ 411 bp overlap
CD74 1 dataset
ChIP CLL_p4 GSE88955.CD74.CLL_p4 218 bp overlap
DAXX 2 datasets
ChIP PC-3 GSE68647.DAXX.PC-3 129 bp overlap
ChIP PC-3_DAXX GSE68647.DAXX.PC-3_DAXX 176 bp overlap
EBF1 2 datasets
ChIP GM12878 ENCFF167CZS 321 bp overlap
ChIP GM12878 ENCFF813OXE 265 bp overlap
ELF1 3 datasets
ChIP GM12878 ENCFF692SMY 170 bp overlap
ChIP GM12878 ENCSR841NDX.ELF1.GM12878 253 bp overlap
ChIP K-562 ENCSR000BMD.ELF1.K-562 123 bp overlap
ELF3 1 dataset
ChIP PDAC GSE64557.ELF3.PDAC 288 bp overlap
ESR1 29 datasets
ChIP Ishikawa ENCSR000BIY.ESR1.Ishikawa 96 bp overlap
ChIP Ishikawa_E2 GSE132426.ESR1.Ishikawa_E2 191 bp overlap
ChIP Ishikawa_ETV4-KO1 GSE129803.ESR1.Ishikawa_ETV4-KO1 89 bp overlap
ChIP Ishikawa_ETV4-KO1_Mili GSE129803.ESR1.Ishikawa_ETV4-KO1_Mili 185 bp overlap
ChIP Ishikawa_ETV4-KO2_Mili GSE129803.ESR1.Ishikawa_ETV4-KO2_Mili 176 bp overlap
ChIP Ishikawa_siETV4-72h GSE129803.ESR1.Ishikawa_siETV4-72h 228 bp overlap
ChIP Ishikawa_siETV4-96h GSE129803.ESR1.Ishikawa_siETV4-96h 156 bp overlap
ChIP Ishikawa_siNTC-72h GSE129803.ESR1.Ishikawa_siNTC-72h 222 bp overlap
ChIP MCF-7 GSE45822.ESR1.MCF-7 158 bp overlap
ChIP MCF-7_E2 GSE59530.ESR1.MCF-7_E2 85 bp overlap
ChIP MCF-7_E2 ERP000209.ESR1.MCF-7_E2 149 bp overlap
ChIP MCF-7_E2 GSE73956.ESR1.MCF-7_E2 158 bp overlap
ChIP MCF-7_E2 GSE60270.ESR1.MCF-7_E2 247 bp overlap
ChIP MCF-7_E2-20min-ERalpha GSE94023.ESR1.MCF-7_E2-20min-ERalpha 59 bp overlap
ChIP MCF-7_E2-40min-ERalpha GSE94023.ESR1.MCF-7_E2-40min-ERalpha 171 bp overlap
ChIP MCF-7_E2-5min-ERalpha GSE94023.ESR1.MCF-7_E2-5min-ERalpha 137 bp overlap
ChIP MCF-7_E2_45min GSE109820.ESR1.MCF-7_E2_45min 87 bp overlap
ChIP MCF-7_SHCTR_E2_TNF GSE59530.ESR1.MCF-7_SHCTR_E2_TNF 131 bp overlap
ChIP MCF-7_SICTR_E2 GSE40129.ESR1.MCF-7_SICTR_E2 117 bp overlap
ChIP MCF-7_SIGATA_E2 GSE40129.ESR1.MCF-7_SIGATA_E2 112 bp overlap
ChIP MCF-7_TamR GSE128445.ESR1.MCF-7_TamR 422 bp overlap
ChIP MCF-7_estradiol_4h GSE99626.ESR1.MCF-7_estradiol_4h 87 bp overlap
ChIP MCF-7_parental GSE123284.ESR1.MCF-7_parental 128 bp overlap
ChIP MDA-MB-134-VI_E2 GSE109103.ESR1.MDA-MB-134-VI_E2 120 bp overlap
ChIP MDA-MB-134-VI_FI GSE109103.ESR1.MDA-MB-134-VI_FI 88 bp overlap
ChIP U2OS_100nM-E2 GSE151039.ESR1.U2OS_100nM-E2 233 bp overlap
ChIP U2OS_10nM-E2 GSE151039.ESR1.U2OS_10nM-E2 242 bp overlap
ChIP U2OS_10nM-E2-B GSE151039.ESR1.U2OS_10nM-E2-B 274 bp overlap
ChIP U2OS_E2 GSE26110.ESR1.U2OS_E2 150 bp overlap
ESR1_Y537N 2 datasets
ChIP MCF-7_dox GSE94493.ESR1_Y537N.MCF-7_dox 241 bp overlap
ChIP MCF-7_dox_E2 GSE94493.ESR1_Y537N.MCF-7_dox_E2 214 bp overlap
ESRRA 2 datasets
Motif DE_60h DE_60h-ESRRA_MA0592.4 9 bp overlap
ChIP SK-BR-3_EGF GSE81651.ESRRA.SK-BR-3_EGF 222 bp overlap
ESRRB 1 dataset
Motif DE_60h DE_60h-ESRRB_MA0141.4 10 bp overlap
ESRRG 1 dataset
ChIP cardiomyocyte GSE113760.ESRRG.cardiomyocyte 231 bp overlap
Elf5 1 dataset
Motif DE_60h DE_60h-Elf5_MA0136.4 8 bp overlap
Esrrg 1 dataset
Motif DE_60h DE_60h-Esrrg_MA0643.2 9 bp overlap
FOXA1 1 dataset
ChIP pancreatic-progenitor_PP1 GSE104840.FOXA1.pancreatic-progenitor_PP1 418 bp overlap
FOXA2 4 datasets
ChIP DE DE-FOXA2-1 591 bp overlap
ChIP DE DE-FOXA2-2 601 bp overlap
ChIP HepG2 ENCFF533COJ 297 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.FOXA2.pancreatic-progenitor_PP1 412 bp overlap
FOXL2 1 dataset
ChIP HGrC1_C134W-TGF GSE138496.FOXL2.HGrC1_C134W-TGF 232 bp overlap
GATA4 7 datasets
ChIP DE DE-GATA4-1 417 bp overlap
ChIP DE DE-GATA4-2 561 bp overlap
Motif DE_60h DE_60h-GATA4_MA0482.3 8 bp overlap
Motif DE_72h DE_72h-GATA4_MA0482.3 8 bp overlap
ChIP foregut GSE117136.GATA4.foregut 395 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA4.pancreatic-progenitor 661 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA4.pancreatic-progenitor_PP1 607 bp overlap
GATA6 7 datasets
ChIP AGS GSE51705.GATA6.AGS 160 bp overlap
ChIP DE DE-GATA6-1 480 bp overlap
ChIP DE DE-GATA6-2 629 bp overlap
ChIP foregut GSE117136.GATA6.foregut 525 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA6.pancreatic-progenitor 592 bp overlap
ChIP pancreatic-progenitor_KO GSE117136.GATA6.pancreatic-progenitor_KO 379 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA6.pancreatic-progenitor_PP1 570 bp overlap
HDGF 1 dataset
ChIP K-562 ENCSR563YDA.HDGF.K-562 155 bp overlap
Hoxa13 1 dataset
Motif DE_60h DE_60h-Hoxa13_MA0650.4 8 bp overlap
ID3 2 datasets
ChIP K-562 ENCSR005NMT.ID3.K-562 529 bp overlap
ChIP K562 ENCFF170RNI 481 bp overlap
IKZF1 2 datasets
ChIP GM12878 ENCFF753XDO 331 bp overlap
ChIP GM12878 ENCFF824TGK 580 bp overlap
IKZF2 5 datasets
Motif DE_60h DE_60h-IKZF2_MA2326.1 6 bp overlap
ChIP GM12878 ENCFF238LYK 586 bp overlap
ChIP GM12878 ENCFF918AID 531 bp overlap
ChIP GM12878 ENCSR680UQE.IKZF2.GM12878 526 bp overlap
ChIP GM12878 ENCSR822AHX.IKZF2.GM12878 390 bp overlap
IRF4 2 datasets
ChIP GM12878 ENCFF769ZDL 321 bp overlap
ChIP GM12878 ENCSR000BGY.IRF4.GM12878 160 bp overlap
JUN 1 dataset
ChIP MCF-7_TamR_BD610326 GSE128445.JUN.MCF-7_TamR_BD610326 189 bp overlap
JUNB 1 dataset
ChIP GM12878 ENCSR897MMC.JUNB.GM12878 193 bp overlap
KDM1A 3 datasets
ChIP pancreatic-progenitor_GT GSE104840.KDM1A.pancreatic-progenitor_GT 344 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.KDM1A.pancreatic-progenitor_PP1 504 bp overlap
ChIP pancreatic-progenitor_PP2-KDM1A-inh GSE104840.KDM1A.pancreatic-progenitor_PP2-KDM1A-inh 299 bp overlap
LCOR 1 dataset
ChIP HepG2 ENCFF499KCU 391 bp overlap
MAF 2 datasets
ChIP CD4_Th1 GSE72266.MAF.CD4_Th1 300 bp overlap
Motif DE_60h DE_60h-MAF_MA1520.2 13 bp overlap
MAZ 2 datasets
ChIP GM12878 ENCFF453CES 457 bp overlap
ChIP GM12878 ENCFF453CES 258 bp overlap
MED1 1 dataset
ChIP CD4_Th1_DMSO GSE62482.MED1.CD4_Th1_DMSO 111 bp overlap
MEF2B 1 dataset
ChIP tonsil GSE110682.MEF2B.tonsil 194 bp overlap
MLLT1 1 dataset
ChIP GM12878 ENCSR552XSN.MLLT1.GM12878 446 bp overlap
MTA2 2 datasets
ChIP GM12878 ENCFF615CWQ 571 bp overlap
ChIP GM12878 ENCSR293QAR.MTA2.GM12878 536 bp overlap
MTA3 1 dataset
ChIP GM12878 ENCSR000BRH.MTA3.GM12878 171 bp overlap
Mafg 1 dataset
Motif DE_60h DE_60h-Mafg_MA0659.4 12 bp overlap
NEUROD1 1 dataset
ChIP D341-Med_shNEUROD1-1579 GSE92582.NEUROD1.D341-Med_shNEUROD1-1579 299 bp overlap
NFIC 1 dataset
ChIP GM12878 ENCSR000BRN.NFIC.GM12878 241 bp overlap
NFKB1 1 dataset
ChIP L1236 GSE63736.NFKB1.L1236 211 bp overlap
NFKB2 1 dataset
ChIP L1236 GSE63736.NFKB2.L1236 252 bp overlap
NIPBL 1 dataset
ChIP LCL GSE38395.NIPBL.LCL 155 bp overlap
NR2C1 2 datasets
Motif DE_60h DE_60h-NR2C1_MA1535.2 6 bp overlap
ChIP GM12878 ENCSR784VIQ.NR2C1.GM12878 497 bp overlap
NR2C2 1 dataset
Motif DE_60h DE_60h-NR2C2_MA1536.2 6 bp overlap
NR2F1 5 datasets
Motif DE_60h DE_60h-NR2F1_MA0017.3 12 bp overlap
ChIP GM12878 ENCFF273VKX 378 bp overlap
ChIP GM12878 ENCSR514VYD.NR2F1.GM12878 566 bp overlap
ChIP K-562 ENCSR970NKQ.NR2F1.K-562 571 bp overlap
ChIP K562 ENCFF221HJH 175 bp overlap
NR2F2 2 datasets
Motif DE_60h DE_60h-NR2F2_MA1111.2 7 bp overlap
ChIP K-562 ENCSR000BRS.NR2F2.K-562 138 bp overlap
NR2F6 2 datasets
ChIP Hep-G2 ENCSR518WPL.NR2F6.Hep-G2 277 bp overlap
ChIP K-562 ENCSR707QWA.NR2F6.K-562 199 bp overlap
NR4A1 2 datasets
Motif DE_60h DE_60h-NR4A1_MA1112.3 8 bp overlap
ChIP Kasumi-1 GSE79491.NR4A1.Kasumi-1 184 bp overlap
NR4A2 1 dataset
Motif DE_60h DE_60h-NR4A2_MA0160.3 8 bp overlap
NRIP1 1 dataset
ChIP MCF-7_E2 ERP005838.NRIP1.MCF-7_E2 118 bp overlap
Nr1H2 1 dataset
Motif DE_60h DE_60h-Nr1H2_MA1996.2 6 bp overlap
Nr1H4 1 dataset
Motif DE_60h DE_60h-Nr1H4_MA1110.3 6 bp overlap
Nr1h3 1 dataset
Motif DE_60h DE_60h-Nr1h3_MA2337.1 6 bp overlap
Nr2e1 1 dataset
Motif DE_60h DE_60h-Nr2e1_MA0676.1 9 bp overlap
PAX5 2 datasets
ChIP GM12878 ENCSR000BHJ.PAX5.GM12878 199 bp overlap
ChIP GM12878 ENCSR000BHD.PAX5.GM12878 107 bp overlap
PDX1 2 datasets
ChIP hiPSC GSE125768.PDX1.hiPSC 340 bp overlap
ChIP hiPSC_derived_PPs GSE106949.PDX1.hiPSC_derived_PPs 331 bp overlap
PKNOX1 1 dataset
ChIP GM12878 ENCFF589FCY 228 bp overlap
POLR2A 1 dataset
ChIP Raji ENCFF613VGX 351 bp overlap
Ppara 1 dataset
Motif DE_60h DE_60h-Ppara_MA2338.1 7 bp overlap
REL 1 dataset
Motif DE_60h DE_60h-REL_MA0101.1 10 bp overlap
RELA 4 datasets
ChIP CD4_Th1_DMSO GSE62482.RELA.CD4_Th1_DMSO 112 bp overlap
ChIP GM12878 ENCSR664POU.RELA.GM12878 228 bp overlap
ChIP GM12878 ENCSR000EAG.RELA.GM12878 218 bp overlap
ChIP GM12891 ENCSR000EAI.RELA.GM12891 267 bp overlap
RELB 3 datasets
ChIP GM12878 ENCFF217ADF 180 bp overlap
ChIP GM12878 ENCSR387QUV.RELB.GM12878 586 bp overlap
ChIP L1236 GSE63736.RELB.L1236 88 bp overlap
REST 2 datasets
ChIP K-562 ENCSR137ZMQ.REST.K-562 203 bp overlap
ChIP K562 ENCFF688UKW 411 bp overlap
RORA 1 dataset
Motif DE_60h DE_60h-RORA_MA0071.1 10 bp overlap
RORB 1 dataset
Motif DE_60h DE_60h-RORB_MA1150.2 10 bp overlap
RUNX3 1 dataset
ChIP GM12878 ENCFF395WHA 371 bp overlap
RXR 1 dataset
ChIP pancreatic-progenitor_PP1 GSE104840.RXR.pancreatic-progenitor_PP1 178 bp overlap
SKIL 2 datasets
ChIP GM12878 ENCFF171OVM 551 bp overlap
ChIP GM12878 ENCSR212YKD.SKIL.GM12878 437 bp overlap
SMAD2-3 2 datasets
ChIP HGrC1_C134W-TGF GSE138496.SMAD2-3.HGrC1_C134W-TGF 138 bp overlap
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 355 bp overlap
SMAD3 1 dataset
ChIP NCI-H441_SIITTF1 GSE51509.SMAD3.NCI-H441_SIITTF1 180 bp overlap
SOX10 1 dataset
Motif DE_60h DE_60h-SOX10_MA0442.3 6 bp overlap
SP1 1 dataset
ChIP GM12878 ENCSR000BHK.SP1.GM12878 174 bp overlap
SPI1 5 datasets
ChIP DC_96h_donorM GSE128834.SPI1.DC_96h_donorM 226 bp overlap
ChIP GM12878 ENCFF134LCP 78 bp overlap
ChIP GM12891 ENCFF563IUT 241 bp overlap
ChIP GM12891 ENCSR000BIJ.SPI1.GM12891 168 bp overlap
ChIP K-562_NABUT GSE74999.SPI1.K-562_NABUT 133 bp overlap
SREBP2 1 dataset
ChIP monocyte_TNF GSE129202.SREBP2.monocyte_TNF 271 bp overlap
SRF 2 datasets
ChIP GM12878 ENCFF878IIX 397 bp overlap
ChIP GM12878 ENCSR041XML.SRF.GM12878 251 bp overlap
STAT3 1 dataset
ChIP B-cell GSE123398.STAT3.B-cell 372 bp overlap
Stat6 1 dataset
Motif DE_60h DE_60h-Stat6_MA0520.2 10 bp overlap
TBX21 2 datasets
ChIP GM12878 ENCFF951HUW 485 bp overlap
ChIP GM12878 ENCSR739IHN.TBX21.GM12878 523 bp overlap
TCF12 3 datasets
ChIP GM12878 ENCSR000BGZ.TCF12.GM12878 155 bp overlap
ChIP K-562 ENCSR744WOO.TCF12.K-562 214 bp overlap
ChIP K562 ENCFF931DJY 391 bp overlap
TCF3 2 datasets
ChIP GM12878 ENCSR000BQT.TCF3.GM12878 111 bp overlap
ChIP K-562 ENCSR970OJY.TCF3.K-562 166 bp overlap
TCF7L1 1 dataset
Motif DE_60h DE_60h-TCF7L1_MA1421.1 12 bp overlap
THRB 1 dataset
Motif DE_60h DE_60h-THRB_MA1575.2 17 bp overlap
Tfcp2l1 2 datasets
Motif DE_60h DE_60h-Tfcp2l1_MA0145.2 14 bp overlap
Motif DE_72h DE_72h-Tfcp2l1_MA0145.2 14 bp overlap
ZEB1 1 dataset
ChIP GM12878 ENCSR000BND.ZEB1.GM12878 105 bp overlap
ZFP36 1 dataset
ChIP GM12878 ENCSR900XDB.ZFP36.GM12878 186 bp overlap
ZNF597 1 dataset
ChIP GM12878 GSE97661.ZNF597.GM12878 139 bp overlap