chr4 : 56,787,104 56,787,870
766 bp 101 TFs 3 linked genes
This 766 bp open chromatin element is linked to REST, NOA1, and POLR2B and is bound by 101 transcription factors.
Linked Genes
3 genes
Gene Expression Dist. to TSS Distance Link type
REST 120.4 kb Distal Multiome
NOA1 190.2 kb Distal Multiome
POLR2B 190.3 kb Distal Multiome
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr4:56,782,104 – 56,792,870
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
101 transcription factors
Source
Cell type
AR 4 datasets
ChIP fibroblast_prostate-cancer_PCDF2 GSE126852.AR.fibroblast_prostate-cancer_PCDF2 221 bp overlap
ChIP prostate GSE56288.AR.prostate 256 bp overlap
ChIP prostate_cancer-associated-fibroblasts GSE90772.AR.prostate_cancer-associated-fibroblasts 289 bp overlap
ChIP prostate_fetal-fibroblast GSE90772.AR.prostate_fetal-fibroblast 225 bp overlap
ASCL1 1 dataset
ChIP NCI-H82 GSE69394.ASCL1.NCI-H82 162 bp overlap
BCOR 1 dataset
ChIP WA01_RNF2-R GSE104690.BCOR.WA01_RNF2-R 324 bp overlap
BRD2 1 dataset
ChIP SUM159PT_MGSK2801 GSE116879.BRD2.SUM159PT_MGSK2801 232 bp overlap
BRD4 13 datasets
ChIP HEK293T GSE51633.BRD4.HEK293T 135 bp overlap
ChIP IMR-90_QUIES GSE74238.BRD4.IMR-90_QUIES 84 bp overlap
ChIP K-562_DMSO GSE99178.BRD4.K-562_DMSO 190 bp overlap
ChIP Kelly_resistant_JQ1 GSE103030.BRD4.Kelly_resistant_JQ1 224 bp overlap
ChIP Kelly_sensitive GSE103030.BRD4.Kelly_sensitive 502 bp overlap
ChIP MDA-MB-231_JQ1_MGSK2801 GSE116879.BRD4.MDA-MB-231_JQ1_MGSK2801 232 bp overlap
ChIP MDA-MB-231_JQ1_MGSK2801_rDNA GSE116879.BRD4.MDA-MB-231_JQ1_MGSK2801_rDNA 232 bp overlap
ChIP MPNST_PDJQ GSE62499.BRD4.MPNST_PDJQ 112 bp overlap
ChIP SUM159PT_DMSO_24h GSE87418.BRD4.SUM159PT_DMSO_24h 228 bp overlap
ChIP SUM159PT_DMSO_8h GSE87418.BRD4.SUM159PT_DMSO_8h 284 bp overlap
ChIP SUM159PT_Dox_48h GSE87418.BRD4.SUM159PT_Dox_48h 137 bp overlap
ChIP SUM159PT_scramble GSE131097.BRD4.SUM159PT_scramble 598 bp overlap
ChIP retina_AB1-RB GSE86981.BRD4.retina_AB1-RB 244 bp overlap
CBFA2T3 2 datasets
ChIP K-562 ENCSR697YLJ.CBFA2T3.K-562 345 bp overlap
ChIP K562 ENCFF673OEZ 411 bp overlap
CDK9 1 dataset
ChIP HEK293T_SICTR GSE51633.CDK9.HEK293T_SICTR 155 bp overlap
CDX2 4 datasets
ChIP Caco-2_DIFF GSE23436.CDX2.Caco-2_DIFF 139 bp overlap
ChIP LS180 GSE31939.CDX2.LS180 210 bp overlap
ChIP LS180_125 GSE31939.CDX2.LS180_125 249 bp overlap
ChIP adult-duodenal-cell GSE115314.CDX2.adult-duodenal-cell 244 bp overlap
CHD7 1 dataset
ChIP hiPSC_derived_neural-crest-cell GSE108506.CHD7.hiPSC_derived_neural-crest-cell 176 bp overlap
CTBP1 1 dataset
ChIP K-562 ENCSR201NQZ.CTBP1.K-562 490 bp overlap
CTCF 3 datasets
ChIP Kasumi-1_ctrl GSE121280.CTCF.Kasumi-1_ctrl 123 bp overlap
ChIP THP-1_macrophage_PMA GSE96800.CTCF.THP-1_macrophage_PMA 220 bp overlap
ChIP anterior-temporal-cortex_adult GSE116825.CTCF.anterior-temporal-cortex_adult 353 bp overlap
Cebpa 1 dataset
ChIP BLaER1 ENCFF262VBH 251 bp overlap
E2F1 1 dataset
ChIP U-87MG_GBM GSE99171.E2F1.U-87MG_GBM 210 bp overlap
E2F7 1 dataset
ChIP IMR-90_SENES_SHCTR GSE40343.E2F7.IMR-90_SENES_SHCTR 52 bp overlap
EGR1 1 dataset
ChIP T-HESCs GSE141063.EGR1.T-HESCs 226 bp overlap
ELK1 1 dataset
ChIP K-562 ENCSR338QAC.ELK1.K-562 199 bp overlap
EP300 1 dataset
ChIP SK-N-SH ENCSR000BUA.EP300.SK-N-SH 57 bp overlap
ERG 7 datasets
ChIP K-562_DOX GSE23730.ERG.K-562_DOX 270 bp overlap
ChIP SKNO-1 GSE23730.ERG.SKNO-1 402 bp overlap
ChIP VCaP GSE83650.ERG.VCaP 168 bp overlap
ChIP VCaP GSE98809.ERG.VCaP 168 bp overlap
ChIP VCaP_DHAT_18H GSE28950.ERG.VCaP_DHAT_18H 135 bp overlap
ChIP VCaP_DHT GSE79128.ERG.VCaP_DHT 420 bp overlap
ChIP VCaP_shARID1A GSE110655.ERG.VCaP_shARID1A 153 bp overlap
ESRRG 1 dataset
ChIP cardiomyocyte GSE113760.ESRRG.cardiomyocyte 205 bp overlap
ETV1 4 datasets
ChIP GIST-T1 GSE106624.ETV1.GIST-T1 312 bp overlap
ChIP GIST-T1 GSE80443.ETV1.GIST-T1 184 bp overlap
ChIP GIST-T1 GSE106624.ETV1.GIST-T1 365 bp overlap
ChIP GIST-T1 GSE80443.ETV1.GIST-T1 168 bp overlap
FEZF1 2 datasets
ChIP HEK293 ENCFF528YED 376 bp overlap
ChIP HEK293 ENCSR827NWO.FEZF1.HEK293 302 bp overlap
FLI1 1 dataset
ChIP SKNO-1 GSE23730.FLI1.SKNO-1 533 bp overlap
FOS 3 datasets
ChIP IMR-90 ENCFF179EDA 128 bp overlap
ChIP IMR-90 ENCSR124AIG.FOS.IMR-90 79 bp overlap
ChIP MCF-10A ENCSR000DON.FOS.MCF-10A 67 bp overlap
FOSL2 4 datasets
ChIP NPC_K755R-pos GSE122631.FOSL2.NPC_K755R-pos 111 bp overlap
ChIP SK-N-SH ENCFF127ZDW 95 bp overlap
ChIP SK-N-SH ENCSR000BVB.FOSL2.SK-N-SH 58 bp overlap
ChIP hESC GSE69539.FOSL2.hESC 75 bp overlap
FOXA2 2 datasets
ChIP DE DE-FOXA2-1 327 bp overlap
ChIP DE DE-FOXA2-2 294 bp overlap
FOXF1 2 datasets
ChIP GIST-T1 GSE106624.FOXF1.GIST-T1 185 bp overlap
ChIP GIST-T1 GSE106624.FOXF1.GIST-T1 147 bp overlap
FOXL2 2 datasets
ChIP HGrC1_C134W-TGF GSE138496.FOXL2.HGrC1_C134W-TGF 620 bp overlap
ChIP HGrC1_C134W-TGF_parental GSE138496.FOXL2.HGrC1_C134W-TGF_parental 274 bp overlap
GABPA 1 dataset
ChIP K562 ENCFF139LXS 674 bp overlap
GATA2 6 datasets
ChIP ME-1_CBFB-MYH11-KD GSE117138.GATA2.ME-1_CBFB-MYH11-KD 283 bp overlap
ChIP SKH1 GSE87283.GATA2.SKH1 217 bp overlap
ChIP TF1 GSE73207.GATA2.TF1 567 bp overlap
ChIP dermal-fibroblast_alone GSE51025.GATA2.dermal-fibroblast_alone 122 bp overlap
ChIP hiPSC_SLC6ebs GSE107639.GATA2.hiPSC_SLC6ebs 396 bp overlap
ChIP hiPSC_SLC9ebs GSE107639.GATA2.hiPSC_SLC9ebs 189 bp overlap
GATA4 5 datasets
ChIP DE DE-GATA4-1 524 bp overlap
ChIP DE DE-GATA4-2 582 bp overlap
ChIP foregut GSE117136.GATA4.foregut 317 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA4.pancreatic-progenitor 530 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA4.pancreatic-progenitor_PP1 604 bp overlap
GATA6 14 datasets
ChIP DE DE-GATA6-1 614 bp overlap
ChIP DE DE-GATA6-2 599 bp overlap
ChIP DE_D1 S09-DE-d1-GATA6-exp1 514 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 452 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 518 bp overlap
ChIP ESO-26 GSE132680.GATA6.ESO-26 723 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 500 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 588 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 515 bp overlap
ChIP foregut GSE117136.GATA6.foregut 414 bp overlap
ChIP foregut_KO GSE117136.GATA6.foregut_KO 352 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA6.pancreatic-progenitor 265 bp overlap
ChIP pancreatic-progenitor_KO GSE117136.GATA6.pancreatic-progenitor_KO 295 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA6.pancreatic-progenitor_PP1 271 bp overlap
GLIS1 2 datasets
ChIP HEK293 ENCFF299RSE 268 bp overlap
ChIP HEK293 ENCSR482BBZ.GLIS1.HEK293 532 bp overlap
HAND2 1 dataset
ChIP Kelly GSE94822.HAND2.Kelly 235 bp overlap
HDAC1 1 dataset
ChIP K-562 GSE140325.HDAC1.K-562 229 bp overlap
HDAC2 2 datasets
ChIP K-562 ENCSR000BMG.HDAC2.K-562 197 bp overlap
ChIP RH4_Entinostat-6H GSE116344.HDAC2.RH4_Entinostat-6H 318 bp overlap
HDAC3 1 dataset
ChIP RH4_DMSO-6H GSE116344.HDAC3.RH4_DMSO-6H 414 bp overlap
HNF4A 1 dataset
ChIP Caco-2_DIFF GSE23436.HNF4A.Caco-2_DIFF 116 bp overlap
IKZF1 3 datasets
ChIP K-562 ENCSR395HWC.IKZF1.K-562 639 bp overlap
ChIP K562 ENCFF348IBL 456 bp overlap
ChIP K562 ENCFF771OHZ 497 bp overlap
IKZF3 1 dataset
ChIP HEK293 ENCSR304AMN.IKZF3.HEK293 254 bp overlap
KLF5 1 dataset
ChIP ESO-26 GSE132680.KLF5.ESO-26 285 bp overlap
MAZ 2 datasets
ChIP HEK293 GSE76494.MAZ.HEK293 91 bp overlap
ChIP IMR-90 ENCSR000EFF.MAZ.IMR-90 209 bp overlap
MED1 1 dataset
ChIP HCT-116_shLuc_thaps GSE121798.MED1.HCT-116_shLuc_thaps 264 bp overlap
MED12 2 datasets
ChIP leiomyoma_PT886 GSE128230.MED12.leiomyoma_PT886 128 bp overlap
ChIP leiomyoma_PT967 GSE128230.MED12.leiomyoma_PT967 58 bp overlap
MYC 1 dataset
ChIP GP5D GSE51234.MYC.GP5D 445 bp overlap
MYCN 1 dataset
ChIP Kelly GSE94822.MYCN.Kelly 214 bp overlap
MYOD1 4 datasets
ChIP IMR-90_GM GSE128527.MYOD1.IMR-90_GM 382 bp overlap
ChIP RH4 GSE83726.MYOD1.RH4 297 bp overlap
ChIP myoblast GSE50413.MYOD1.myoblast 144 bp overlap
ChIP rhabdomyosarcoma GSE50413.MYOD1.rhabdomyosarcoma 65 bp overlap
NANOG 1 dataset
ChIP HUES-8 GSE109524.NANOG.HUES-8 303 bp overlap
NCOA2 1 dataset
ChIP MCF-7 ERP000901.NCOA2.MCF-7 163 bp overlap
NCOR1 1 dataset
ChIP K-562 ENCSR798ILC.NCOR1.K-562 285 bp overlap
NEUROD1 1 dataset
ChIP D341-Med_shGFP GSE92582.NEUROD1.D341-Med_shGFP 550 bp overlap
NEUROG2 7 datasets
ChIP MRC-5_NFD_02DPT GSE75910.NEUROG2.MRC-5_NFD_02DPT 196 bp overlap
ChIP MRC-5_NFD_02DPT GSE75910.NEUROG2.MRC-5_NFD_02DPT 207 bp overlap
ChIP MRC-5_NFD_05DPT GSE75910.NEUROG2.MRC-5_NFD_05DPT 279 bp overlap
ChIP MRC-5_NFD_1DPT GSE75910.NEUROG2.MRC-5_NFD_1DPT 226 bp overlap
ChIP MRC-5_N_02DPT GSE75910.NEUROG2.MRC-5_N_02DPT 254 bp overlap
ChIP MRC-5_N_05DPT GSE75910.NEUROG2.MRC-5_N_05DPT 306 bp overlap
ChIP MRC-5_N_1DPT GSE75910.NEUROG2.MRC-5_N_1DPT 498 bp overlap
NIPBL 1 dataset
ChIP GP5D GSE51234.NIPBL.GP5D 530 bp overlap
NKX2-1 1 dataset
ChIP NCI-H3122 GSE39998.NKX2-1.NCI-H3122 72 bp overlap
NKX3-1 1 dataset
ChIP islet ERP004003.NKX3-1.islet 184 bp overlap
NR2F2 5 datasets
ChIP K-562 ENCSR000BRS.NR2F2.K-562 215 bp overlap
ChIP liver ENCFF427MRU 421 bp overlap
ChIP liver ENCFF565JGD 491 bp overlap
ChIP liver ENCSR168SMX.NR2F2.liver 669 bp overlap
ChIP liver ENCSR338MMB.NR2F2.liver 316 bp overlap
NR2F6 1 dataset
ChIP K-562 ENCSR707QWA.NR2F6.K-562 149 bp overlap
NR3C1 2 datasets
ChIP IMR-90 ERP007093.NR3C1.IMR-90 64 bp overlap
ChIP MDA-MB-231 GSE152203.NR3C1.MDA-MB-231 53 bp overlap
OSR2 1 dataset
ChIP HEK293 GSE76494.OSR2.HEK293 204 bp overlap
PGR 2 datasets
ChIP myometrium_NP1 GSE137550.PGR.myometrium_NP1 406 bp overlap
ChIP myometrium_NP2 GSE137550.PGR.myometrium_NP2 341 bp overlap
POLR2A 3 datasets
ChIP sigmoid colon ENCFF748YVT 415 bp overlap
ChIP transverse colon ENCFF607LKE 214 bp overlap
ChIP upper lobe of left lung ENCFF199JUI 227 bp overlap
POU5F1 1 dataset
ChIP hESC_NAIVE GSE69646.POU5F1.hESC_NAIVE 267 bp overlap
PRDM6 2 datasets
ChIP HEK293 ENCFF283AJL 142 bp overlap
ChIP HEK293 ENCSR892QHR.PRDM6.HEK293 222 bp overlap
PSIP1 1 dataset
ChIP T-REx-293 GSE117155.PSIP1.T-REx-293 417 bp overlap
RAD21 2 datasets
ChIP GP5D GSE51234.RAD21.GP5D 315 bp overlap
ChIP IMR-90 ENCSR000EFJ.RAD21.IMR-90 61 bp overlap
RARA 1 dataset
ChIP hiPSC_D3 GSE109172.RARA.hiPSC_D3 400 bp overlap
RCOR1 2 datasets
ChIP IMR-90 ENCFF644MZN 132 bp overlap
ChIP IMR-90 ENCSR000EFG.RCOR1.IMR-90 70 bp overlap
REST 6 datasets
ChIP K-562 ENCSR000BMW.REST.K-562 166 bp overlap
ChIP PFSK1 ENCSR000BOX.REST.PFSK1 89 bp overlap
ChIP colorectal-cancer_CRC121_dissociated GSE112555.REST.colorectal-cancer_CRC121_dissociated 183 bp overlap
ChIP colorectal-cancer_shCDH1 GSE112555.REST.colorectal-cancer_shCDH1 495 bp overlap
ChIP liver ENCFF240FWT 465 bp overlap
ChIP liver ENCSR867WPH.REST.liver 183 bp overlap
SIN3A 1 dataset
ChIP hiPSC_IB12 GSE106870.SIN3A.hiPSC_IB12 137 bp overlap
SKIL 1 dataset
ChIP K-562 ENCSR336DXE.SKIL.K-562 298 bp overlap
SMAD2-3 3 datasets
ChIP HGrC1_C134W-TGF GSE138496.SMAD2-3.HGrC1_C134W-TGF 123 bp overlap
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 506 bp overlap
ChIP KGN_TGF GSE138496.SMAD2-3.KGN_TGF 208 bp overlap
SMAD4 1 dataset
ChIP HGrC1_C134W-TGF GSE138496.SMAD4.HGrC1_C134W-TGF 300 bp overlap
SMARCA2 1 dataset
ChIP NPC_K755R-pos GSE122631.SMARCA2.NPC_K755R-pos 110 bp overlap
SMARCA4 5 datasets
ChIP TTC-1240 GSE124903.SMARCA4.TTC-1240 766 bp overlap
ChIP TTC-1240_R377H GSE124903.SMARCA4.TTC-1240_R377H 746 bp overlap
ChIP TTC-1240_SMARCB1-FL GSE124903.SMARCA4.TTC-1240_SMARCB1-FL 532 bp overlap
ChIP TTC-1240_SMARCB1-FL GSE124903.SMARCA4.TTC-1240_SMARCB1-FL 136 bp overlap
ChIP TTC-1240_delC GSE124903.SMARCA4.TTC-1240_delC 766 bp overlap
SMARCB1 2 datasets
ChIP TTC-1240_R377H GSE124903.SMARCB1.TTC-1240_R377H 216 bp overlap
ChIP proliferating-human-fibroblast_ASO_Ctrl GSE131711.SMARCB1.proliferating-human-fibroblast_ASO_Ctrl 335 bp overlap
SMARCC1 4 datasets
ChIP TTC-1240 GSE124903.SMARCC1.TTC-1240 699 bp overlap
ChIP TTC-1240_R377H GSE124903.SMARCC1.TTC-1240_R377H 690 bp overlap
ChIP TTC-1240_SMARCB1-FL GSE124903.SMARCC1.TTC-1240_SMARCB1-FL 534 bp overlap
ChIP TTC-1240_delC GSE124903.SMARCC1.TTC-1240_delC 766 bp overlap
SMC3 3 datasets
ChIP GP5D GSE51234.SMC3.GP5D 438 bp overlap
ChIP IMR-90 ENCFF627LON 91 bp overlap
ChIP IMR-90 ENCSR000HPG.SMC3.IMR-90 64 bp overlap
SOX17 1 dataset
ChIP DE_D2 DED2-SOX17_Batch_II 527 bp overlap
SOX6 1 dataset
ChIP K-562 ENCSR788RSW.SOX6.K-562 332 bp overlap
SP7 1 dataset
ChIP HEK293 ENCSR468IJT.SP7.HEK293 356 bp overlap
SPI1 1 dataset
ChIP K-562_NABUT GSE74999.SPI1.K-562_NABUT 182 bp overlap
SS18 1 dataset
ChIP SYO-1_shSSX GSE108025.SS18.SYO-1_shSSX 100 bp overlap
SS18-SSX 1 dataset
ChIP fibroblast_23aa-SSX-Tail GSE139053.SS18-SSX.fibroblast_23aa-SSX-Tail 63 bp overlap
STAT3 2 datasets
ChIP MCF-10A ENCSR000DOQ.STAT3.MCF-10A 101 bp overlap
ChIP MCF10A-Er-Src_TAM GSE115597.STAT3.MCF10A-Er-Src_TAM 103 bp overlap
TAL1 3 datasets
ChIP CHRF28811 ERP008568.TAL1.CHRF28811 281 bp overlap
ChIP K-562 GSE107726.TAL1.K-562 572 bp overlap
ChIP K-562 ENCSR106FRG.TAL1.K-562 278 bp overlap
TBX2 1 dataset
ChIP Kelly GSE94822.TBX2.Kelly 164 bp overlap
TCF21 1 dataset
ChIP HCASMC GSE124011.TCF21.HCASMC 121 bp overlap
TCF3 2 datasets
ChIP K-562 ENCSR970OJY.TCF3.K-562 250 bp overlap
ChIP K562 ENCFF319QZT 381 bp overlap
TEAD1 1 dataset
ChIP HEK293_siE1A GSE130135.TEAD1.HEK293_siE1A 114 bp overlap
TFAP4 1 dataset
ChIP DLD-1 GSE46935.TFAP4.DLD-1 475 bp overlap
VEZF1 1 dataset
ChIP K562 ENCFF053XDV 438 bp overlap
ZBTB24 1 dataset
ChIP HCT-116 GSE111683.ZBTB24.HCT-116 154 bp overlap
ZBTB42 1 dataset
ChIP HEK293 GSE76494.ZBTB42.HEK293 187 bp overlap
ZIC2 1 dataset
ChIP HEK293 ENCFF033NQQ 459 bp overlap
ZNF711 1 dataset
ChIP HEK293T GSE145160.ZNF711.HEK293T 514 bp overlap
ZNF778 1 dataset
ChIP HEK293 GSE76494.ZNF778.HEK293 157 bp overlap