chr11 : 11,691,525 11,692,376
851 bp 109 TFs 2 linked genes
This 851 bp open chromatin element is linked to GALNT18 and USP47 and is bound by 109 transcription factors.
Linked Genes
2 genes
Gene Expression Dist. to TSS Distance Link type
GALNT18 70.0 kb Distal Multiome
USP47 150.0 kb Distal Multiome
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr11:11,686,525 – 11,697,376
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
109 transcription factors
Source
Cell type
AR 21 datasets
ChIP LNCaP ERP001226.AR.LNCaP 104 bp overlap
ChIP LNCaP GSE110655.AR.LNCaP 59 bp overlap
ChIP LNCaP-95_NT GSE125552.AR.LNCaP-95_NT 91 bp overlap
ChIP LNCaP-clone-FGC_DHT-ABBV-744 GSE118247.AR.LNCaP-clone-FGC_DHT-ABBV-744 75 bp overlap
ChIP LNCaP-clone-FGC_DHT-DMSO GSE118247.AR.LNCaP-clone-FGC_DHT-DMSO 123 bp overlap
ChIP LNCaP_1F5_SIFOXA1 GSE30623.AR.LNCaP_1F5_SIFOXA1 52 bp overlap
ChIP LNCaP_ETOH GSE69043.AR.LNCaP_ETOH 106 bp overlap
ChIP LNCaP_FOXA1 ERP003503.AR.LNCaP_FOXA1 81 bp overlap
ChIP LNCaP_M253K_shFOXA1_Ethanol GSE128883.AR.LNCaP_M253K_shFOXA1_Ethanol 114 bp overlap
ChIP LNCaP_R1881 GSE69043.AR.LNCaP_R1881 105 bp overlap
ChIP LNCaP_SHGATA2_ETOH GSE69043.AR.LNCaP_SHGATA2_ETOH 64 bp overlap
ChIP LNCaP_SHGATA2_R1881 GSE69043.AR.LNCaP_SHGATA2_R1881 103 bp overlap
ChIP LNCaP_Talen_DHT GSE89938.AR.LNCaP_Talen_DHT 80 bp overlap
ChIP LNCaP_androgen-N_hypoxia-N GSE114732.AR.LNCaP_androgen-N_hypoxia-N 67 bp overlap
ChIP LNCaP_androgen-N_hypoxia-Y GSE114732.AR.LNCaP_androgen-N_hypoxia-Y 72 bp overlap
ChIP LNCaP_sgCHD1 GSE117430.AR.LNCaP_sgCHD1 103 bp overlap
ChIP LNCaP_sgCtrl GSE117430.AR.LNCaP_sgCtrl 86 bp overlap
ChIP prostate-cancer GSE136128.AR.prostate-cancer 61 bp overlap
ChIP prostate-cancer_C4-2-CON GSE136128.AR.prostate-cancer_C4-2-CON 89 bp overlap
ChIP prostate-cancer_PDX_167 GSE130408.AR.prostate-cancer_PDX_167 75 bp overlap
ChIP prostate-cancer_PDX_78 GSE130408.AR.prostate-cancer_PDX_78 66 bp overlap
ASH2L 1 dataset
ChIP H1 ENCFF399KAM 394 bp overlap
Ascl2 2 datasets
Motif DE_48h DE_48h-Ascl2_MA0816.1 10 bp overlap
Motif DE_72h DE_72h-Ascl2_MA0816.1 10 bp overlap
BAP1 2 datasets
ChIP NCI-H1963_shASXL3 GSE145028.BAP1.NCI-H1963_shASXL3 313 bp overlap
ChIP NCI-H1963_shNONT GSE145028.BAP1.NCI-H1963_shNONT 94 bp overlap
BHLHE22 2 datasets
Motif DE_48h DE_48h-BHLHE22_MA1635.2 6 bp overlap
Motif DE_72h DE_72h-BHLHE22_MA1635.2 6 bp overlap
BRD4 5 datasets
ChIP LNCaP-C4-2_F133V GSE88871.BRD4.LNCaP-C4-2_F133V 100 bp overlap
ChIP LNCaP-C4-2_OE GSE88871.BRD4.LNCaP-C4-2_OE 167 bp overlap
ChIP NCI-H1963 GSE145028.BRD4.NCI-H1963 116 bp overlap
ChIP NCI-H1963_shASXL3 GSE145028.BRD4.NCI-H1963_shASXL3 221 bp overlap
ChIP NCI-H1963_shNONT GSE145028.BRD4.NCI-H1963_shNONT 352 bp overlap
CXXC5 1 dataset
ChIP prostate-cancer GSE136128.CXXC5.prostate-cancer 72 bp overlap
Cebpa 3 datasets
ChIP BLaER1 ENCFF364PUR 251 bp overlap
ChIP BLaER1 ENCFF460KDD 378 bp overlap
ChIP BLaER1 ENCFF680YXW 218 bp overlap
EGR2 1 dataset
ChIP HEK293 ENCFF336LFH 425 bp overlap
ESR1_Y537C 1 dataset
ChIP SUM44PE__ESR1_mutant_LTED GSE100074.ESR1_Y537C.SUM44PE__ESR1_mutant_LTED 162 bp overlap
ETV6 1 dataset
ChIP WTC11 ENCFF812SCD 437 bp overlap
FOXA1 13 datasets
Motif DE_48h DE_48h-FOXA1_MA0148.5 8 bp overlap
Motif DE_72h DE_72h-FOXA1_MA0148.5 8 bp overlap
ChIP HepG2 ENCFF207NVJ 281 bp overlap
ChIP HepG2 ENCFF740VZW 285 bp overlap
ChIP LNCaP-C4-2B_CST GSE123618.FOXA1.LNCaP-C4-2B_CST 111 bp overlap
ChIP LNCaP-C4-2B_TFS GSE123618.FOXA1.LNCaP-C4-2B_TFS 74 bp overlap
ChIP LNCaP_1F5 GSE30623.FOXA1.LNCaP_1F5 73 bp overlap
ChIP LNCaP_R1881 GSE69043.FOXA1.LNCaP_R1881 57 bp overlap
ChIP LNCaP_S2101-4H GSE114266.FOXA1.LNCaP_S2101-4H 200 bp overlap
ChIP LNCaP_S2101-4H GSE114266.FOXA1.LNCaP_S2101-4H 223 bp overlap
ChIP LNCaP_SHFOXA1_R1881 GSE37345.FOXA1.LNCaP_SHFOXA1_R1881 68 bp overlap
ChIP LNCaP_TFS GSE123618.FOXA1.LNCaP_TFS 109 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.FOXA1.pancreatic-progenitor_PP1 209 bp overlap
FOXA2 6 datasets
ChIP DE DE-FOXA2-1 636 bp overlap
ChIP DE DE-FOXA2-2 645 bp overlap
Motif DE_48h DE_48h-FOXA2_MA0047.4 8 bp overlap
Motif DE_72h DE_72h-FOXA2_MA0047.4 8 bp overlap
ChIP HepG2 ENCFF570ABM 405 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.FOXA2.pancreatic-progenitor_PP1 268 bp overlap
FOXA3 2 datasets
Motif DE_48h DE_48h-FOXA3_MA1683.2 7 bp overlap
Motif DE_72h DE_72h-FOXA3_MA1683.2 7 bp overlap
FOXB1 2 datasets
Motif DE_48h DE_48h-FOXB1_MA0845.1 11 bp overlap
Motif DE_72h DE_72h-FOXB1_MA0845.1 11 bp overlap
FOXC1 2 datasets
Motif DE_48h DE_48h-FOXC1_MA0032.2 11 bp overlap
Motif DE_72h DE_72h-FOXC1_MA0032.2 11 bp overlap
FOXC2 2 datasets
Motif DE_48h DE_48h-FOXC2_MA0846.2 11 bp overlap
Motif DE_72h DE_72h-FOXC2_MA0846.2 11 bp overlap
FOXD1 2 datasets
Motif DE_48h DE_48h-FOXD1_MA0031.2 7 bp overlap
Motif DE_72h DE_72h-FOXD1_MA0031.2 7 bp overlap
FOXD2 2 datasets
Motif DE_48h DE_48h-FOXD2_MA0847.4 11 bp overlap
Motif DE_72h DE_72h-FOXD2_MA0847.4 11 bp overlap
FOXD3 2 datasets
Motif DE_48h DE_48h-FOXD3_MA0041.3 14 bp overlap
Motif DE_72h DE_72h-FOXD3_MA0041.3 14 bp overlap
FOXE1 2 datasets
Motif DE_48h DE_48h-FOXE1_MA1487.3 12 bp overlap
Motif DE_72h DE_72h-FOXE1_MA1487.3 12 bp overlap
FOXF2 2 datasets
Motif DE_48h DE_48h-FOXF2_MA0030.2 9 bp overlap
Motif DE_72h DE_72h-FOXF2_MA0030.2 9 bp overlap
FOXG1 2 datasets
Motif DE_48h DE_48h-FOXG1_MA0613.1 8 bp overlap
Motif DE_72h DE_72h-FOXG1_MA0613.1 8 bp overlap
FOXI1 2 datasets
Motif DE_48h DE_48h-FOXI1_MA0042.2 7 bp overlap
Motif DE_72h DE_72h-FOXI1_MA0042.2 7 bp overlap
FOXJ3 1 dataset
ChIP WTC11 ENCFF909BTK 405 bp overlap
FOXK1 3 datasets
Motif DE_48h DE_48h-FOXK1_MA0852.3 7 bp overlap
Motif DE_72h DE_72h-FOXK1_MA0852.3 7 bp overlap
ChIP WTC11 ENCFF875IGU 391 bp overlap
FOXK2 2 datasets
Motif DE_48h DE_48h-FOXK2_MA1103.3 7 bp overlap
Motif DE_72h DE_72h-FOXK2_MA1103.3 7 bp overlap
FOXL1 2 datasets
Motif DE_48h DE_48h-FOXL1_MA0033.2 7 bp overlap
Motif DE_72h DE_72h-FOXL1_MA0033.2 7 bp overlap
FOXN3 2 datasets
Motif DE_48h DE_48h-FOXN3_MA1489.1 8 bp overlap
Motif DE_72h DE_72h-FOXN3_MA1489.1 8 bp overlap
FOXO4 2 datasets
Motif DE_48h DE_48h-FOXO4_MA0848.1 7 bp overlap
Motif DE_72h DE_72h-FOXO4_MA0848.1 7 bp overlap
FOXO6 2 datasets
Motif DE_48h DE_48h-FOXO6_MA0849.1 7 bp overlap
Motif DE_72h DE_72h-FOXO6_MA0849.1 7 bp overlap
FOXP1 3 datasets
Motif DE_48h DE_48h-FOXP1_MA0481.4 7 bp overlap
Motif DE_72h DE_72h-FOXP1_MA0481.4 7 bp overlap
ChIP WTC11 ENCFF338WGC 525 bp overlap
FOXP2 2 datasets
Motif DE_48h DE_48h-FOXP2_MA0593.2 9 bp overlap
Motif DE_72h DE_72h-FOXP2_MA0593.2 9 bp overlap
FOXP3 2 datasets
Motif DE_48h DE_48h-FOXP3_MA0850.1 7 bp overlap
Motif DE_72h DE_72h-FOXP3_MA0850.1 7 bp overlap
FOXP4 3 datasets
Motif DE_48h DE_48h-FOXP4_MA2117.1 7 bp overlap
Motif DE_72h DE_72h-FOXP4_MA2117.1 7 bp overlap
ChIP WTC11 ENCFF708TAF 377 bp overlap
FOXS1 2 datasets
Motif DE_48h DE_48h-FOXS1_MA2118.1 8 bp overlap
Motif DE_72h DE_72h-FOXS1_MA2118.1 8 bp overlap
Foxf1 2 datasets
Motif DE_48h DE_48h-Foxf1_MA1606.2 7 bp overlap
Motif DE_72h DE_72h-Foxf1_MA1606.2 7 bp overlap
Foxj2 5 datasets
Motif DE_48h DE_48h-Foxj2_MA0614.1 8 bp overlap
Motif DE_48h DE_48h-Foxj2_MA0614.1 8 bp overlap
Motif DE_72h DE_72h-Foxj2_MA0614.1 8 bp overlap
Motif DE_72h DE_72h-Foxj2_MA0614.1 8 bp overlap
Motif ES_0h ES_0h-Foxj2_MA0614.1 8 bp overlap
Foxj3 2 datasets
Motif DE_48h DE_48h-Foxj3_MA0851.2 9 bp overlap
Motif DE_72h DE_72h-Foxj3_MA0851.2 9 bp overlap
Foxl2 2 datasets
Motif DE_48h DE_48h-Foxl2_MA1607.2 10 bp overlap
Motif DE_72h DE_72h-Foxl2_MA1607.2 10 bp overlap
Foxn1 1 dataset
Motif DE_72h DE_72h-Foxn1_MA1684.1 6 bp overlap
Foxo1 2 datasets
Motif DE_48h DE_48h-Foxo1_MA0480.3 7 bp overlap
Motif DE_72h DE_72h-Foxo1_MA0480.3 7 bp overlap
Foxo3 2 datasets
Motif DE_48h DE_48h-Foxo3_MA0157.4 7 bp overlap
Motif DE_72h DE_72h-Foxo3_MA0157.4 7 bp overlap
GATA2 4 datasets
ChIP LNCaP GSE38391.GATA2.LNCaP 106 bp overlap
ChIP LNCaP GSE52725.GATA2.LNCaP 106 bp overlap
ChIP LNCaP_CSFCS GSE69043.GATA2.LNCaP_CSFCS 129 bp overlap
ChIP LNCaP_FBS GSE69043.GATA2.LNCaP_FBS 146 bp overlap
GATA4 3 datasets
ChIP DE DE-GATA4-1 417 bp overlap
ChIP DE DE-GATA4-2 718 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA4.pancreatic-progenitor_PP1 121 bp overlap
GATA6 2 datasets
ChIP DE DE-GATA6-1 295 bp overlap
ChIP DE DE-GATA6-2 494 bp overlap
Gfi1B 1 dataset
Motif DE_72h DE_72h-Gfi1B_MA0483.2 10 bp overlap
HOXB13 1 dataset
ChIP LNCaP GSE56288.HOXB13.LNCaP 53 bp overlap
IKZF1 2 datasets
ChIP K-562 ENCSR395HWC.IKZF1.K-562 78 bp overlap
ChIP K562 ENCFF348IBL 61 bp overlap
INSM1 1 dataset
Motif DE_72h DE_72h-INSM1_MA0155.1 12 bp overlap
JUND 1 dataset
ChIP H1 ENCFF468JZD 231 bp overlap
KLF5 1 dataset
Motif DE_72h DE_72h-KLF5_MA0599.1 10 bp overlap
MAF 1 dataset
Motif DE_72h DE_72h-MAF_MA1520.2 13 bp overlap
MAZ 1 dataset
Motif DE_72h DE_72h-MAZ_MA1522.2 8 bp overlap
MYOG 2 datasets
Motif DE_48h DE_48h-MYOG_MA0500.3 8 bp overlap
Motif DE_72h DE_72h-MYOG_MA0500.3 8 bp overlap
NHLH1 2 datasets
Motif DE_48h DE_48h-NHLH1_MA0048.3 9 bp overlap
Motif DE_72h DE_72h-NHLH1_MA0048.3 9 bp overlap
NR2F1 2 datasets
ChIP K-562 ENCSR970NKQ.NR2F1.K-562 89 bp overlap
ChIP K562 ENCFF221HJH 106 bp overlap
NR2F2 1 dataset
ChIP K562 ENCFF004YPK 132 bp overlap
Neurod2 2 datasets
Motif DE_48h DE_48h-Neurod2_MA1993.2 6 bp overlap
Motif DE_72h DE_72h-Neurod2_MA1993.2 6 bp overlap
Olig2 2 datasets
Motif DE_48h DE_48h-Olig2_MA1997.2 6 bp overlap
Motif DE_72h DE_72h-Olig2_MA1997.2 6 bp overlap
POLR2A 2 datasets
ChIP H1 ENCFF566JSR 260 bp overlap
ChIP upper lobe of left lung ENCFF199JUI 405 bp overlap
POU5F1_M 1 dataset
ChIP DE_D1 DED1-OCT4-M_Batch_II 485 bp overlap
RBBP5 3 datasets
ChIP H1 ENCFF905HFL 351 bp overlap
ChIP WA01 ENCSR000AQC.RBBP5.WA01 126 bp overlap
ChIP WA01 ENCSR000AQC.RBBP5.WA01 227 bp overlap
RBPJ 2 datasets
Motif DE_48h DE_48h-RBPJ_MA1116.2 6 bp overlap
Motif DE_72h DE_72h-RBPJ_MA1116.2 6 bp overlap
RELA 2 datasets
ChIP LNCaP_SICTR_TNFA GSE83860.RELA.LNCaP_SICTR_TNFA 83 bp overlap
ChIP LNCaP_SIFOXA1_TNFA GSE83860.RELA.LNCaP_SIFOXA1_TNFA 96 bp overlap
RUNX2 1 dataset
ChIP LNCaP-C4-2B GSE33889.RUNX2.LNCaP-C4-2B 211 bp overlap
SMAD2-3 1 dataset
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 333 bp overlap
SMAD2_3 5 datasets
ChIP DE_D1 S07-DE-d1-SMAD2_3-exp1 441 bp overlap
ChIP DE_D1 S12-DE-d1-SMAD2_3-exp1 434 bp overlap
ChIP DE_D1 S39-DE-d1-SMAD2_3-exp2 550 bp overlap
ChIP DE_D2 S01-DE-d2-SMAD2_3-exp1 600 bp overlap
ChIP DE_D2 S53-DE-d2-SMAD2_3-exp2 628 bp overlap
SMAD4 1 dataset
ChIP endoderm GSE29422.SMAD4.endoderm 250 bp overlap
SOX10 2 datasets
Motif DE_48h DE_48h-SOX10_MA0442.3 6 bp overlap
Motif DE_72h DE_72h-SOX10_MA0442.3 6 bp overlap
SOX13 1 dataset
Motif DE_72h DE_72h-SOX13_MA1120.2 7 bp overlap
SOX17 1 dataset
ChIP DE_D2 DED2-SOX17_Batch_II 369 bp overlap
SOX17_M 1 dataset
ChIP DE_D2 DED2-SOX17-M_Batch_II 718 bp overlap
SOX2 2 datasets
Motif DE_72h DE_72h-SOX2_MA0143.5 7 bp overlap
ChIP HNSC GSE69479.SOX2.HNSC 321 bp overlap
SOX21 1 dataset
ChIP neuroepithelilal-cells GSE110505.SOX21.neuroepithelilal-cells 286 bp overlap
SOX4 3 datasets
Motif DE_48h DE_48h-SOX4_MA0867.3 8 bp overlap
Motif DE_72h DE_72h-SOX4_MA0867.3 8 bp overlap
Motif DE_72h DE_72h-SOX4_MA0867.3 8 bp overlap
SOX8 1 dataset
Motif DE_72h DE_72h-SOX8_MA0868.3 7 bp overlap
SOX9 1 dataset
Motif DE_72h DE_72h-SOX9_MA0077.2 8 bp overlap
SP1 1 dataset
ChIP H1 ENCFF263FUH 321 bp overlap
SP8 1 dataset
Motif DE_72h DE_72h-SP8_MA0747.2 11 bp overlap
STAT3 5 datasets
ChIP HCC1187 GSE152203.STAT3.HCC1187 122 bp overlap
ChIP MCF-7_jc5834 GSE126004.STAT3.MCF-7_jc5834 55 bp overlap
ChIP MCF-7_jc5836 GSE126004.STAT3.MCF-7_jc5836 78 bp overlap
ChIP MCF-7_jc5838 GSE126004.STAT3.MCF-7_jc5838 76 bp overlap
ChIP MCF-7_jc5840 GSE126004.STAT3.MCF-7_jc5840 146 bp overlap
Sox11 2 datasets
Motif DE_48h DE_48h-Sox11_MA0869.3 8 bp overlap
Motif DE_72h DE_72h-Sox11_MA0869.3 8 bp overlap
Sox17 1 dataset
Motif DE_72h DE_72h-Sox17_MA0078.3 10 bp overlap
Sox3 1 dataset
Motif DE_72h DE_72h-Sox3_MA0514.3 7 bp overlap
Sox5 1 dataset
Motif DE_72h DE_72h-Sox5_MA0087.3 8 bp overlap
Sox7 1 dataset
Motif DE_72h DE_72h-Sox7_MA2095.1 10 bp overlap
TAF1 2 datasets
ChIP H1 ENCFF478SZO 485 bp overlap
ChIP WA01 ENCSR000BHO.TAF1.WA01 206 bp overlap
TAL1 1 dataset
ChIP K-562 GSE107726.TAL1.K-562 54 bp overlap
TCF12 1 dataset
ChIP H1 ENCFF203EBH 251 bp overlap
TFAP4::FLI1 2 datasets
Motif DE_48h DE_48h-TFAP4FLI1_MA1967.2 14 bp overlap
Motif DE_72h DE_72h-TFAP4FLI1_MA1967.2 14 bp overlap
TGIF2 1 dataset
ChIP WTC11 ENCFF649SHI 441 bp overlap
TLE3 1 dataset
ChIP LNCaP_r1881 GSE94682.TLE3.LNCaP_r1881 71 bp overlap
Tcf12 2 datasets
Motif DE_48h DE_48h-Tcf12_MA0521.3 6 bp overlap
Motif DE_72h DE_72h-Tcf12_MA0521.3 6 bp overlap
Twist2 2 datasets
Motif DE_48h DE_48h-Twist2_MA0633.3 6 bp overlap
Motif DE_72h DE_72h-Twist2_MA0633.3 6 bp overlap
Wt1 1 dataset
Motif DE_72h DE_72h-Wt1_MA1627.2 10 bp overlap
YY1 3 datasets
ChIP H1 ENCFF524BTL 337 bp overlap
ChIP WA01 ENCSR000BKD.YY1.WA01 93 bp overlap
ChIP WA01 ENCSR000BKD.YY1.WA01 136 bp overlap
ZBTB21 1 dataset
ChIP WTC11 ENCFF677ZYY 401 bp overlap
ZKSCAN1 5 datasets
Motif DE_48h DE_48h-ZKSCAN1_MA1585.2 9 bp overlap
Motif DE_48h DE_48h-ZKSCAN1_MA1585.2 9 bp overlap
Motif DE_72h DE_72h-ZKSCAN1_MA1585.2 9 bp overlap
Motif DE_72h DE_72h-ZKSCAN1_MA1585.2 9 bp overlap
Motif ES_0h ES_0h-ZKSCAN1_MA1585.2 9 bp overlap
ZNF281 2 datasets
ChIP HepG2 ENCFF585QNU 325 bp overlap
ChIP WTC11 ENCFF551GAV 377 bp overlap
ZNF331 2 datasets
Motif DE_48h DE_48h-ZNF331_MA1726.2 10 bp overlap
Motif DE_72h DE_72h-ZNF331_MA1726.2 10 bp overlap
ZNF416 2 datasets
Motif DE_48h DE_48h-ZNF416_MA1979.2 10 bp overlap
Motif DE_72h DE_72h-ZNF416_MA1979.2 10 bp overlap
ZNF418 3 datasets
Motif DE_48h DE_48h-ZNF418_MA1980.1 15 bp overlap
Motif DE_72h DE_72h-ZNF418_MA1980.1 15 bp overlap
Motif ES_0h ES_0h-ZNF418_MA1980.1 15 bp overlap
ZNF530 1 dataset
Motif DE_72h DE_72h-ZNF530_MA1981.2 14 bp overlap
ZNF768 3 datasets
Motif DE_48h DE_48h-ZNF768_MA1731.2 9 bp overlap
Motif DE_72h DE_72h-ZNF768_MA1731.2 9 bp overlap
Motif ES_0h ES_0h-ZNF768_MA1731.2 9 bp overlap
ZNF770 1 dataset
Motif DE_72h DE_72h-ZNF770_MA2099.1 8 bp overlap
Zfp809 2 datasets
Motif DE_48h DE_48h-Zfp809_MA2125.1 9 bp overlap
Motif DE_72h DE_72h-Zfp809_MA2125.1 9 bp overlap