chr10 : 69,568,733 69,569,392
659 bp 99 TFs 4 linked genes
This 659 bp open chromatin element is linked to 4 target genes and is bound by 99 transcription factors.
Linked Genes
4 genes
Distance
Link type
Gene Expression Dist. to TSS Distance Link type
NEUROG3 4.0 kb Proximal Proximity
FAM241B 61.2 kb Distal Multiome
COL13A1 232.9 kb Distal Multiome
HK1 250.2 kb Distal Multiome
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr10:69,563,733 – 69,574,392
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
99 transcription factors
Source
Cell type
ARID1A 1 dataset
ChIP HAP1 GSE108387.ARID1A.HAP1 316 bp overlap
ASH2L 2 datasets
ChIP H1 ENCFF399KAM 406 bp overlap
ChIP WA01 ENCSR850KIP.ASH2L.WA01 659 bp overlap
ATF2 1 dataset
ChIP WA01 ENCSR000BQU.ATF2.WA01 265 bp overlap
Ar 1 dataset
Motif ES_0h ES_0h-Ar_MA0007.4 16 bp overlap
BCOR 1 dataset
ChIP WA01_KDM2B_KO GSE104690.BCOR.WA01_KDM2B_KO 659 bp overlap
BRD1 2 datasets
ChIP HUES-64 GSE104059.BRD1.HUES-64 201 bp overlap
ChIP HUES-64 GSE104059.BRD1.HUES-64 243 bp overlap
BRD4 4 datasets
ChIP CLL_patient2_4h_CpG_BET GSE109411.BRD4.CLL_patient2_4h_CpG_BET 340 bp overlap
ChIP HAP1 GSE108387.BRD4.HAP1 346 bp overlap
ChIP hESC GSE33281.BRD4.hESC 221 bp overlap
ChIP retina_AB1-FW15 GSE86981.BRD4.retina_AB1-FW15 290 bp overlap
CHD1 2 datasets
ChIP WA01 ENCSR000AQK.CHD1.WA01 85 bp overlap
ChIP WA01 ENCSR000AQK.CHD1.WA01 196 bp overlap
CHD2 1 dataset
ChIP WA01 ENCSR000EBT.CHD2.WA01 335 bp overlap
CHD7 1 dataset
ChIP WA01 ENCSR000AVA.CHD7.WA01 147 bp overlap
CREB1 1 dataset
ChIP WA01 ENCSR000BSN.CREB1.WA01 369 bp overlap
CTBP2 3 datasets
ChIP H1 ENCFF329MAX 531 bp overlap
ChIP H1 ENCFF329MAX 392 bp overlap
ChIP WA01 ENCSR000EUO.CTBP2.WA01 659 bp overlap
CTNNB1 1 dataset
ChIP hiPSC_D2 GSE132532.CTNNB1.hiPSC_D2 427 bp overlap
E2F6 1 dataset
ChIP WA01 ENCSR000BSI.E2F6.WA01 577 bp overlap
EP300 4 datasets
ChIP H1 ENCFF927IYK 297 bp overlap
ChIP H1 ENCFF937OPV 205 bp overlap
ChIP WA01 ENCSR000AUQ.EP300.WA01 435 bp overlap
ChIP WA01 ENCSR000BKK.EP300.WA01 389 bp overlap
ESR1 1 dataset
ChIP Ishikawa_DMSO_D538G_clone3 GSE132426.ESR1.Ishikawa_DMSO_D538G_clone3 289 bp overlap
EZH2 30 datasets
ChIP A673 ENCFF790MVL 397 bp overlap
ChIP A673 ENCFF790MVL 521 bp overlap
ChIP A673 ENCFF790MVL 114 bp overlap
ChIP A673 ENCFF955JRZ 365 bp overlap
ChIP A673 ENCFF955JRZ 114 bp overlap
ChIP GM23248 ENCFF404ZHM 501 bp overlap
ChIP GM23248 ENCFF404ZHM 227 bp overlap
ChIP GM23248 ENCSR131FFJ.EZH2.GM23248 308 bp overlap
ChIP H1 ENCFF232NZA 242 bp overlap
ChIP HUVEC-C GSE109625.EZH2.HUVEC-C 232 bp overlap
ChIP HUVEC-C_VEGF_1h GSE109625.EZH2.HUVEC-C_VEGF_1h 62 bp overlap
ChIP Hep-3B2-1-7 GSE128137.EZH2.Hep-3B2-1-7 275 bp overlap
ChIP LNCaP GSE39459.EZH2.LNCaP 241 bp overlap
ChIP LNCaP GSE39459.EZH2.LNCaP 61 bp overlap
ChIP LNCaP-abl GSE39459.EZH2.LNCaP-abl 278 bp overlap
ChIP LNCaP-abl GSE39459.EZH2.LNCaP-abl 291 bp overlap
ChIP T98G GSE112240.EZH2.T98G 269 bp overlap
ChIP astrocyte ENCFF365JTP 416 bp overlap
ChIP endothelial cell of umbilical vein ENCFF539AKL 107 bp overlap
ChIP epithelial_mammary ENCSR000ARE.EZH2.epithelial_mammary 235 bp overlap
ChIP epithelial_mammary ENCSR000ARE.EZH2.epithelial_mammary 273 bp overlap
ChIP fibroblast_LUNG ENCSR000ARO.EZH2.fibroblast_LUNG 177 bp overlap
ChIP hepatocyte ENCFF552DZB 66 bp overlap
ChIP hepatocyte ENCFF552DZB 119 bp overlap
ChIP mammary epithelial cell ENCFF224GAI 56 bp overlap
ChIP myotube ENCSR000ASZ.EZH2.myotube 282 bp overlap
ChIP myotube ENCSR000ASZ.EZH2.myotube 243 bp overlap
ChIP neural ENCSR511CUH.EZH2.neural 595 bp overlap
ChIP neural progenitor cell ENCFF018MKA 360 bp overlap
ChIP neural progenitor cell ENCFF472NFV 659 bp overlap
FOXP1 3 datasets
ChIP H9 GSE31006.FOXP1.H9 216 bp overlap
ChIP H9 GSE31006.FOXP1.H9 172 bp overlap
ChIP WTC11 ENCFF338WGC 525 bp overlap
GATA6 2 datasets
ChIP DE_D1 S41-DE-d1-GATA6-exp2 271 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 268 bp overlap
GLIS3 1 dataset
ChIP H9_plus GSE109562.GLIS3.H9_plus 659 bp overlap
GTF2I 1 dataset
ChIP WTC11 ENCFF255XXZ 285 bp overlap
GTF3C2 1 dataset
ChIP H9 GSE94418.GTF3C2.H9 258 bp overlap
HDAC2 3 datasets
ChIP H1 ENCFF353UJQ 289 bp overlap
ChIP WA01 ENCSR000AVB.HDAC2.WA01 629 bp overlap
ChIP WA01 ENCSR000BNR.HDAC2.WA01 135 bp overlap
Hic1 2 datasets
Motif DE_12h DE_12h-Hic1_MA0739.2 8 bp overlap
Motif ES_0h ES_0h-Hic1_MA0739.2 8 bp overlap
JARID2 4 datasets
ChIP EDOMIPS2 GSE48516.JARID2.EDOMIPS2 250 bp overlap
ChIP MRC-5_IPS25_MEG3 GSE48515.JARID2.MRC-5_IPS25_MEG3 659 bp overlap
ChIP UTEIPS11 GSE48516.JARID2.UTEIPS11 84 bp overlap
ChIP UTEIPS7 GSE48516.JARID2.UTEIPS7 388 bp overlap
JUN 4 datasets
ChIP DE_D1 S40-DE-d1-JUN-exp2 140 bp overlap
ChIP ESC S24-ESC-d0-JUN-exp1 508 bp overlap
ChIP HUES-8 GSE109524.JUN.HUES-8 615 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.JUN.HUES-8_DE_ctrl 514 bp overlap
JUND 2 datasets
ChIP H1 ENCFF468JZD 231 bp overlap
ChIP WA01 ENCSR000BKP.JUND.WA01 258 bp overlap
KDM1A 3 datasets
ChIP H1 ENCFF696SGD 343 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.KDM1A.pancreatic-progenitor_PP1 281 bp overlap
ChIP pancreatic-progenitor_PP2-KDM1A-inh GSE104840.KDM1A.pancreatic-progenitor_PP2-KDM1A-inh 235 bp overlap
KDM4A 5 datasets
ChIP H1 ENCFF078LED 290 bp overlap
ChIP H1 ENCFF078LED 315 bp overlap
ChIP WA01 ENCSR000AVC.KDM4A.WA01 128 bp overlap
ChIP WA01 ENCSR000AVC.KDM4A.WA01 481 bp overlap
ChIP hiPSC_IB12 GSE106870.KDM4A.hiPSC_IB12 166 bp overlap
KLF4 2 datasets
ChIP HAP1 GSE130417.KLF4.HAP1 192 bp overlap
ChIP hiPSC GSE56567.KLF4.hiPSC 245 bp overlap
MAX 3 datasets
ChIP H1 ENCFF914VQY 135 bp overlap
ChIP WA01 ENCSR000EUP.MAX.WA01 158 bp overlap
ChIP WTC11 ENCFF223QFY 531 bp overlap
MYC 1 dataset
ChIP H1 ENCFF794ZJT 265 bp overlap
NANOG 8 datasets
ChIP GM23338 ENCFF065NZG 311 bp overlap
ChIP H1 ENCFF747ZPQ 241 bp overlap
ChIP HUES-8 GSE109524.NANOG.HUES-8 659 bp overlap
ChIP WA01 ENCSR000BMT.NANOG.WA01 220 bp overlap
ChIP WA01 ERP004238.NANOG.WA01 245 bp overlap
ChIP WA01_3IL ERP004238.NANOG.WA01_3IL 599 bp overlap
ChIP WA09 GSE105028.NANOG.WA09 426 bp overlap
ChIP WA09_heat-shock GSE105028.NANOG.WA09_heat-shock 242 bp overlap
NIPBL 4 datasets
ChIP WA09_heat-shock GSE105028.NIPBL.WA09_heat-shock 527 bp overlap
ChIP hESC GSE64758.NIPBL.hESC 231 bp overlap
ChIP hESC_WNT3A GSE64758.NIPBL.hESC_WNT3A 320 bp overlap
ChIP hESC_WNT3A_ACTIVIN GSE64758.NIPBL.hESC_WNT3A_ACTIVIN 211 bp overlap
NR3C1 1 dataset
Motif ES_0h ES_0h-NR3C1_MA0113.4 15 bp overlap
NR3C2 1 dataset
Motif ES_0h ES_0h-NR3C2_MA0727.2 15 bp overlap
PDX1 1 dataset
ChIP hiPSC_derived_PPs GSE106949.PDX1.hiPSC_derived_PPs 396 bp overlap
PHF8 2 datasets
ChIP H1 ENCFF427UFV 445 bp overlap
ChIP WA01 ENCSR000ATK.PHF8.WA01 134 bp overlap
POU2F1 2 datasets
Motif DE_12h DE_12h-POU2F1_MA0785.2 9 bp overlap
Motif ES_0h ES_0h-POU2F1_MA0785.2 9 bp overlap
POU2F1::SOX2 2 datasets
Motif DE_12h DE_12h-POU2F1SOX2_MA1962.1 17 bp overlap
Motif ES_0h ES_0h-POU2F1SOX2_MA1962.1 17 bp overlap
POU3F1 2 datasets
Motif DE_12h DE_12h-POU3F1_MA0786.2 10 bp overlap
Motif ES_0h ES_0h-POU3F1_MA0786.2 10 bp overlap
POU3F2 2 datasets
Motif DE_12h DE_12h-POU3F2_MA0787.1 12 bp overlap
Motif ES_0h ES_0h-POU3F2_MA0787.1 12 bp overlap
POU3F3 2 datasets
Motif DE_12h DE_12h-POU3F3_MA0788.1 13 bp overlap
Motif ES_0h ES_0h-POU3F3_MA0788.1 13 bp overlap
POU3F4 2 datasets
Motif DE_12h DE_12h-POU3F4_MA0789.1 9 bp overlap
Motif ES_0h ES_0h-POU3F4_MA0789.1 9 bp overlap
POU5F1 13 datasets
ChIP BG03 GSE21614.POU5F1.BG03 304 bp overlap
ChIP DE_D1 DED1-OCT4_Batch_II 550 bp overlap
ChIP GM23338 ENCFF333SNB 311 bp overlap
ChIP NCCIT GSE36134.POU5F1.NCCIT 514 bp overlap
ChIP SKM-1 GSE93706.POU5F1.SKM-1 284 bp overlap
ChIP WA01_3IL ERP004238.POU5F1.WA01_3IL 312 bp overlap
ChIP WA01_3IL ERP004238.POU5F1.WA01_3IL 173 bp overlap
ChIP WA09 GSE105028.POU5F1.WA09 268 bp overlap
ChIP WA09_heat-shock GSE105028.POU5F1.WA09_heat-shock 196 bp overlap
ChIP hESC ENCSR264RJX.POU5F1.hESC 210 bp overlap
ChIP hESC_PRIMED GSE69646.POU5F1.hESC_PRIMED 536 bp overlap
ChIP hiPSC GSE56567.POU5F1.hiPSC 286 bp overlap
ChIP hiPSC_3s2 GSE81899.POU5F1.hiPSC_3s2 146 bp overlap
POU5F1B 2 datasets
Motif DE_12h DE_12h-POU5F1B_MA0792.1 9 bp overlap
Motif ES_0h ES_0h-POU5F1B_MA0792.1 9 bp overlap
POU5F1_M 1 dataset
ChIP DE_D1 DED1-OCT4-M_Batch_II 603 bp overlap
PRDM14 2 datasets
ChIP NCCIT GSE71675.PRDM14.NCCIT 411 bp overlap
ChIP hESC GSE138674.PRDM14.hESC 538 bp overlap
Pgr 1 dataset
Motif ES_0h ES_0h-Pgr_MA2323.1 17 bp overlap
Plagl1 1 dataset
Motif ES_0h ES_0h-Plagl1_MA1615.2 8 bp overlap
Pou5f1::Sox2 2 datasets
Motif DE_12h DE_12h-Pou5f1Sox2_MA0142.1 15 bp overlap
Motif ES_0h ES_0h-Pou5f1Sox2_MA0142.1 15 bp overlap
RAD21 1 dataset
ChIP hiPSC_IIIB12 GSE106870.RAD21.hiPSC_IIIB12 173 bp overlap
RARA 2 datasets
ChIP hiPSC_D3 GSE109172.RARA.hiPSC_D3 429 bp overlap
ChIP hiPSC_D4 GSE109172.RARA.hiPSC_D4 377 bp overlap
RBBP5 2 datasets
ChIP H1 ENCFF905HFL 451 bp overlap
ChIP WA01 ENCSR000AQC.RBBP5.WA01 635 bp overlap
REST 14 datasets
ChIP A-549 ENCSR000BQP.REST.A-549 173 bp overlap
ChIP CD4 GSE49570.REST.CD4 163 bp overlap
ChIP GM23338 ENCFF024TCL 265 bp overlap
ChIP GM23338 ENCFF024TCL 265 bp overlap
ChIP GM23338 ENCSR871KYB.REST.GM23338 388 bp overlap
ChIP HL-60 ENCFF589LOF 240 bp overlap
ChIP K-562 ENCSR000BMW.REST.K-562 164 bp overlap
ChIP PANC-1 ENCSR000BJO.REST.PANC-1 242 bp overlap
ChIP PANC-1 ENCSR000BPK.REST.PANC-1 101 bp overlap
ChIP PFSK1 ENCSR000BOX.REST.PFSK1 145 bp overlap
ChIP PFSK1 ENCSR000BJP.REST.PFSK1 103 bp overlap
ChIP colorectal-cancer_shCTRL_intact GSE112555.REST.colorectal-cancer_shCTRL_intact 509 bp overlap
ChIP hiPSC_IB12 GSE106870.REST.hiPSC_IB12 215 bp overlap
ChIP hiPSC_IID12 GSE106870.REST.hiPSC_IID12 254 bp overlap
RNF2 2 datasets
ChIP HUES-64 GSE104059.RNF2.HUES-64 285 bp overlap
ChIP WA01 GSE104690.RNF2.WA01 541 bp overlap
RUNX1 1 dataset
ChIP Jurkat GSE85524.RUNX1.Jurkat 189 bp overlap
RXRA 1 dataset
ChIP WA01 ENCSR000BJW.RXRA.WA01 172 bp overlap
SALL3 1 dataset
ChIP hiPSC GSE104863.SALL3.hiPSC 517 bp overlap
SIN3A 3 datasets
ChIP H1 ENCFF896IJG 301 bp overlap
ChIP WA01 ENCSR000EBO.SIN3A.WA01 209 bp overlap
ChIP WA01 ENCSR000BIS.SIN3A.WA01 120 bp overlap
SMAD2 3 datasets
Motif DE_12h DE_12h-SMAD2_MA1964.2 6 bp overlap
ChIP hESC GSE29422.SMAD2.hESC 239 bp overlap
ChIP hESC_YAP-_activinA_15h GSE99202.SMAD2.hESC_YAP-_activinA_15h 374 bp overlap
SMAD2-3 2 datasets
ChIP HUES-8 GSE109524.SMAD2-3.HUES-8 659 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.SMAD2-3.HUES-8_DE_ctrl 659 bp overlap
SMAD2_3 3 datasets
ChIP DE_D1 S39-DE-d1-SMAD2_3-exp2 659 bp overlap
ChIP ESC S23-ESC-d0-SMAD2_3-exp1 456 bp overlap
ChIP ESC S33-ESC-d0-SMAD2_3-exp2 659 bp overlap
SMAD3 4 datasets
ChIP BG03 GSE21614.SMAD3.BG03 181 bp overlap
ChIP BG03_DIFF_0H GSE36578.SMAD3.BG03_DIFF_0H 272 bp overlap
ChIP BG03_DIFF_2H GSE36578.SMAD3.BG03_DIFF_2H 210 bp overlap
ChIP hESC GSE29422.SMAD3.hESC 301 bp overlap
SMARCA4 4 datasets
ChIP 22Rv1 GSE115615.SMARCA4.22Rv1 187 bp overlap
ChIP NSC GSE125033.SMARCA4.NSC 451 bp overlap
ChIP hiPSC GSE124903.SMARCA4.hiPSC 400 bp overlap
ChIP hiPSC_WT GSE124903.SMARCA4.hiPSC_WT 655 bp overlap
SMARCB1 2 datasets
ChIP hiPSC GSE124903.SMARCB1.hiPSC 468 bp overlap
ChIP hiPSC_WT GSE124903.SMARCB1.hiPSC_WT 525 bp overlap
SMARCC1 2 datasets
ChIP hiPSC GSE124903.SMARCC1.hiPSC 443 bp overlap
ChIP hiPSC_WT GSE124903.SMARCC1.hiPSC_WT 514 bp overlap
SOX10 1 dataset
Motif DE_12h DE_12h-SOX10_MA0442.3 6 bp overlap
SOX2 3 datasets
ChIP H9 GSE46837.SOX2.H9 387 bp overlap
ChIP hESC GSE69479.SOX2.hESC 195 bp overlap
ChIP hiPSC GSE56567.SOX2.hiPSC 414 bp overlap
SP1 4 datasets
ChIP H1 ENCFF263FUH 321 bp overlap
ChIP WA01 ENCSR000BIR.SP1.WA01 508 bp overlap
ChIP WTC11 ENCFF688PEU 501 bp overlap
ChIP WTC11 ENCFF688PEU 414 bp overlap
SP4 1 dataset
ChIP WA01 ENCSR000BQV.SP4.WA01 198 bp overlap
SREBP2 1 dataset
ChIP HCC70 GSE126380.SREBP2.HCC70 354 bp overlap
SS18 1 dataset
ChIP SYO-1_shRING1A-B GSE139053.SS18.SYO-1_shRING1A-B 321 bp overlap
SUZ12 4 datasets
ChIP H1 ENCFF881NFR 233 bp overlap
ChIP LNCaP GSE39459.SUZ12.LNCaP 427 bp overlap
ChIP LNCaP-abl GSE39459.SUZ12.LNCaP-abl 140 bp overlap
ChIP NT2-D1 ENCSR000EXH.SUZ12.NT2-D1 129 bp overlap
Sox11 2 datasets
Motif DE_12h DE_12h-Sox11_MA0869.3 8 bp overlap
Motif ES_0h ES_0h-Sox11_MA0869.3 8 bp overlap
TAF1 3 datasets
ChIP H1 ENCFF478SZO 485 bp overlap
ChIP H1 ENCFF478SZO 336 bp overlap
ChIP WA01 ENCSR000BHO.TAF1.WA01 475 bp overlap
TBP 4 datasets
ChIP H1 ENCFF859IIO 377 bp overlap
ChIP WA01 ENCSR000ECB.TBP.WA01 386 bp overlap
ChIP hiPSC_WTa_RNase-neg GSE128135.TBP.hiPSC_WTa_RNase-neg 258 bp overlap
ChIP hiPSC_WTb_RNase-neg GSE128135.TBP.hiPSC_WTb_RNase-neg 412 bp overlap
TCF12 2 datasets
ChIP H1 ENCFF203EBH 251 bp overlap
ChIP WA01 ENCSR000BIT.TCF12.WA01 391 bp overlap
TCF3 1 dataset
ChIP NPC GSE154479.TCF3.NPC 259 bp overlap
TEAD1 2 datasets
Motif DE_12h DE_12h-TEAD1_MA0090.4 9 bp overlap
Motif ES_0h ES_0h-TEAD1_MA0090.4 9 bp overlap
TEAD3 2 datasets
Motif DE_12h DE_12h-TEAD3_MA0808.1 8 bp overlap
Motif ES_0h ES_0h-TEAD3_MA0808.1 8 bp overlap
TEAD4 4 datasets
Motif DE_12h DE_12h-TEAD4_MA0809.3 8 bp overlap
ChIP ESC S26-ESC-d0-TEAD4-exp1 287 bp overlap
Motif ES_0h ES_0h-TEAD4_MA0809.3 8 bp overlap
ChIP H1 ENCFF778PAX 245 bp overlap
TFAP2C 2 datasets
ChIP UCLA1-hESCs_TFAP2C--_1d_TFAP2C_Induc GSE101074.TFAP2C.UCLA1-hESCs_TFAP2C--_1d_TFAP2C_Induc 659 bp overlap
ChIP UCLA1-hESCs_TFAP2C--_2d_TFAP2C_Induc GSE101074.TFAP2C.UCLA1-hESCs_TFAP2C--_2d_TFAP2C_Induc 590 bp overlap
TGIF2 1 dataset
ChIP WTC11 ENCFF649SHI 405 bp overlap
TP63 1 dataset
ChIP breast-organoid GSE113909.TP63.breast-organoid 116 bp overlap
TRIM28 1 dataset
ChIP hESC GSE115387.TRIM28.hESC 418 bp overlap
USF1 1 dataset
ChIP WA01 ENCSR000BIU.USF1.WA01 238 bp overlap
USF2 1 dataset
ChIP WTC11 ENCFF139JAW 417 bp overlap
YY1 2 datasets
ChIP H1 ENCFF524BTL 337 bp overlap
ChIP WA01 ENCSR000BKD.YY1.WA01 336 bp overlap
ZBTB21 1 dataset
ChIP WTC11 ENCFF677ZYY 401 bp overlap
ZNF143 2 datasets
ChIP WA01 ENCSR000EBW.ZNF143.WA01 225 bp overlap
ChIP WA09_heat-shock GSE105028.ZNF143.WA09_heat-shock 266 bp overlap
ZNF189 1 dataset
Motif ES_0h ES_0h-ZNF189_MA1725.2 9 bp overlap
ZNF207 1 dataset
ChIP WA09 GSE118632.ZNF207.WA09 331 bp overlap
ZNF281 2 datasets
ChIP WTC11 ENCFF551GAV 377 bp overlap
ChIP WTC11 ENCFF551GAV 360 bp overlap
ZNF423 1 dataset
ChIP WTC11 ENCFF574PBR 317 bp overlap
ZNF454 1 dataset
Motif DE_12h DE_12h-ZNF454_MA1712.2 17 bp overlap