chr8 : 48,643,345 48,643,944
599 bp 84 TFs 0 linked genes
This 599 bp open chromatin element has no linked target genes and is bound by 84 transcription factors.
Linked Genes
No linked genes
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr8:48,638,345 – 48,648,944
This element Other elements ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
84 transcription factors
Source
Cell type
AR 2 datasets
ChIP endometrial-stromal-cell GSE119432.AR.endometrial-stromal-cell 218 bp overlap
ChIP fibroblast_prostate-cancer_PCDF2 GSE126852.AR.fibroblast_prostate-cancer_PCDF2 193 bp overlap
ARNT 1 dataset
ChIP SK-MEL-28 GSE85352.ARNT.SK-MEL-28 313 bp overlap
ATRX 3 datasets
ChIP metastatic-neuroblastoma_CHLA90 GSE100148.ATRX.metastatic-neuroblastoma_CHLA90 599 bp overlap
ChIP metastatic-neuroblastoma_SKNFI GSE100148.ATRX.metastatic-neuroblastoma_SKNFI 274 bp overlap
ChIP metastatic-neuroblastoma_SKNMM GSE100148.ATRX.metastatic-neuroblastoma_SKNMM 206 bp overlap
BNC2 2 datasets
ChIP SK-N-SH ENCFF174EMC 425 bp overlap
ChIP SK-N-SH_all-trans-retinoic-acid_6-uM_48-hour ENCSR754GYI.BNC2.SK-N-SH_all-trans-retinoic-acid_6-uM_48-hour 247 bp overlap
BRD2 1 dataset
ChIP SUM159PT_JQ1_MGSK2801 GSE116879.BRD2.SUM159PT_JQ1_MGSK2801 199 bp overlap
BRD4 10 datasets
ChIP IMR-90_QUIES GSE74238.BRD4.IMR-90_QUIES 119 bp overlap
ChIP Kelly_sensitive GSE103030.BRD4.Kelly_sensitive 138 bp overlap
ChIP MCF-10A ERP003925.BRD4.MCF-10A 199 bp overlap
ChIP SK-N-BE2-C_Naive-veh GSE107706.BRD4.SK-N-BE2-C_Naive-veh 138 bp overlap
ChIP SUM159PT_100nMtrametinib300nMJQ1_24h GSE87418.BRD4.SUM159PT_100nMtrametinib300nMJQ1_24h 400 bp overlap
ChIP SUM159PT_100nMtrametinib_24h GSE87418.BRD4.SUM159PT_100nMtrametinib_24h 599 bp overlap
ChIP SUM159PT_100nMtrametinib_72h GSE87418.BRD4.SUM159PT_100nMtrametinib_72h 496 bp overlap
ChIP SUM159PT_100nMtrametinib_8h GSE87418.BRD4.SUM159PT_100nMtrametinib_8h 511 bp overlap
ChIP SUM159PT_DMSO_8h GSE87418.BRD4.SUM159PT_DMSO_8h 388 bp overlap
ChIP SUM159PT_shMYC_dox_48h GSE87418.BRD4.SUM159PT_shMYC_dox_48h 236 bp overlap
CDK8 1 dataset
ChIP myometrium_PT1063 GSE128230.CDK8.myometrium_PT1063 62 bp overlap
CHD2 2 datasets
ChIP SK-N-SH ENCFF669KMB 361 bp overlap
ChIP SK-N-SH ENCSR274SLQ.CHD2.SK-N-SH 236 bp overlap
CREB5 2 datasets
ChIP SK-N-SH ENCFF144PMI 345 bp overlap
ChIP SK-N-SH_all-trans-retinoic-acid_6-uM_48-hour ENCSR758GOA.CREB5.SK-N-SH_all-trans-retinoic-acid_6-uM_48-hour 355 bp overlap
CSDC2 2 datasets
ChIP SK-N-SH ENCFF868MXA 351 bp overlap
ChIP SK-N-SH_all-trans-retinoic-acid_6-uM_48-hour ENCSR474KBG.CSDC2.SK-N-SH_all-trans-retinoic-acid_6-uM_48-hour 228 bp overlap
CTCF 8 datasets
ChIP adrenal-gland ENCSR899JSO.CTCF.adrenal-gland 222 bp overlap
ChIP gastroesophageal-sphincter ENCSR206ETG.CTCF.gastroesophageal-sphincter 351 bp overlap
ChIP hESC_D5 GSE116862.CTCF.hESC_D5 452 bp overlap
ChIP heart ENCSR232OFD.CTCF.heart 245 bp overlap
ChIP heart left ventricle ENCFF354HOQ 461 bp overlap
ChIP myotube ENCSR000ANS.CTCF.myotube 157 bp overlap
ChIP uterus ENCSR527TPP.CTCF.uterus 327 bp overlap
ChIP uterus ENCSR684PGO.CTCF.uterus 211 bp overlap
Cebpa 1 dataset
ChIP BLaER1 ENCFF031ISE 258 bp overlap
EP300 7 datasets
ChIP SK-N-SH ENCFF451CNG 385 bp overlap
ChIP SK-N-SH ENCFF829RWA 377 bp overlap
ChIP SK-N-SH ENCSR000EHV.EP300.SK-N-SH 365 bp overlap
ChIP SK-N-SH ENCSR000BUA.EP300.SK-N-SH 331 bp overlap
ChIP sigmoid colon ENCFF524QSR 271 bp overlap
ChIP sigmoid colon ENCFF682PXQ 231 bp overlap
ChIP tibial nerve ENCFF346AYA 505 bp overlap
ERG 1 dataset
ChIP TSU-1621MT_ATRA GSE60477.ERG.TSU-1621MT_ATRA 140 bp overlap
ESRRG 1 dataset
ChIP cardiomyocyte GSE113760.ESRRG.cardiomyocyte 568 bp overlap
ETS1 1 dataset
ChIP cardiomyocyte_D9 GSE129986.ETS1.cardiomyocyte_D9 599 bp overlap
FOSL2 2 datasets
ChIP SK-N-SH ENCSR000BVB.FOSL2.SK-N-SH 118 bp overlap
ChIP hESC GSE69539.FOSL2.hESC 156 bp overlap
GATA1 1 dataset
ChIP CD34_Day7_30min GSE104676.GATA1.CD34_Day7_30min 55 bp overlap
GATA2 12 datasets
ChIP ESF GSE108408.GATA2.ESF 472 bp overlap
ChIP SH-SY5Y ENCFF485YIB 405 bp overlap
ChIP SH-SY5Y ENCSR000EYB.GATA2.SH-SY5Y 383 bp overlap
ChIP SK-N-SH ENCFF764OZD 417 bp overlap
ChIP SK-N-SH ENCSR706VOO.GATA2.SK-N-SH 256 bp overlap
ChIP WA09 GSE105081.GATA2.WA09 156 bp overlap
ChIP dermal-fibroblast_alone GSE51025.GATA2.dermal-fibroblast_alone 558 bp overlap
ChIP hiPSC_SLC6ebs GSE107639.GATA2.hiPSC_SLC6ebs 432 bp overlap
ChIP primary-endometrial-stromal-cell_IVD_P1 GSE145673.GATA2.primary-endometrial-stromal-cell_IVD_P1 506 bp overlap
ChIP primary-endometrial-stromal-cell_IVD_P2 GSE145673.GATA2.primary-endometrial-stromal-cell_IVD_P2 487 bp overlap
ChIP primary-endometrial-stromal-cell_Veh_P1 GSE145673.GATA2.primary-endometrial-stromal-cell_Veh_P1 528 bp overlap
ChIP primary-endometrial-stromal-cell_Veh_P2 GSE145673.GATA2.primary-endometrial-stromal-cell_Veh_P2 551 bp overlap
GATA3 2 datasets
ChIP CLB-Ga GSE90683.GATA3.CLB-Ga 255 bp overlap
ChIP SK-N-SH ENCFF040SSB 238 bp overlap
GATA4 11 datasets
ChIP DE DE-GATA4-1 526 bp overlap
ChIP DE DE-GATA4-2 564 bp overlap
ChIP G296S GSE85628.GATA4.G296S 463 bp overlap
ChIP G296S_2 GSE85628.GATA4.G296S_2 463 bp overlap
ChIP cardiomyocyte GSE85628.GATA4.cardiomyocyte 536 bp overlap
ChIP cardiomyocyte_1 GSE85628.GATA4.cardiomyocyte_1 537 bp overlap
ChIP cardiomyocyte_5 GSE85628.GATA4.cardiomyocyte_5 570 bp overlap
ChIP cardiomyocyte_7 GSE85628.GATA4.cardiomyocyte_7 583 bp overlap
ChIP foregut GSE117136.GATA4.foregut 309 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA4.pancreatic-progenitor 427 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA4.pancreatic-progenitor_PP1 545 bp overlap
GATA6 13 datasets
ChIP DE DE-GATA6-1 436 bp overlap
ChIP DE DE-GATA6-2 599 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 534 bp overlap
ChIP DE_D1 S41-DE-d1-GATA6-exp2 525 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 570 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 595 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 521 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 532 bp overlap
ChIP foregut GSE117136.GATA6.foregut 501 bp overlap
ChIP foregut_KO GSE117136.GATA6.foregut_KO 589 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA6.pancreatic-progenitor 333 bp overlap
ChIP pancreatic-progenitor_KO GSE117136.GATA6.pancreatic-progenitor_KO 599 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA6.pancreatic-progenitor_PP1 435 bp overlap
GLIS3 1 dataset
ChIP SK-N-SH ENCFF370MHZ 285 bp overlap
GPS2 1 dataset
ChIP hMADS_D0_E GSE152517.GPS2.hMADS_D0_E 422 bp overlap
HAND2 3 datasets
ChIP CLB-Ga GSE90683.HAND2.CLB-Ga 554 bp overlap
ChIP Kelly GSE94822.HAND2.Kelly 249 bp overlap
ChIP SK-N-BE2-C GSE94822.HAND2.SK-N-BE2-C 320 bp overlap
HSF1 2 datasets
ChIP MCF-7_hyperthermia-15min GSE137558.HSF1.MCF-7_hyperthermia-15min 322 bp overlap
ChIP NCI-H838 GSE38901.HSF1.NCI-H838 277 bp overlap
ISL1 1 dataset
ChIP SK-N-SH ENCFF285GEQ 277 bp overlap
JUND 2 datasets
ChIP SK-N-SH ENCFF551NEQ 321 bp overlap
ChIP SK-N-SH ENCSR000BSK.JUND.SK-N-SH 222 bp overlap
KDM1A 1 dataset
ChIP SH-SY5Y_B3 GSE58258.KDM1A.SH-SY5Y_B3 409 bp overlap
MAX 3 datasets
ChIP SK-N-SH ENCFF285LXR 384 bp overlap
ChIP SK-N-SH ENCSR000BVD.MAX.SK-N-SH 178 bp overlap
ChIP liver ENCSR847DIT.MAX.liver 162 bp overlap
MED1 11 datasets
ChIP SUM159PT_100nMtrametinib300nMJQ1_24h GSE87418.MED1.SUM159PT_100nMtrametinib300nMJQ1_24h 317 bp overlap
ChIP SUM159PT_100nMtrametinib_24h GSE87418.MED1.SUM159PT_100nMtrametinib_24h 211 bp overlap
ChIP hMSC-TERT4_4h GSE104537.MED1.hMSC-TERT4_4h 599 bp overlap
ChIP hMSC-TERT4_D14 GSE104537.MED1.hMSC-TERT4_D14 411 bp overlap
ChIP hMSC-TERT4_D3 GSE104537.MED1.hMSC-TERT4_D3 526 bp overlap
ChIP hMSC-TERT4_D7 GSE104537.MED1.hMSC-TERT4_D7 267 bp overlap
ChIP hMSC-TERT4_adipocyte-4H GSE113253.MED1.hMSC-TERT4_adipocyte-4H 599 bp overlap
ChIP hMSC-TERT4_adipocyte-D1 GSE113253.MED1.hMSC-TERT4_adipocyte-D1 599 bp overlap
ChIP hMSC-TERT4_adipocyte-D14 GSE113253.MED1.hMSC-TERT4_adipocyte-D14 203 bp overlap
ChIP hMSC-TERT4_adipocyte-D3 GSE113253.MED1.hMSC-TERT4_adipocyte-D3 215 bp overlap
ChIP hMSC-TERT4_adipocyte-D7 GSE113253.MED1.hMSC-TERT4_adipocyte-D7 376 bp overlap
MED12 1 dataset
ChIP myometrium_PT916 GSE128230.MED12.myometrium_PT916 107 bp overlap
MED26 1 dataset
ChIP HEK293T GSE121024.MED26.HEK293T 216 bp overlap
MXI1 2 datasets
ChIP SK-N-SH ENCFF746HVJ 415 bp overlap
ChIP SK-N-SH ENCFF746HVJ 419 bp overlap
MYCN 2 datasets
ChIP Kelly GSE94822.MYCN.Kelly 164 bp overlap
ChIP LA-N-5 GSE138295.MYCN.LA-N-5 331 bp overlap
NEUROG2 3 datasets
ChIP MRC-5_NFD_02DPT GSE75910.NEUROG2.MRC-5_NFD_02DPT 178 bp overlap
ChIP MRC-5_NFD_05DPT GSE75910.NEUROG2.MRC-5_NFD_05DPT 183 bp overlap
ChIP MRC-5_N_05DPT GSE75910.NEUROG2.MRC-5_N_05DPT 199 bp overlap
NFIC 1 dataset
ChIP SK-N-SH ENCSR000BSV.NFIC.SK-N-SH 214 bp overlap
NKX2-5 2 datasets
ChIP hESC_ab3584 GSE89457.NKX2-5.hESC_ab3584 509 bp overlap
ChIP hESC_sc-14033 GSE89457.NKX2-5.hESC_sc-14033 420 bp overlap
NR2F2 6 datasets
ChIP hiPSC GSE81585.NR2F2.hiPSC 350 bp overlap
ChIP liver ENCFF565JGD 281 bp overlap
ChIP liver ENCFF565JGD 373 bp overlap
ChIP liver ENCSR168SMX.NR2F2.liver 557 bp overlap
ChIP liver ENCSR338MMB.NR2F2.liver 335 bp overlap
ChIP uterus_STROMA_ENDOMETRIUM GSE52008.NR2F2.uterus_STROMA_ENDOMETRIUM 225 bp overlap
NR3C1 6 datasets
ChIP HCC1937 GSE152203.NR3C1.HCC1937 190 bp overlap
ChIP IMR-90 ERP007093.NR3C1.IMR-90 321 bp overlap
ChIP IMR-90_GLUCC ERP007081.NR3C1.IMR-90_GLUCC 155 bp overlap
ChIP MDA-MB-231 GSE152203.NR3C1.MDA-MB-231 296 bp overlap
ChIP SUM159PT GSE152203.NR3C1.SUM159PT 497 bp overlap
ChIP hMSC_DMI GSE68864.NR3C1.hMSC_DMI 264 bp overlap
PBX3 1 dataset
ChIP SK-N-SH ENCFF876BMC 203 bp overlap
PGR 6 datasets
ChIP hESC GSE69539.PGR.hESC 298 bp overlap
ChIP leiomyoma_RU486 GSE40724.PGR.leiomyoma_RU486 304 bp overlap
ChIP myometrium_NP1 GSE137550.PGR.myometrium_NP1 364 bp overlap
ChIP myometrium_NP2 GSE137550.PGR.myometrium_NP2 507 bp overlap
ChIP myometrium_TP1 GSE137550.PGR.myometrium_TP1 301 bp overlap
ChIP myometrium_TP2 GSE137550.PGR.myometrium_TP2 170 bp overlap
PGR_A 1 dataset
ChIP hESC GSE62475.PGR_A.hESC 284 bp overlap
PGR_B 1 dataset
ChIP hESC GSE62475.PGR_B.hESC 333 bp overlap
PHOX2B 1 dataset
ChIP CLB-Ga GSE90683.PHOX2B.CLB-Ga 369 bp overlap
PITX3 1 dataset
ChIP SH-SY5Y GSE93275.PITX3.SH-SY5Y 437 bp overlap
POLR2A 20 datasets
ChIP SK-N-SH ENCFF683PFH 134 bp overlap
ChIP breast epithelium ENCFF960NNA 176 bp overlap
ChIP esophagus muscularis mucosa ENCFF432ZCY 121 bp overlap
ChIP esophagus muscularis mucosa ENCFF759BBR 123 bp overlap
ChIP esophagus muscularis mucosa ENCFF791ZXN 360 bp overlap
ChIP gastroesophageal sphincter ENCFF070PCA 207 bp overlap
ChIP sigmoid colon ENCFF101ILL 321 bp overlap
ChIP sigmoid colon ENCFF725QFT 439 bp overlap
ChIP sigmoid colon ENCFF748YVT 431 bp overlap
ChIP sigmoid colon ENCFF754JQR 238 bp overlap
ChIP spleen ENCFF044PYR 269 bp overlap
ChIP spleen ENCFF446ZGT 326 bp overlap
ChIP stomach ENCFF607ZPU 325 bp overlap
ChIP stomach ENCFF820WZN 179 bp overlap
ChIP transverse colon ENCFF607LKE 277 bp overlap
ChIP transverse colon ENCFF610RWV 279 bp overlap
ChIP transverse colon ENCFF840PXT 321 bp overlap
ChIP upper lobe of left lung ENCFF095BJW 371 bp overlap
ChIP upper lobe of left lung ENCFF199JUI 405 bp overlap
ChIP vagina ENCFF384GAB 599 bp overlap
PPARA 1 dataset
ChIP SK-N-SH_all-trans-retinoic-acid_6-uM_48-hour ENCSR602QEJ.PPARA.SK-N-SH_all-trans-retinoic-acid_6-uM_48-hour 222 bp overlap
PPARG 1 dataset
ChIP ASC GSE21366.PPARG.ASC 314 bp overlap
PRDM14 1 dataset
ChIP primordial-germ-cell-like-cell GSE138674.PRDM14.primordial-germ-cell-like-cell 250 bp overlap
RAD21 1 dataset
ChIP SK-N-SH ENCSR000EHX.RAD21.SK-N-SH 182 bp overlap
RARA 1 dataset
ChIP hiPSC_D5 GSE132532.RARA.hiPSC_D5 235 bp overlap
RBPJ 1 dataset
ChIP SCC_4h GSE156486.RBPJ.SCC_4h 181 bp overlap
RCOR1 3 datasets
ChIP IMR-90 ENCSR000EFG.RCOR1.IMR-90 264 bp overlap
ChIP SK-N-SH ENCFF518EXB 245 bp overlap
ChIP SK-N-SH ENCSR009TKN.RCOR1.SK-N-SH 332 bp overlap
RELA 2 datasets
ChIP SGBS_TNF GSE64233.RELA.SGBS_TNF 206 bp overlap
ChIP aortic-endothelial-cell_IL1B_D21 GSE139377.RELA.aortic-endothelial-cell_IL1B_D21 302 bp overlap
RUNX1 1 dataset
ChIP H9_DOX-5_HA GSE137670.RUNX1.H9_DOX-5_HA 196 bp overlap
RXRA 2 datasets
ChIP liver ENCFF077DAP 465 bp overlap
ChIP liver ENCFF077DAP 370 bp overlap
SIN3A 2 datasets
ChIP SK-N-SH ENCFF931NFD 341 bp overlap
ChIP SK-N-SH ENCSR000BPB.SIN3A.SK-N-SH 180 bp overlap
SMAD2-3 1 dataset
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 559 bp overlap
SMARCC1 1 dataset
ChIP TTC-549_Dox GSE71504.SMARCC1.TTC-549_Dox 166 bp overlap
SMC3 1 dataset
ChIP SK-N-SH ENCFF791WFB 241 bp overlap
SNAI2 2 datasets
ChIP SK-N-SH ENCFF449PID 337 bp overlap
ChIP SK-N-SH_all-trans-retinoic-acid_6-uM_48-hour ENCSR106GVM.SNAI2.SK-N-SH_all-trans-retinoic-acid_6-uM_48-hour 307 bp overlap
SOX17_M 1 dataset
ChIP DE_D2 DED2-SOX17-M_Batch_II 233 bp overlap
SP1 2 datasets
ChIP liver ENCFF769YSM 506 bp overlap
ChIP liver ENCFF769YSM 404 bp overlap
STAT1 1 dataset
ChIP THP-1_IFNb GSE128111.STAT1.THP-1_IFNb 312 bp overlap
TBX5 5 datasets
ChIP G296S GSE85628.TBX5.G296S 390 bp overlap
ChIP G296S_2 GSE85628.TBX5.G296S_2 390 bp overlap
ChIP G296S_4 GSE85628.TBX5.G296S_4 190 bp overlap
ChIP cardiomyocyte_7 GSE85628.TBX5.cardiomyocyte_7 381 bp overlap
ChIP hiPSC GSE81585.TBX5.hiPSC 511 bp overlap
TCF12 2 datasets
ChIP SK-N-SH ENCFF147AHB 337 bp overlap
ChIP WA01 ENCSR000BIT.TCF12.WA01 135 bp overlap
TCF21 1 dataset
ChIP HCASMC GSE124011.TCF21.HCASMC 372 bp overlap
TCF4 2 datasets
ChIP SH-SY5Y GSE96915.TCF4.SH-SY5Y 342 bp overlap
ChIP SK-N-SH ENCFF270OWF 403 bp overlap
TEAD1 1 dataset
ChIP adipocyte GSE140782.TEAD1.adipocyte 404 bp overlap
TEAD4 2 datasets
ChIP BJ_fibroblast_CD13-neg GSE114367.TEAD4.BJ_fibroblast_CD13-neg 250 bp overlap
ChIP SK-N-SH ENCSR000BUQ.TEAD4.SK-N-SH 186 bp overlap
TFAP2A 1 dataset
ChIP WA09 GSE105081.TFAP2A.WA09 432 bp overlap
TFAP2B 1 dataset
ChIP SK-N-SH ENCFF869XXQ 277 bp overlap
TSHZ2 2 datasets
ChIP SK-N-SH ENCFF182EBB 381 bp overlap
ChIP SK-N-SH_all-trans-retinoic-acid_6-uM_48-hour ENCSR282NLQ.TSHZ2.SK-N-SH_all-trans-retinoic-acid_6-uM_48-hour 370 bp overlap
TWIST1 4 datasets
ChIP SHEP-21N GSE80151.TWIST1.SHEP-21N 274 bp overlap
ChIP SHEP-21N GSE80151.TWIST1.SHEP-21N 251 bp overlap
ChIP SHEP-21N_24h GSE80151.TWIST1.SHEP-21N_24h 230 bp overlap
ChIP SHEP-21N_DOX_0H GSE80151.TWIST1.SHEP-21N_DOX_0H 274 bp overlap
YAP1 1 dataset
ChIP hiPSC GSE111930.YAP1.hiPSC 123 bp overlap
YY1 1 dataset
ChIP SK-N-SH ENCSR000BLZ.YY1.SK-N-SH 136 bp overlap
ZFP3 2 datasets
ChIP SK-N-SH ENCFF981MBE 439 bp overlap
ChIP SK-N-SH_all-trans-retinoic-acid_6-uM_48-hour ENCSR581GUY.ZFP3.SK-N-SH_all-trans-retinoic-acid_6-uM_48-hour 394 bp overlap
ZNF257 1 dataset
ChIP HEK293T GSE78099.ZNF257.HEK293T 116 bp overlap
ZNF283 1 dataset
ChIP HEK293T GSE78099.ZNF283.HEK293T 157 bp overlap
ZNF565 2 datasets
ChIP SK-N-SH ENCFF372UGG 277 bp overlap
ChIP SK-N-SH ENCFF372UGG 212 bp overlap