chr4 : 134,149,104 134,150,162
1,058 bp 171 TFs 0 linked genes
This 1.1 kb open chromatin element has no linked target genes and is bound by 171 transcription factors.
Linked Genes
No linked genes
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr4:134,144,104 – 134,155,162
This element Other elements ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
171 transcription factors
Source
Cell type
Arid5a 1 dataset
Motif DE_12h DE_12h-Arid5a_MA0602.2 8 bp overlap
Arx 1 dataset
Motif DE_12h DE_12h-Arx_MA0874.2 10 bp overlap
BARX1 1 dataset
Motif DE_12h DE_12h-BARX1_MA0875.2 6 bp overlap
BSX 1 dataset
Motif DE_12h DE_12h-BSX_MA0876.2 6 bp overlap
CHD7 1 dataset
ChIP hiPSC_derived_lt-NES GSE108506.CHD7.hiPSC_derived_lt-NES 322 bp overlap
CTNNB1 2 datasets
ChIP hESC_YAP-_activinA_15h GSE99202.CTNNB1.hESC_YAP-_activinA_15h 502 bp overlap
ChIP hiPSC_D2 GSE132532.CTNNB1.hiPSC_D2 253 bp overlap
CUX1 1 dataset
Motif DE_12h DE_12h-CUX1_MA0754.3 9 bp overlap
CUX2 1 dataset
Motif DE_12h DE_12h-CUX2_MA0755.2 9 bp overlap
Cebpa 1 dataset
ChIP BLaER1 ENCFF460KDD 120 bp overlap
DBP 1 dataset
Motif DE_12h DE_12h-DBP_MA0639.2 10 bp overlap
DLX1 1 dataset
Motif DE_12h DE_12h-DLX1_MA0879.3 6 bp overlap
DLX6 1 dataset
Motif DE_12h DE_12h-DLX6_MA0882.2 6 bp overlap
DUX4 1 dataset
Motif DE_12h DE_12h-DUX4_MA0468.1 11 bp overlap
DUXA 1 dataset
Motif DE_12h DE_12h-DUXA_MA0884.2 13 bp overlap
Dlx2 1 dataset
Motif DE_12h DE_12h-Dlx2_MA0885.3 8 bp overlap
Dlx3 1 dataset
Motif DE_12h DE_12h-Dlx3_MA0880.2 6 bp overlap
Dlx4 1 dataset
Motif DE_12h DE_12h-Dlx4_MA0881.2 6 bp overlap
Dlx5 1 dataset
Motif DE_12h DE_12h-Dlx5_MA1476.3 8 bp overlap
Dux 1 dataset
Motif DE_12h DE_12h-Dux_MA0611.3 11 bp overlap
EN2 1 dataset
Motif DE_12h DE_12h-EN2_MA0642.3 7 bp overlap
ESR1 2 datasets
ChIP endometrioid-adenocarcinoma_tumor_2 GSE94031.ESR1.endometrioid-adenocarcinoma_tumor_2 302 bp overlap
ChIP primary-endometrium-cancer_E1_DSG GSE114737.ESR1.primary-endometrium-cancer_E1_DSG 369 bp overlap
Ebf4 1 dataset
Motif DE_12h DE_12h-Ebf4_MA2122.1 11 bp overlap
Erg 1 dataset
Motif DE_12h DE_12h-Erg_MA0474.4 10 bp overlap
FIGLA 1 dataset
Motif DE_12h DE_12h-FIGLA_MA0820.2 6 bp overlap
FOXA1 1 dataset
Motif DE_12h DE_12h-FOXA1_MA0148.5 8 bp overlap
FOXA2 4 datasets
ChIP DE DE-FOXA2-1 515 bp overlap
ChIP DE DE-FOXA2-2 449 bp overlap
Motif DE_12h DE_12h-FOXA2_MA0047.4 8 bp overlap
Motif DE_12h DE_12h-FOXA2_MA0047.4 8 bp overlap
FOXA3 2 datasets
Motif DE_12h DE_12h-FOXA3_MA1683.2 7 bp overlap
Motif DE_12h DE_12h-FOXA3_MA1683.2 7 bp overlap
FOXB1 2 datasets
Motif DE_12h DE_12h-FOXB1_MA0845.1 11 bp overlap
Motif DE_12h DE_12h-FOXB1_MA0845.1 11 bp overlap
FOXC1 1 dataset
Motif DE_12h DE_12h-FOXC1_MA0032.2 11 bp overlap
FOXC2 1 dataset
Motif DE_12h DE_12h-FOXC2_MA0846.2 11 bp overlap
FOXD1 1 dataset
Motif DE_12h DE_12h-FOXD1_MA0031.2 7 bp overlap
FOXD2 1 dataset
Motif DE_12h DE_12h-FOXD2_MA0847.4 11 bp overlap
FOXF2 1 dataset
Motif DE_12h DE_12h-FOXF2_MA0030.2 9 bp overlap
FOXG1 1 dataset
Motif DE_12h DE_12h-FOXG1_MA0613.1 8 bp overlap
FOXH1 1 dataset
Motif DE_12h DE_12h-FOXH1_MA0479.2 8 bp overlap
FOXI1 2 datasets
Motif DE_12h DE_12h-FOXI1_MA0042.2 7 bp overlap
Motif DE_12h DE_12h-FOXI1_MA0042.2 7 bp overlap
FOXK1 1 dataset
Motif DE_12h DE_12h-FOXK1_MA0852.3 7 bp overlap
FOXK2 1 dataset
Motif DE_12h DE_12h-FOXK2_MA1103.3 7 bp overlap
FOXL1 1 dataset
Motif DE_12h DE_12h-FOXL1_MA0033.2 7 bp overlap
FOXN3 3 datasets
Motif DE_12h DE_12h-FOXN3_MA1489.1 8 bp overlap
Motif DE_12h DE_12h-FOXN3_MA1489.1 8 bp overlap
Motif DE_12h DE_12h-FOXN3_MA1489.1 8 bp overlap
FOXO4 1 dataset
Motif DE_12h DE_12h-FOXO4_MA0848.1 7 bp overlap
FOXO6 1 dataset
Motif DE_12h DE_12h-FOXO6_MA0849.1 7 bp overlap
FOXP1 2 datasets
Motif DE_12h DE_12h-FOXP1_MA0481.4 7 bp overlap
Motif DE_12h DE_12h-FOXP1_MA0481.4 7 bp overlap
FOXP2 2 datasets
Motif DE_12h DE_12h-FOXP2_MA0593.2 9 bp overlap
Motif DE_12h DE_12h-FOXP2_MA0593.2 9 bp overlap
FOXP3 1 dataset
Motif DE_12h DE_12h-FOXP3_MA0850.1 7 bp overlap
FOXP4 2 datasets
Motif DE_12h DE_12h-FOXP4_MA2117.1 7 bp overlap
Motif DE_12h DE_12h-FOXP4_MA2117.1 7 bp overlap
FOXS1 1 dataset
Motif DE_12h DE_12h-FOXS1_MA2118.1 8 bp overlap
Foxf1 1 dataset
Motif DE_12h DE_12h-Foxf1_MA1606.2 7 bp overlap
Foxj2 1 dataset
Motif DE_12h DE_12h-Foxj2_MA0614.1 8 bp overlap
Foxj3 1 dataset
Motif DE_12h DE_12h-Foxj3_MA0851.2 9 bp overlap
Foxl2 3 datasets
Motif DE_12h DE_12h-Foxl2_MA1607.2 10 bp overlap
Motif DE_12h DE_12h-Foxl2_MA1607.2 10 bp overlap
Motif DE_12h DE_12h-Foxl2_MA1607.2 10 bp overlap
Foxo1 1 dataset
Motif DE_12h DE_12h-Foxo1_MA0480.3 7 bp overlap
Foxo3 1 dataset
Motif DE_12h DE_12h-Foxo3_MA0157.4 7 bp overlap
Foxq1 2 datasets
Motif DE_12h DE_12h-Foxq1_MA0040.2 10 bp overlap
Motif DE_12h DE_12h-Foxq1_MA0040.2 10 bp overlap
GABPA 1 dataset
Motif DE_12h DE_12h-GABPA_MA0062.4 10 bp overlap
GATA1 1 dataset
Motif DE_12h DE_12h-GATA1_MA0035.5 7 bp overlap
GATA6 7 datasets
ChIP DE DE-GATA6-2 300 bp overlap
ChIP DE_D1 S09-DE-d1-GATA6-exp1 588 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 697 bp overlap
ChIP DE_D1 S41-DE-d1-GATA6-exp2 557 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 247 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 771 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 600 bp overlap
GBX1 1 dataset
Motif DE_12h DE_12h-GBX1_MA0889.2 7 bp overlap
GBX2 1 dataset
Motif DE_12h DE_12h-GBX2_MA0890.2 6 bp overlap
GRHL2 1 dataset
Motif DE_12h DE_12h-GRHL2_MA1105.3 8 bp overlap
HESX1 1 dataset
Motif DE_12h DE_12h-HESX1_MA0894.2 6 bp overlap
HLF 1 dataset
Motif DE_12h DE_12h-HLF_MA0043.4 9 bp overlap
HNF1A 1 dataset
Motif DE_12h DE_12h-HNF1A_MA0046.3 13 bp overlap
HNF1B 1 dataset
Motif DE_12h DE_12h-HNF1B_MA0153.2 13 bp overlap
HNF4A 1 dataset
Motif DE_12h DE_12h-HNF4A_MA1494.2 14 bp overlap
HOXA7 1 dataset
Motif DE_12h DE_12h-HOXA7_MA1498.3 6 bp overlap
HOXC11 1 dataset
Motif DE_12h DE_12h-HOXC11_MA0651.3 11 bp overlap
HOXC12 1 dataset
Motif DE_12h DE_12h-HOXC12_MA0906.2 10 bp overlap
Hnf1A 1 dataset
Motif DE_12h DE_12h-Hnf1A_MA1991.2 10 bp overlap
IKZF1 1 dataset
Motif DE_12h DE_12h-IKZF1_MA1508.2 8 bp overlap
IRF3 1 dataset
Motif DE_12h DE_12h-IRF3_MA1418.2 17 bp overlap
JUN 2 datasets
ChIP DE_D1 S40-DE-d1-JUN-exp2 333 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.JUN.HUES-8_DE_ctrl 490 bp overlap
LBX1 1 dataset
Motif DE_12h DE_12h-LBX1_MA0618.2 7 bp overlap
LBX2 1 dataset
Motif DE_12h DE_12h-LBX2_MA0699.2 6 bp overlap
LEF1 1 dataset
ChIP hESC_WNT3A GSE64758.LEF1.hESC_WNT3A 230 bp overlap
LHX2 1 dataset
Motif DE_12h DE_12h-LHX2_MA0700.3 6 bp overlap
LHX9 1 dataset
Motif DE_12h DE_12h-LHX9_MA0701.3 7 bp overlap
LMO2 1 dataset
ChIP CCRF-CEM GSE33850.LMO2.CCRF-CEM 411 bp overlap
Lef1 1 dataset
Motif DE_12h DE_12h-Lef1_MA0768.3 8 bp overlap
Lhx3 1 dataset
Motif DE_12h DE_12h-Lhx3_MA0135.2 12 bp overlap
MEIS1 1 dataset
Motif DE_12h DE_12h-MEIS1_MA0498.3 5 bp overlap
MSX1 1 dataset
Motif DE_12h DE_12h-MSX1_MA0666.3 6 bp overlap
MSX2 1 dataset
Motif DE_12h DE_12h-MSX2_MA0708.3 6 bp overlap
MYB 1 dataset
Motif DE_12h DE_12h-MYB_MA0100.4 6 bp overlap
Msx3 1 dataset
Motif DE_12h DE_12h-Msx3_MA0709.2 6 bp overlap
NANOG 6 datasets
ChIP HUES-8 GSE109524.NANOG.HUES-8 455 bp overlap
ChIP WA01 ERP004238.NANOG.WA01 221 bp overlap
ChIP WA01_3IL ERP004238.NANOG.WA01_3IL 773 bp overlap
ChIP WA09 GSE105028.NANOG.WA09 509 bp overlap
ChIP WA09_heat-shock GSE105028.NANOG.WA09_heat-shock 620 bp overlap
ChIP hESC GSE18292.NANOG.hESC 195 bp overlap
NCOR1 1 dataset
ChIP LS180_125 GSE39277.NCOR1.LS180_125 170 bp overlap
NFATC3 1 dataset
Motif DE_12h DE_12h-NFATC3_MA0625.3 6 bp overlap
NFATC4 1 dataset
Motif DE_12h DE_12h-NFATC4_MA1525.3 9 bp overlap
NFIL3 1 dataset
Motif DE_12h DE_12h-NFIL3_MA0025.3 9 bp overlap
NIPBL 1 dataset
ChIP hESC_WNT3A_ACTIVIN GSE64758.NIPBL.hESC_WNT3A_ACTIVIN 248 bp overlap
NR1I3 1 dataset
Motif DE_12h DE_12h-NR1I3_MA1534.2 8 bp overlap
NR2F1 1 dataset
Motif DE_12h DE_12h-NR2F1_MA0017.3 12 bp overlap
Nfatc1 1 dataset
Motif DE_12h DE_12h-Nfatc1_MA0624.3 6 bp overlap
Nfatc2 1 dataset
Motif DE_12h DE_12h-Nfatc2_MA0152.3 8 bp overlap
Nobox 1 dataset
Motif DE_12h DE_12h-Nobox_MA0125.2 6 bp overlap
ONECUT1 4 datasets
Motif DE_12h DE_12h-ONECUT1_MA0679.3 9 bp overlap
ChIP H9 ERP004206.ONECUT1.H9 433 bp overlap
ChIP H9 ERP004206.ONECUT1.H9 238 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.ONECUT1.pancreatic-progenitor_PP1 768 bp overlap
ONECUT3 1 dataset
Motif DE_12h DE_12h-ONECUT3_MA0757.2 12 bp overlap
PAX1 1 dataset
Motif DE_12h DE_12h-PAX1_MA0779.2 16 bp overlap
PAX3 1 dataset
Motif DE_12h DE_12h-PAX3_MA0780.1 10 bp overlap
PAX8 1 dataset
Motif DE_12h DE_12h-PAX8_MA2094.1 16 bp overlap
PAX9 1 dataset
Motif DE_12h DE_12h-PAX9_MA0781.2 16 bp overlap
PBX2 1 dataset
Motif DE_12h DE_12h-PBX2_MA1113.3 9 bp overlap
PBX3 1 dataset
Motif DE_12h DE_12h-PBX3_MA1114.2 11 bp overlap
PHOX2B 1 dataset
Motif DE_12h DE_12h-PHOX2B_MA0681.3 12 bp overlap
PKNOX1 2 datasets
Motif DE_12h DE_12h-PKNOX1_MA0782.3 10 bp overlap
ChIP HEK293T ENCSR233FAG.PKNOX1.HEK293T 375 bp overlap
POU1F1 1 dataset
Motif DE_12h DE_12h-POU1F1_MA0784.3 14 bp overlap
POU2F1 1 dataset
Motif DE_12h DE_12h-POU2F1_MA0785.2 9 bp overlap
POU2F2 2 datasets
Motif DE_12h DE_12h-POU2F2_MA0507.3 13 bp overlap
Motif DE_12h DE_12h-POU2F2_MA0507.3 13 bp overlap
POU2F3 1 dataset
Motif DE_12h DE_12h-POU2F3_MA0627.3 9 bp overlap
POU3F1 1 dataset
Motif DE_12h DE_12h-POU3F1_MA0786.2 10 bp overlap
POU3F2 2 datasets
Motif DE_12h DE_12h-POU3F2_MA0787.1 12 bp overlap
Motif DE_12h DE_12h-POU3F2_MA0787.1 12 bp overlap
POU3F3 2 datasets
Motif DE_12h DE_12h-POU3F3_MA0788.1 13 bp overlap
Motif DE_12h DE_12h-POU3F3_MA0788.1 13 bp overlap
POU3F4 1 dataset
Motif DE_12h DE_12h-POU3F4_MA0789.1 9 bp overlap
POU4F1 1 dataset
Motif DE_12h DE_12h-POU4F1_MA0790.2 12 bp overlap
POU4F3 1 dataset
Motif DE_12h DE_12h-POU4F3_MA0791.2 12 bp overlap
POU5F1 7 datasets
Motif DE_12h DE_12h-POU5F1_MA1115.2 7 bp overlap
Motif DE_12h DE_12h-POU5F1_MA1115.2 7 bp overlap
ChIP HUES-8 GSE109524.POU5F1.HUES-8 355 bp overlap
ChIP WA09 GSE105028.POU5F1.WA09 234 bp overlap
ChIP WA09_heat-shock GSE105028.POU5F1.WA09_heat-shock 333 bp overlap
ChIP hESC ENCSR264RJX.POU5F1.hESC 122 bp overlap
ChIP hESC_PRIMED GSE69646.POU5F1.hESC_PRIMED 550 bp overlap
PPARD 1 dataset
Motif DE_12h DE_12h-PPARD_MA1550.2 14 bp overlap
PROP1 1 dataset
Motif DE_12h DE_12h-PROP1_MA0715.1 11 bp overlap
PRRX2 1 dataset
Motif DE_12h DE_12h-PRRX2_MA0075.4 7 bp overlap
Pax7 2 datasets
Motif DE_12h DE_12h-Pax7_MA0680.3 10 bp overlap
Motif DE_12h DE_12h-Pax7_MA0680.3 10 bp overlap
Pou5f1::Sox2 2 datasets
Motif DE_12h DE_12h-Pou5f1Sox2_MA0142.1 15 bp overlap
Motif DE_12h DE_12h-Pou5f1Sox2_MA0142.1 15 bp overlap
Prdm4 1 dataset
Motif DE_12h DE_12h-Prdm4_MA1647.3 11 bp overlap
RAX 1 dataset
Motif DE_12h DE_12h-RAX_MA0718.2 6 bp overlap
RBPJ 1 dataset
Motif DE_12h DE_12h-RBPJ_MA1116.2 6 bp overlap
RFX5 1 dataset
Motif DE_12h DE_12h-RFX5_MA0510.3 14 bp overlap
RXRB 1 dataset
Motif DE_12h DE_12h-RXRB_MA0855.1 14 bp overlap
RXRG 1 dataset
Motif DE_12h DE_12h-RXRG_MA0856.1 14 bp overlap
Rarb 1 dataset
Motif DE_12h DE_12h-Rarb_MA0857.1 16 bp overlap
Rxra 1 dataset
Motif DE_12h DE_12h-Rxra_MA0512.2 14 bp overlap
SMAD2-3 3 datasets
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 298 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.SMAD2-3.HUES-8_DE_JNKi 893 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.SMAD2-3.HUES-8_DE_ctrl 873 bp overlap
SMAD2_3 3 datasets
ChIP DE_D1 S07-DE-d1-SMAD2_3-exp1 666 bp overlap
ChIP DE_D1 S12-DE-d1-SMAD2_3-exp1 855 bp overlap
ChIP DE_D1 S39-DE-d1-SMAD2_3-exp2 770 bp overlap
SMARCA4 3 datasets
ChIP WA09_heat-shock GSE105028.SMARCA4.WA09_heat-shock 297 bp overlap
ChIP hiPSC GSE124903.SMARCA4.hiPSC 190 bp overlap
ChIP hiPSC_WT GSE124903.SMARCA4.hiPSC_WT 176 bp overlap
SMARCC1 2 datasets
ChIP DE_D1 S15-DE-d1-BAF155-exp1 669 bp overlap
ChIP hiPSC_WT GSE124903.SMARCC1.hiPSC_WT 214 bp overlap
SOX13 1 dataset
Motif DE_12h DE_12h-SOX13_MA1120.2 7 bp overlap
SOX14 1 dataset
Motif DE_12h DE_12h-SOX14_MA1562.2 9 bp overlap
SOX2 2 datasets
Motif DE_12h DE_12h-SOX2_MA0143.5 7 bp overlap
ChIP HNSC GSE69479.SOX2.HNSC 341 bp overlap
SOX21 2 datasets
Motif DE_12h DE_12h-SOX21_MA0866.1 15 bp overlap
Motif DE_12h DE_12h-SOX21_MA0866.1 15 bp overlap
SP5 1 dataset
Motif DE_12h DE_12h-SP5_MA1965.2 6 bp overlap
SREBF1 1 dataset
Motif DE_12h DE_12h-SREBF1_MA0829.3 10 bp overlap
Sox1 2 datasets
Motif DE_12h DE_12h-Sox1_MA0870.1 15 bp overlap
Motif DE_12h DE_12h-Sox1_MA0870.1 15 bp overlap
Sox3 1 dataset
Motif DE_12h DE_12h-Sox3_MA0514.3 7 bp overlap
Spz1 2 datasets
Motif DE_12h DE_12h-Spz1_MA0111.1 11 bp overlap
Motif DE_12h DE_12h-Spz1_MA0111.1 11 bp overlap
Stat4 2 datasets
Motif DE_12h DE_12h-Stat4_MA0518.2 10 bp overlap
Motif DE_12h DE_12h-Stat4_MA0518.2 10 bp overlap
Stat5a 1 dataset
Motif DE_12h DE_12h-Stat5a_MA1624.2 9 bp overlap
Stat6 1 dataset
Motif DE_12h DE_12h-Stat6_MA0520.2 10 bp overlap
T 1 dataset
ChIP H9_ENDODERM GSE60606.T.H9_ENDODERM 248 bp overlap
TCF7 1 dataset
Motif DE_12h DE_12h-TCF7_MA0769.3 7 bp overlap
TCF7L1 1 dataset
Motif DE_12h DE_12h-TCF7L1_MA1421.1 12 bp overlap
TCF7L2 3 datasets
Motif DE_12h DE_12h-TCF7L2_MA0523.2 9 bp overlap
Motif DE_12h DE_12h-TCF7L2_MA0523.2 9 bp overlap
Motif DE_12h DE_12h-TCF7L2_MA0523.2 9 bp overlap
THRB 1 dataset
Motif DE_12h DE_12h-THRB_MA1574.2 13 bp overlap
TP53 2 datasets
Motif DE_12h DE_12h-TP53_MA0106.3 18 bp overlap
ChIP H9_ectoderm GSE142050.TP53.H9_ectoderm 566 bp overlap
VENTX 2 datasets
Motif DE_12h DE_12h-VENTX_MA0724.1 9 bp overlap
Motif DE_12h DE_12h-VENTX_MA0724.1 9 bp overlap
ZBTB24 1 dataset
Motif DE_12h DE_12h-ZBTB24_MA2330.1 10 bp overlap
ZEB1 1 dataset
Motif DE_12h DE_12h-ZEB1_MA0103.4 6 bp overlap
ZNF157 1 dataset
Motif DE_12h DE_12h-ZNF157_MA2331.1 21 bp overlap
ZNF24 1 dataset
Motif DE_12h DE_12h-ZNF24_MA1124.1 13 bp overlap
ZNF547 1 dataset
Motif DE_12h DE_12h-ZNF547_MA2334.1 13 bp overlap
ZNF558 1 dataset
Motif DE_12h DE_12h-ZNF558_MA2335.1 29 bp overlap
ZNF667 1 dataset
Motif DE_12h DE_12h-ZNF667_MA1984.2 11 bp overlap
ZNF675 2 datasets
Motif DE_12h DE_12h-ZNF675_MA1714.2 19 bp overlap
Motif DE_12h DE_12h-ZNF675_MA1714.2 19 bp overlap
ZNF680 1 dataset
Motif DE_12h DE_12h-ZNF680_MA1729.2 11 bp overlap
ZNF766 1 dataset
Motif DE_12h DE_12h-ZNF766_MA2098.1 9 bp overlap
Zfp335 1 dataset
Motif DE_12h DE_12h-Zfp335_MA2002.2 7 bp overlap