chr1 : 79,044,973 – 79,045,812
839 bp 145 TFs 0 linked genes
This 839 bp open chromatin element has no linked target genes and is bound by 145 transcription factors.
Linked Genes
No linked genes
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr1:79,039,973 – 79,050,812
This element Other elements ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
145 transcription factors
Source
Cell type
ARGFX 4 datasets
Motif DE_12h DE_12h-ARGFX_MA1463.2 8 bp overlap
Motif DE_36h DE_36h-ARGFX_MA1463.2 8 bp overlap
Motif DE_48h DE_48h-ARGFX_MA1463.2 8 bp overlap
Motif ES_0h ES_0h-ARGFX_MA1463.2 8 bp overlap
Atf3 1 dataset
Motif DE_12h DE_12h-Atf3_MA1988.2 7 bp overlap
Atoh1 3 datasets
Motif DE_12h DE_12h-Atoh1_MA1467.3 7 bp overlap
Motif DE_36h DE_36h-Atoh1_MA1467.3 7 bp overlap
Motif ES_0h ES_0h-Atoh1_MA1467.3 7 bp overlap
BACH1 1 dataset
Motif DE_12h DE_12h-BACH1_MA1633.2 9 bp overlap
BACH2 1 dataset
Motif DE_12h DE_12h-BACH2_MA1101.3 11 bp overlap
BARX1 4 datasets
Motif DE_12h DE_12h-BARX1_MA0875.2 6 bp overlap
Motif DE_36h DE_36h-BARX1_MA0875.2 6 bp overlap
Motif DE_48h DE_48h-BARX1_MA0875.2 6 bp overlap
Motif ES_0h ES_0h-BARX1_MA0875.2 6 bp overlap
BATF 1 dataset
Motif DE_12h DE_12h-BATF_MA1634.2 7 bp overlap
BATF3 1 dataset
Motif DE_12h DE_12h-BATF3_MA0835.3 7 bp overlap
BATF::JUN 1 dataset
Motif DE_12h DE_12h-BATFJUN_MA0462.3 7 bp overlap
BCL11A 3 datasets
Motif DE_12h DE_12h-BCL11A_MA2324.1 7 bp overlap
Motif DE_36h DE_36h-BCL11A_MA2324.1 7 bp overlap
Motif ES_0h ES_0h-BCL11A_MA2324.1 7 bp overlap
BCL6 2 datasets
Motif DE_12h DE_12h-BCL6_MA0463.3 13 bp overlap
Motif ES_0h ES_0h-BCL6_MA0463.3 13 bp overlap
BNC2 1 dataset
Motif DE_12h DE_12h-BNC2_MA1928.2 7 bp overlap
BRD4 3 datasets
ChIP 402-91 GSE111253.BRD4.402-91 269 bp overlap
ChIP HEK293_sgNT GSE129407.BRD4.HEK293_sgNT 279 bp overlap
ChIP SUM1315_DMSO GSE63581.BRD4.SUM1315_DMSO 678 bp overlap
BSX 4 datasets
Motif DE_12h DE_12h-BSX_MA0876.2 6 bp overlap
Motif DE_36h DE_36h-BSX_MA0876.2 6 bp overlap
Motif DE_48h DE_48h-BSX_MA0876.2 6 bp overlap
Motif ES_0h ES_0h-BSX_MA0876.2 6 bp overlap
CDK6 1 dataset
ChIP KB_IL GSE52469.CDK6.KB_IL 100 bp overlap
CTCF 170 datasets
ChIP 22Rv1_hydroxy_10nM_4h ENCSR847XGE.CTCF.22Rv1_hydroxy_10nM_4h 341 bp overlap
ChIP 786-O_NORMOXIA GSE78113.CTCF.786-O_NORMOXIA 226 bp overlap
ChIP A-549 ENCSR000AUF.CTCF.A-549 381 bp overlap
ChIP A-673 ENCSR611JJS.CTCF.A-673 463 bp overlap
ChIP A673 ENCFF123WOM 227 bp overlap
ChIP BC-3 GSE135740.CTCF.BC-3 468 bp overlap
ChIP BE2C ENCFF757SRF 317 bp overlap
ChIP C4-2B ENCFF821XVN 660 bp overlap
ChIP C4-2B ENCFF821XVN 819 bp overlap
ChIP D54 ENCSR000DKN.CTCF.D54 239 bp overlap
Motif DE_12h DE_12h-CTCF_MA0139.2 15 bp overlap
Motif DE_48h DE_48h-CTCF_MA0139.2 15 bp overlap
Motif ES_0h ES_0h-CTCF_MA0139.2 15 bp overlap
ChIP GM04648 GSE148179.CTCF.GM04648 384 bp overlap
ChIP GM09236__B-lymphocytes GSE111170.CTCF.GM09236__B-lymphocytes 150 bp overlap
ChIP GM09237_B-lymphocytes GSE111170.CTCF.GM09237_B-lymphocytes 151 bp overlap
ChIP GM10266 ENCFF241YYF 177 bp overlap
ChIP GM12866 ENCSR000DRF.CTCF.GM12866 100 bp overlap
ChIP GM12870 ENCSR000DRK.CTCF.GM12870 106 bp overlap
ChIP GM12874 ENCFF942MTD 261 bp overlap
ChIP GM12878 ENCFF485TGR 251 bp overlap
ChIP GM12878 ENCFF635MMB 217 bp overlap
ChIP GM12878 GSE76922.CTCF.GM12878 312 bp overlap
ChIP GM13977 ENCFF528ESQ 171 bp overlap
ChIP GM17942 GSE76922.CTCF.GM17942 377 bp overlap
ChIP GM23338 ENCFF531QOI 201 bp overlap
ChIP GM23338 ENCFF772DML 152 bp overlap
ChIP GP5D GSE51234.CTCF.GP5D 243 bp overlap
ChIP GSC23 GSE139416.CTCF.GSC23 700 bp overlap
ChIP H1 ENCFF230QSV 165 bp overlap
ChIP H1 ENCFF414GZI 217 bp overlap
ChIP H1 ENCFF764RHO 277 bp overlap
ChIP H54 ENCFF255TVO 127 bp overlap
ChIP H9 ENCFF152GTF 369 bp overlap
ChIP HAP1 GSE152721.CTCF.HAP1 170 bp overlap
ChIP HAP1_clone21 GSE152721.CTCF.HAP1_clone21 180 bp overlap
ChIP HAP1_clone21_TSS-CRISPR GSE152721.CTCF.HAP1_clone21_TSS-CRISPR 242 bp overlap
ChIP HCT-116 ENCSR240PRQ.CTCF.HCT-116 379 bp overlap
ChIP HCT-116 GSE131606.CTCF.HCT-116 571 bp overlap
ChIP HCT-116 ENCSR000DTO.CTCF.HCT-116 239 bp overlap
ChIP HCT-116_RAD21-mAC GSE104888.CTCF.HCT-116_RAD21-mAC 295 bp overlap
ChIP HCT-116_RAD21-mAC_500uM_auxin GSE104888.CTCF.HCT-116_RAD21-mAC_500uM_auxin 234 bp overlap
ChIP HCT116 ENCFF003KHP 421 bp overlap
ChIP HCT116 ENCFF209YMI 285 bp overlap
ChIP HEC-1-B_R1-mutation GSE140868.CTCF.HEC-1-B_R1-mutation 54 bp overlap
ChIP HEC-1-B_RF-insertion GSE140868.CTCF.HEC-1-B_RF-insertion 216 bp overlap
ChIP HEK293 ENCFF498RMM 261 bp overlap
ChIP HEK293 ENCFF821TIC 360 bp overlap
ChIP HEK293 ENCSR617IFZ.CTCF.HEK293 362 bp overlap
ChIP HeLa GSE126990.CTCF.HeLa 394 bp overlap
ChIP HeLa-Kyoto GSE102884.CTCF.HeLa-Kyoto 266 bp overlap
ChIP HeLa-Kyoto GSE138405.CTCF.HeLa-Kyoto 394 bp overlap
ChIP HeLa-Kyoto_PDS5-depleted GSE102884.CTCF.HeLa-Kyoto_PDS5-depleted 276 bp overlap
ChIP HeLa-Kyoto_WAPL-depleted GSE102884.CTCF.HeLa-Kyoto_WAPL-depleted 384 bp overlap
ChIP HeLa-Kyoto_WAPL_PDS-depleted GSE102884.CTCF.HeLa-Kyoto_WAPL_PDS-depleted 469 bp overlap
ChIP HeLa-S3 ENCFF255ASZ 425 bp overlap
ChIP HeLa-S3 ENCFF565UFR 161 bp overlap
ChIP HeLa-S3 ENCFF626XQK 251 bp overlap
ChIP HeLa-S3 ENCSR000AOA.CTCF.HeLa-S3 242 bp overlap
ChIP HeLa-S3 ENCSR000DUB.CTCF.HeLa-S3 176 bp overlap
ChIP HeLa-S3_biotin GSE108869.CTCF.HeLa-S3_biotin 232 bp overlap
ChIP HeLa-S3_shCtrl GSE137848.CTCF.HeLa-S3_shCtrl 301 bp overlap
ChIP HeLa-S3_unsynchro GSE108173.CTCF.HeLa-S3_unsynchro 347 bp overlap
ChIP HepG2 ENCFF348BUL 197 bp overlap
ChIP ID00014 GSE76922.CTCF.ID00014 393 bp overlap
ChIP ID00016 GSE76922.CTCF.ID00016 315 bp overlap
ChIP LNCAP ENCFF223HIG 508 bp overlap
ChIP LNCAP ENCFF700QXT 506 bp overlap
ChIP LNCaP ENCSR315NAC.CTCF.LNCaP 367 bp overlap
ChIP LNCaP clone FGC ENCFF519YVI 181 bp overlap
ChIP LNCaP clone FGC ENCFF957KCI 181 bp overlap
ChIP MCF-7 ENCFF139NQI 277 bp overlap
ChIP MCF-7 ENCFF162GNE 245 bp overlap
ChIP MCF-7 ENCFF198DQX 231 bp overlap
ChIP MCF-7 ENCFF210JUZ 136 bp overlap
ChIP MCF-7 ENCFF414SZG 191 bp overlap
ChIP MCF-7 ENCFF424NQR 191 bp overlap
ChIP MCF-7 ENCFF494VXA 231 bp overlap
ChIP MCF-7 ENCFF844STM 191 bp overlap
ChIP MCF-7 ENCFF954TUV 205 bp overlap
ChIP MCF-7 ENCSR560BUE.CTCF.MCF-7 362 bp overlap
ChIP MCF-7 GSE137216.CTCF.MCF-7 345 bp overlap
ChIP MCF-7 ENCSR000DWH.CTCF.MCF-7 300 bp overlap
ChIP MCF-7 ENCSR000DMR.CTCF.MCF-7 249 bp overlap
ChIP MCF-7 ENCSR000AHD.CTCF.MCF-7 133 bp overlap
ChIP MCF-7 ENCSR000DMS.CTCF.MCF-7 165 bp overlap
ChIP MCF-7 GSE70764.CTCF.MCF-7 97 bp overlap
ChIP MCF-7L_t0 GSE108787.CTCF.MCF-7L_t0 359 bp overlap
ChIP MCF-7L_t1 GSE108787.CTCF.MCF-7L_t1 460 bp overlap
ChIP MCF-7L_t16 GSE108787.CTCF.MCF-7L_t16 510 bp overlap
ChIP MCF-7L_t24 GSE108787.CTCF.MCF-7L_t24 525 bp overlap
ChIP MCF-7_E2 ERP000380.CTCF.MCF-7_E2 150 bp overlap
ChIP MCF-7_TAM ERP000783.CTCF.MCF-7_TAM 180 bp overlap
ChIP OCI-LY1 ENCFF455ESK 451 bp overlap
ChIP OCI-LY3 ENCFF939BYJ 301 bp overlap
ChIP OCI-Ly1 ENCSR072EUE.CTCF.OCI-Ly1 339 bp overlap
ChIP OCI-Ly3 ENCSR756ZKG.CTCF.OCI-Ly3 625 bp overlap
ChIP OVCAR-8 GSE70764.CTCF.OVCAR-8 226 bp overlap
ChIP PC-3 ENCFF487TUI 441 bp overlap
ChIP PC-3 ENCSR359LOD.CTCF.PC-3 342 bp overlap
ChIP RH4 GSE83726.CTCF.RH4 240 bp overlap
ChIP RWPE-2 ENCSR856JJB.CTCF.RWPE-2 417 bp overlap
ChIP RWPE2 ENCFF911IEE 573 bp overlap
ChIP SK-N-AS GSE101295.CTCF.SK-N-AS 439 bp overlap
ChIP SK-N-SH ENCFF731NJX 251 bp overlap
ChIP SU-DHL-6 ENCFF116KKR 510 bp overlap
ChIP SU-DHL-6 ENCFF116KKR 561 bp overlap
ChIP T-47D GSE111923.CTCF.T-47D 431 bp overlap
ChIP T-47D_NaCl-isotonic GSE111923.CTCF.T-47D_NaCl-isotonic 790 bp overlap
ChIP T-47D_triptolide GSE111923.CTCF.T-47D_triptolide 455 bp overlap
ChIP THP-1_Pam3csk-000m-Flavo-000m GSE103477.CTCF.THP-1_Pam3csk-000m-Flavo-000m 249 bp overlap
ChIP THP-1_Pam3csk-000m-Flavo-240m GSE103477.CTCF.THP-1_Pam3csk-000m-Flavo-240m 182 bp overlap
ChIP THP-1_Pam3csk-020m-Flavo-000m GSE103477.CTCF.THP-1_Pam3csk-020m-Flavo-000m 281 bp overlap
ChIP THP-1_Pam3csk-045m-Flavo-000m GSE103477.CTCF.THP-1_Pam3csk-045m-Flavo-000m 246 bp overlap
ChIP THP-1_Pam3csk-060m-Flavo-000m GSE103477.CTCF.THP-1_Pam3csk-060m-Flavo-000m 174 bp overlap
ChIP THP-1_Pam3csk-120m-Flavo-000m GSE103477.CTCF.THP-1_Pam3csk-120m-Flavo-000m 277 bp overlap
ChIP THP-1_Pam3csk-150m-Flavo-030m GSE103477.CTCF.THP-1_Pam3csk-150m-Flavo-030m 183 bp overlap
ChIP THP-1_Pam3csk-180m-Flavo-060m GSE103477.CTCF.THP-1_Pam3csk-180m-Flavo-060m 232 bp overlap
ChIP THP-1_Pam3csk-360m-Flavo-240m GSE103477.CTCF.THP-1_Pam3csk-360m-Flavo-240m 277 bp overlap
ChIP THP-1_siCtrl-eGFP-Pam3csk-0h GSE103477.CTCF.THP-1_siCtrl-eGFP-Pam3csk-0h 190 bp overlap
ChIP VCaP ENCFF858YQT 521 bp overlap
ChIP VCaP ENCSR265ARE.CTCF.VCaP 323 bp overlap
ChIP WA01 ENCSR000DLK.CTCF.WA01 109 bp overlap
ChIP WA01 ENCSR000BNH.CTCF.WA01 188 bp overlap
ChIP WA01_NEUR GSE52457.CTCF.WA01_NEUR 264 bp overlap
ChIP WERI-Rb-1 ENCFF349QKF 231 bp overlap
ChIP WERI-Rb-1 ENCSR000DXW.CTCF.WERI-Rb-1 117 bp overlap
ChIP astrocyte_spinal_cord ENCSR000DSU.CTCF.astrocyte_spinal_cord 111 bp overlap
ChIP brain ENCFF163BBN 430 bp overlap
ChIP brain ENCFF685VRG 431 bp overlap
ChIP brain microvascular endothelial cell ENCFF526VHE 281 bp overlap
ChIP cardiac muscle cell ENCFF777TNC 365 bp overlap
ChIP cardiac-muscle ENCSR713SXF.CTCF.cardiac-muscle 407 bp overlap
ChIP chondrocyte ENCFF134ORZ 514 bp overlap
ChIP choroid plexus epithelial cell ENCFF407YNR 277 bp overlap
ChIP choroid-plexus_epithelial ENCSR000DTL.CTCF.choroid-plexus_epithelial 309 bp overlap
ChIP delta-47 GSE70764.CTCF.delta-47 232 bp overlap
ChIP dorsolateral prefrontal cortex ENCFF282ONV 448 bp overlap
ChIP endodermal cell ENCFF471YCZ 327 bp overlap
ChIP endothelial cell of umbilical vein ENCFF455OQM 197 bp overlap
ChIP endothelial_brain-microvascular ENCSR000DTA.CTCF.endothelial_brain-microvascular 193 bp overlap
ChIP epithelial cell of prostate ENCFF086GTI 489 bp overlap
ChIP epithelial cell of proximal tubule ENCFF763ZKS 331 bp overlap
ChIP epithelial_proximal_tubule ENCSR000DXD.CTCF.epithelial_proximal_tubule 143 bp overlap
ChIP fibroblast of lung ENCFF356FDN 317 bp overlap
ChIP fibroblast of lung ENCFF356FDN 317 bp overlap
ChIP fibroblast of lung ENCFF505HVQ 261 bp overlap
ChIP fibroblast of pulmonary artery ENCFF742RSV 297 bp overlap
ChIP fibroblast of the aortic adventitia ENCFF639DMR 217 bp overlap
ChIP hESC_D0 GSE116862.CTCF.hESC_D0 175 bp overlap
ChIP hESC_D5 GSE116862.CTCF.hESC_D5 428 bp overlap
ChIP heart left ventricle ENCFF244ZHV 437 bp overlap
ChIP heart_left-ventricle ENCSR544APK.CTCF.heart_left-ventricle 201 bp overlap
ChIP hiPSC GSE115407.CTCF.hiPSC 275 bp overlap
ChIP hiPSC_IB7 GSE106870.CTCF.hiPSC_IB7 121 bp overlap
ChIP hiPSC_IV8 GSE106870.CTCF.hiPSC_IV8 202 bp overlap
ChIP hiPSC_failed-genome-editing GSE106870.CTCF.hiPSC_failed-genome-editing 149 bp overlap
ChIP hiPSC_parental-line GSE106870.CTCF.hiPSC_parental-line 185 bp overlap
ChIP left ventricle myocardium inferior ENCFF161DPW 471 bp overlap
ChIP leukemia_CEBPA-24h GSE142161.CTCF.leukemia_CEBPA-24h 190 bp overlap
ChIP neural progenitor cell ENCFF420RBO 267 bp overlap
ChIP neural_progenitor ENCSR125NBL.CTCF.neural_progenitor 383 bp overlap
ChIP neuron_bipolar_doxy_4d ENCSR619IUE.CTCF.neuron_bipolar_doxy_4d 252 bp overlap
ChIP osteocyte ENCFF929FPD 454 bp overlap
ChIP primary-glioblastoma_G523 GSE121601.CTCF.primary-glioblastoma_G523 156 bp overlap
ChIP primary-glioblastoma_G583 GSE121601.CTCF.primary-glioblastoma_G583 513 bp overlap
ChIP prostate_epithelial ENCSR196HOM.CTCF.prostate_epithelial 340 bp overlap
ChIP retina_Hu6 GSE137311.CTCF.retina_Hu6 87 bp overlap
ChIP spleen ENCSR482PMN.CTCF.spleen 189 bp overlap
ChIP type B pancreatic cell ENCFF910FNQ 334 bp overlap
CTCFL 1 dataset
ChIP Kelly_shLUC-res GSE115248.CTCFL.Kelly_shLUC-res 240 bp overlap
CTCF_s 1 dataset
ChIP HeLa-S3_biotin GSE108869.CTCF_s.HeLa-S3_biotin 199 bp overlap
Cebpa 3 datasets
ChIP BLaER1 ENCFF093OYK 262 bp overlap
ChIP BLaER1 ENCFF274GAT 290 bp overlap
ChIP BLaER1 ENCFF335XTP 308 bp overlap
DLX1 4 datasets
Motif DE_12h DE_12h-DLX1_MA0879.3 6 bp overlap
Motif DE_36h DE_36h-DLX1_MA0879.3 6 bp overlap
Motif DE_48h DE_48h-DLX1_MA0879.3 6 bp overlap
Motif ES_0h ES_0h-DLX1_MA0879.3 6 bp overlap
DLX6 4 datasets
Motif DE_12h DE_12h-DLX6_MA0882.2 6 bp overlap
Motif DE_36h DE_36h-DLX6_MA0882.2 6 bp overlap
Motif DE_48h DE_48h-DLX6_MA0882.2 6 bp overlap
Motif ES_0h ES_0h-DLX6_MA0882.2 6 bp overlap
Dlx2 4 datasets
Motif DE_12h DE_12h-Dlx2_MA0885.3 8 bp overlap
Motif DE_36h DE_36h-Dlx2_MA0885.3 8 bp overlap
Motif DE_48h DE_48h-Dlx2_MA0885.3 8 bp overlap
Motif ES_0h ES_0h-Dlx2_MA0885.3 8 bp overlap
Dlx3 4 datasets
Motif DE_12h DE_12h-Dlx3_MA0880.2 6 bp overlap
Motif DE_36h DE_36h-Dlx3_MA0880.2 6 bp overlap
Motif DE_48h DE_48h-Dlx3_MA0880.2 6 bp overlap
Motif ES_0h ES_0h-Dlx3_MA0880.2 6 bp overlap
Dlx4 4 datasets
Motif DE_12h DE_12h-Dlx4_MA0881.2 6 bp overlap
Motif DE_36h DE_36h-Dlx4_MA0881.2 6 bp overlap
Motif DE_48h DE_48h-Dlx4_MA0881.2 6 bp overlap
Motif ES_0h ES_0h-Dlx4_MA0881.2 6 bp overlap
Dlx5 4 datasets
Motif DE_12h DE_12h-Dlx5_MA1476.3 8 bp overlap
Motif DE_36h DE_36h-Dlx5_MA1476.3 8 bp overlap
Motif DE_48h DE_48h-Dlx5_MA1476.3 8 bp overlap
Motif ES_0h ES_0h-Dlx5_MA1476.3 8 bp overlap
E2F1 2 datasets
ChIP HeLa GSE22478.E2F1.HeLa 211 bp overlap
ChIP LNCaP_shRB GSE94958.E2F1.LNCaP_shRB 161 bp overlap
E2F6 1 dataset
ChIP WA01 ENCSR000BSI.E2F6.WA01 188 bp overlap
EGR2 1 dataset
Motif ES_0h ES_0h-EGR2_MA0472.2 11 bp overlap
EGR3 1 dataset
Motif ES_0h ES_0h-EGR3_MA0732.2 11 bp overlap
EGR4 1 dataset
Motif ES_0h ES_0h-EGR4_MA0733.2 11 bp overlap
ERG 1 dataset
ChIP SKNO-1 GSE23730.ERG.SKNO-1 376 bp overlap
ESR1 1 dataset
ChIP Ishikawa_siETV4-72h GSE129803.ESR1.Ishikawa_siETV4-72h 242 bp overlap
FOS 1 dataset
Motif DE_12h DE_12h-FOS_MA0476.2 8 bp overlap
FOS::JUN 1 dataset
Motif DE_12h DE_12h-FOSJUN_MA0099.4 9 bp overlap
FOS::JUNB 1 dataset
Motif DE_12h DE_12h-FOSJUNB_MA1134.2 9 bp overlap
FOS::JUND 1 dataset
Motif DE_12h DE_12h-FOSJUND_MA1141.2 9 bp overlap
FOSB::JUNB 1 dataset
Motif DE_12h DE_12h-FOSBJUNB_MA1135.2 9 bp overlap
FOSL1 1 dataset
Motif DE_12h DE_12h-FOSL1_MA0477.3 9 bp overlap
FOSL1::JUN 1 dataset
Motif DE_12h DE_12h-FOSL1JUN_MA1128.2 9 bp overlap
FOSL1::JUNB 1 dataset
Motif DE_12h DE_12h-FOSL1JUNB_MA1137.2 9 bp overlap
FOSL1::JUND 2 datasets
Motif DE_12h DE_12h-FOSL1JUND_MA1142.2 8 bp overlap
Motif DE_12h DE_12h-FOSL1JUND_MA1143.2 9 bp overlap
FOSL2 1 dataset
Motif DE_12h DE_12h-FOSL2_MA0478.2 10 bp overlap
FOSL2::JUN 1 dataset
Motif DE_12h DE_12h-FOSL2JUN_MA1130.2 9 bp overlap
FOSL2::JUNB 1 dataset
Motif DE_12h DE_12h-FOSL2JUNB_MA1138.2 9 bp overlap
FOSL2::JUND 1 dataset
Motif DE_12h DE_12h-FOSL2JUND_MA1144.2 9 bp overlap
FOXA1 3 datasets
Motif DE_12h DE_12h-FOXA1_MA0148.5 8 bp overlap
Motif DE_60h DE_60h-FOXA1_MA0148.5 8 bp overlap
Motif ES_0h ES_0h-FOXA1_MA0148.5 8 bp overlap
FOXA2 3 datasets
Motif DE_12h DE_12h-FOXA2_MA0047.4 8 bp overlap
Motif DE_60h DE_60h-FOXA2_MA0047.4 8 bp overlap
Motif ES_0h ES_0h-FOXA2_MA0047.4 8 bp overlap
FOXA3 3 datasets
Motif DE_12h DE_12h-FOXA3_MA1683.2 7 bp overlap
Motif DE_60h DE_60h-FOXA3_MA1683.2 7 bp overlap
Motif ES_0h ES_0h-FOXA3_MA1683.2 7 bp overlap
FOXD1 3 datasets
Motif DE_12h DE_12h-FOXD1_MA0031.2 7 bp overlap
Motif DE_60h DE_60h-FOXD1_MA0031.2 7 bp overlap
Motif ES_0h ES_0h-FOXD1_MA0031.2 7 bp overlap
FOXG1 3 datasets
Motif DE_12h DE_12h-FOXG1_MA0613.1 8 bp overlap
Motif DE_60h DE_60h-FOXG1_MA0613.1 8 bp overlap
Motif ES_0h ES_0h-FOXG1_MA0613.1 8 bp overlap
FOXI1 3 datasets
Motif DE_12h DE_12h-FOXI1_MA0042.2 7 bp overlap
Motif DE_60h DE_60h-FOXI1_MA0042.2 7 bp overlap
Motif ES_0h ES_0h-FOXI1_MA0042.2 7 bp overlap
FOXK1 3 datasets
Motif DE_12h DE_12h-FOXK1_MA0852.3 7 bp overlap
Motif DE_60h DE_60h-FOXK1_MA0852.3 7 bp overlap
Motif ES_0h ES_0h-FOXK1_MA0852.3 7 bp overlap
FOXK2 3 datasets
Motif DE_12h DE_12h-FOXK2_MA1103.3 7 bp overlap
Motif DE_60h DE_60h-FOXK2_MA1103.3 7 bp overlap
Motif ES_0h ES_0h-FOXK2_MA1103.3 7 bp overlap
FOXL1 3 datasets
Motif DE_12h DE_12h-FOXL1_MA0033.2 7 bp overlap
Motif DE_60h DE_60h-FOXL1_MA0033.2 7 bp overlap
Motif ES_0h ES_0h-FOXL1_MA0033.2 7 bp overlap
FOXO4 3 datasets
Motif DE_12h DE_12h-FOXO4_MA0848.1 7 bp overlap
Motif DE_60h DE_60h-FOXO4_MA0848.1 7 bp overlap
Motif ES_0h ES_0h-FOXO4_MA0848.1 7 bp overlap
FOXO6 3 datasets
Motif DE_12h DE_12h-FOXO6_MA0849.1 7 bp overlap
Motif DE_60h DE_60h-FOXO6_MA0849.1 7 bp overlap
Motif ES_0h ES_0h-FOXO6_MA0849.1 7 bp overlap
FOXP1 3 datasets
Motif DE_12h DE_12h-FOXP1_MA0481.4 7 bp overlap
Motif DE_60h DE_60h-FOXP1_MA0481.4 7 bp overlap
Motif ES_0h ES_0h-FOXP1_MA0481.4 7 bp overlap
FOXP2 3 datasets
Motif DE_12h DE_12h-FOXP2_MA0593.2 9 bp overlap
Motif DE_60h DE_60h-FOXP2_MA0593.2 9 bp overlap
Motif ES_0h ES_0h-FOXP2_MA0593.2 9 bp overlap
FOXP3 3 datasets
Motif DE_12h DE_12h-FOXP3_MA0850.1 7 bp overlap
Motif DE_60h DE_60h-FOXP3_MA0850.1 7 bp overlap
Motif ES_0h ES_0h-FOXP3_MA0850.1 7 bp overlap
FOXP4 3 datasets
Motif DE_12h DE_12h-FOXP4_MA2117.1 7 bp overlap
Motif DE_60h DE_60h-FOXP4_MA2117.1 7 bp overlap
Motif ES_0h ES_0h-FOXP4_MA2117.1 7 bp overlap
FOXS1 3 datasets
Motif DE_12h DE_12h-FOXS1_MA2118.1 8 bp overlap
Motif DE_60h DE_60h-FOXS1_MA2118.1 8 bp overlap
Motif ES_0h ES_0h-FOXS1_MA2118.1 8 bp overlap
Foxf1 3 datasets
Motif DE_12h DE_12h-Foxf1_MA1606.2 7 bp overlap
Motif DE_60h DE_60h-Foxf1_MA1606.2 7 bp overlap
Motif ES_0h ES_0h-Foxf1_MA1606.2 7 bp overlap
Foxj2 3 datasets
Motif DE_12h DE_12h-Foxj2_MA0614.1 8 bp overlap
Motif DE_60h DE_60h-Foxj2_MA0614.1 8 bp overlap
Motif ES_0h ES_0h-Foxj2_MA0614.1 8 bp overlap
Foxj3 3 datasets
Motif DE_12h DE_12h-Foxj3_MA0851.2 9 bp overlap
Motif DE_60h DE_60h-Foxj3_MA0851.2 9 bp overlap
Motif ES_0h ES_0h-Foxj3_MA0851.2 9 bp overlap
Foxl2 3 datasets
Motif DE_12h DE_12h-Foxl2_MA1607.2 10 bp overlap
Motif DE_60h DE_60h-Foxl2_MA1607.2 10 bp overlap
Motif ES_0h ES_0h-Foxl2_MA1607.2 10 bp overlap
Foxo1 3 datasets
Motif DE_12h DE_12h-Foxo1_MA0480.3 7 bp overlap
Motif DE_60h DE_60h-Foxo1_MA0480.3 7 bp overlap
Motif ES_0h ES_0h-Foxo1_MA0480.3 7 bp overlap
Foxo3 3 datasets
Motif DE_12h DE_12h-Foxo3_MA0157.4 7 bp overlap
Motif DE_60h DE_60h-Foxo3_MA0157.4 7 bp overlap
Motif ES_0h ES_0h-Foxo3_MA0157.4 7 bp overlap
GBX2 4 datasets
Motif DE_12h DE_12h-GBX2_MA0890.2 6 bp overlap
Motif DE_36h DE_36h-GBX2_MA0890.2 6 bp overlap
Motif DE_48h DE_48h-GBX2_MA0890.2 6 bp overlap
Motif ES_0h ES_0h-GBX2_MA0890.2 6 bp overlap
GLIS1 1 dataset
ChIP HEK293 ENCSR482BBZ.GLIS1.HEK293 203 bp overlap
GLIS2 3 datasets
ChIP HCT-116_5FU GSE125927.GLIS2.HCT-116_5FU 249 bp overlap
ChIP HEK293 ENCFF446EIF 438 bp overlap
ChIP HEK293 ENCSR535DIA.GLIS2.HEK293 337 bp overlap
GTF3C2 1 dataset
ChIP H9 GSE94418.GTF3C2.H9 211 bp overlap
HAND2 3 datasets
Motif DE_12h DE_12h-HAND2_MA1638.2 6 bp overlap
Motif DE_36h DE_36h-HAND2_MA1638.2 6 bp overlap
Motif ES_0h ES_0h-HAND2_MA1638.2 6 bp overlap
HESX1 4 datasets
Motif DE_12h DE_12h-HESX1_MA0894.2 6 bp overlap
Motif DE_36h DE_36h-HESX1_MA0894.2 6 bp overlap
Motif DE_48h DE_48h-HESX1_MA0894.2 6 bp overlap
Motif ES_0h ES_0h-HESX1_MA0894.2 6 bp overlap
HIC2 3 datasets
Motif DE_12h DE_12h-HIC2_MA0738.2 6 bp overlap
Motif DE_60h DE_60h-HIC2_MA0738.2 6 bp overlap
Motif ES_0h ES_0h-HIC2_MA0738.2 6 bp overlap
HNF4A 1 dataset
Motif ES_0h ES_0h-HNF4A_MA1494.2 14 bp overlap
HOXA7 4 datasets
Motif DE_12h DE_12h-HOXA7_MA1498.3 6 bp overlap
Motif DE_36h DE_36h-HOXA7_MA1498.3 6 bp overlap
Motif DE_48h DE_48h-HOXA7_MA1498.3 6 bp overlap
Motif ES_0h ES_0h-HOXA7_MA1498.3 6 bp overlap
Hic1 3 datasets
Motif DE_12h DE_12h-Hic1_MA0739.2 8 bp overlap
Motif DE_60h DE_60h-Hic1_MA0739.2 8 bp overlap
Motif ES_0h ES_0h-Hic1_MA0739.2 8 bp overlap
JDP2 1 dataset
Motif DE_12h DE_12h-JDP2_MA0655.1 9 bp overlap
JUN 1 dataset
ChIP MDA-MB-231 GSE112444.JUN.MDA-MB-231 241 bp overlap
JUN::JUNB 1 dataset
Motif DE_12h DE_12h-JUNJUNB_MA1132.2 8 bp overlap
JUNB 1 dataset
Motif DE_12h DE_12h-JUNB_MA0490.3 9 bp overlap
JUND 1 dataset
Motif DE_12h DE_12h-JUND_MA0491.3 9 bp overlap
Jun 1 dataset
Motif DE_12h DE_12h-Jun_MA0489.3 8 bp overlap
KDM4C 1 dataset
ChIP KYSE-150 GSE53938.KDM4C.KYSE-150 238 bp overlap
KLF17 2 datasets
Motif DE_12h DE_12h-KLF17_MA1514.2 14 bp overlap
Motif ES_0h ES_0h-KLF17_MA1514.2 14 bp overlap
KLF9 2 datasets
Motif DE_12h DE_12h-KLF9_MA1107.3 11 bp overlap
Motif ES_0h ES_0h-KLF9_MA1107.3 11 bp overlap
LBX2 4 datasets
Motif DE_12h DE_12h-LBX2_MA0699.2 6 bp overlap
Motif DE_36h DE_36h-LBX2_MA0699.2 6 bp overlap
Motif DE_48h DE_48h-LBX2_MA0699.2 6 bp overlap
Motif ES_0h ES_0h-LBX2_MA0699.2 6 bp overlap
LHX2 4 datasets
Motif DE_12h DE_12h-LHX2_MA0700.3 6 bp overlap
Motif DE_36h DE_36h-LHX2_MA0700.3 6 bp overlap
Motif DE_48h DE_48h-LHX2_MA0700.3 6 bp overlap
Motif ES_0h ES_0h-LHX2_MA0700.3 6 bp overlap
MAF::NFE2 1 dataset
Motif DE_12h DE_12h-MAFNFE2_MA0501.2 11 bp overlap
MAFG::NFE2L1 1 dataset
Motif DE_12h DE_12h-MAFGNFE2L1_MA0089.3 11 bp overlap
MAFK 1 dataset
Motif DE_12h DE_12h-MAFK_MA0496.4 10 bp overlap
MAX 1 dataset
ChIP WTC11 ENCFF223QFY 585 bp overlap
MEIS1 2 datasets
Motif DE_12h DE_12h-MEIS1_MA0498.3 5 bp overlap
Motif ES_0h ES_0h-MEIS1_MA0498.3 5 bp overlap
MGA 1 dataset
ChIP A-549_empty GSE112188.MGA.A-549_empty 280 bp overlap
MSX1 4 datasets
Motif DE_12h DE_12h-MSX1_MA0666.3 6 bp overlap
Motif DE_36h DE_36h-MSX1_MA0666.3 6 bp overlap
Motif DE_48h DE_48h-MSX1_MA0666.3 6 bp overlap
Motif ES_0h ES_0h-MSX1_MA0666.3 6 bp overlap
MSX2 4 datasets
Motif DE_12h DE_12h-MSX2_MA0708.3 6 bp overlap
Motif DE_36h DE_36h-MSX2_MA0708.3 6 bp overlap
Motif DE_48h DE_48h-MSX2_MA0708.3 6 bp overlap
Motif ES_0h ES_0h-MSX2_MA0708.3 6 bp overlap
MXI1 2 datasets
Motif DE_12h DE_12h-MXI1_MA1108.3 6 bp overlap
Motif ES_0h ES_0h-MXI1_MA1108.3 6 bp overlap
MYC 2 datasets
ChIP CD34 GSE85488.MYC.CD34 113 bp overlap
ChIP Jurkat GSE83777.MYC.Jurkat 113 bp overlap
Msx3 4 datasets
Motif DE_12h DE_12h-Msx3_MA0709.2 6 bp overlap
Motif DE_36h DE_36h-Msx3_MA0709.2 6 bp overlap
Motif DE_48h DE_48h-Msx3_MA0709.2 6 bp overlap
Motif ES_0h ES_0h-Msx3_MA0709.2 6 bp overlap
NEUROD1 3 datasets
Motif DE_12h DE_12h-NEUROD1_MA1109.2 8 bp overlap
Motif DE_36h DE_36h-NEUROD1_MA1109.2 8 bp overlap
Motif ES_0h ES_0h-NEUROD1_MA1109.2 8 bp overlap
NEUROG2 3 datasets
Motif DE_12h DE_12h-NEUROG2_MA1642.2 7 bp overlap
Motif DE_36h DE_36h-NEUROG2_MA1642.2 7 bp overlap
Motif ES_0h ES_0h-NEUROG2_MA1642.2 7 bp overlap
NFE2 1 dataset
Motif DE_12h DE_12h-NFE2_MA0841.2 10 bp overlap
NFIA 1 dataset
Motif DE_12h DE_12h-NFIA_MA0670.2 6 bp overlap
NFIX 1 dataset
Motif DE_12h DE_12h-NFIX_MA0671.2 6 bp overlap
NKX2-2 4 datasets
Motif DE_12h DE_12h-NKX2-2_MA1645.2 8 bp overlap
Motif DE_60h DE_60h-NKX2-2_MA1645.2 8 bp overlap
Motif ES_0h ES_0h-NKX2-2_MA1645.2 8 bp overlap
Motif ES_0h ES_0h-NKX2-2_MA1645.2 8 bp overlap
NKX2-5 4 datasets
Motif DE_12h DE_12h-NKX2-5_MA0063.3 7 bp overlap
Motif DE_60h DE_60h-NKX2-5_MA0063.3 7 bp overlap
Motif ES_0h ES_0h-NKX2-5_MA0063.3 7 bp overlap
Motif ES_0h ES_0h-NKX2-5_MA0063.3 7 bp overlap
Neurod2 3 datasets
Motif DE_12h DE_12h-Neurod2_MA0668.3 8 bp overlap
Motif DE_36h DE_36h-Neurod2_MA0668.3 8 bp overlap
Motif ES_0h ES_0h-Neurod2_MA0668.3 8 bp overlap
Nfe2l2 1 dataset
Motif DE_12h DE_12h-Nfe2l2_MA0150.3 11 bp overlap
Nobox 4 datasets
Motif DE_12h DE_12h-Nobox_MA0125.2 6 bp overlap
Motif DE_36h DE_36h-Nobox_MA0125.2 6 bp overlap
Motif DE_48h DE_48h-Nobox_MA0125.2 6 bp overlap
Motif ES_0h ES_0h-Nobox_MA0125.2 6 bp overlap
POU5F1 1 dataset
ChIP BG03 GSE21614.POU5F1.BG03 157 bp overlap
Ptf1A 3 datasets
Motif DE_12h DE_12h-Ptf1A_MA1618.2 9 bp overlap
Motif DE_36h DE_36h-Ptf1A_MA1618.2 9 bp overlap
Motif ES_0h ES_0h-Ptf1A_MA1618.2 9 bp overlap
RAD21 21 datasets
ChIP A549 ENCFF047SFC 251 bp overlap
ChIP GM12878 ENCFF101UQZ 191 bp overlap
ChIP H1 ENCFF698EWO 127 bp overlap
ChIP H1 ENCFF967OJF 245 bp overlap
ChIP HCT-116 ENCSR000BSB.RAD21.HCT-116 186 bp overlap
ChIP HCT-116_RAD21-mAC GSE104888.RAD21.HCT-116_RAD21-mAC 297 bp overlap
ChIP HCT116 ENCFF568PEO 311 bp overlap
ChIP HEK293_siE1A GSE130135.RAD21.HEK293_siE1A 418 bp overlap
ChIP HeLa-Tet-On GSE112028.RAD21.HeLa-Tet-On 473 bp overlap
ChIP Ishikawa ENCFF570JVV 225 bp overlap
ChIP MCF-7 ENCFF694KOM 337 bp overlap
ChIP MCF-7 ENCFF724VCQ 98 bp overlap
ChIP MCF-7 ERP000209.RAD21.MCF-7 165 bp overlap
ChIP MCF-7 GSE72082.RAD21.MCF-7 175 bp overlap
ChIP MCF-7 ENCSR000BTQ.RAD21.MCF-7 117 bp overlap
ChIP MCF-7 GSE137216.RAD21.MCF-7 167 bp overlap
ChIP RH4 GSE83726.RAD21.RH4 232 bp overlap
ChIP T-47D_triptolide GSE111923.RAD21.T-47D_triptolide 281 bp overlap
ChIP THP-1_Pam3csk-020m-Flavo-000m GSE103477.RAD21.THP-1_Pam3csk-020m-Flavo-000m 199 bp overlap
ChIP THP-1_eGFP-IFNb GSE103477.RAD21.THP-1_eGFP-IFNb 160 bp overlap
ChIP hiPSC_parental-line GSE106870.RAD21.hiPSC_parental-line 190 bp overlap
RARA 1 dataset
Motif ES_0h ES_0h-RARA_MA0729.1 18 bp overlap
RAX 4 datasets
Motif DE_12h DE_12h-RAX_MA0718.2 6 bp overlap
Motif DE_36h DE_36h-RAX_MA0718.2 6 bp overlap
Motif DE_48h DE_48h-RAX_MA0718.2 6 bp overlap
Motif ES_0h ES_0h-RAX_MA0718.2 6 bp overlap
RBBP5 1 dataset
ChIP WA01 ENCSR000AQC.RBBP5.WA01 135 bp overlap
RBPJ 1 dataset
Motif DE_12h DE_12h-RBPJ_MA1116.2 6 bp overlap
Rarg 1 dataset
Motif ES_0h ES_0h-Rarg_MA0859.2 15 bp overlap
Rhox11 3 datasets
Motif DE_12h DE_12h-Rhox11_MA0629.2 9 bp overlap
Motif DE_60h DE_60h-Rhox11_MA0629.2 9 bp overlap
Motif ES_0h ES_0h-Rhox11_MA0629.2 9 bp overlap
SMARCA4 1 dataset
ChIP MCF-10A GSE74716.SMARCA4.MCF-10A 188 bp overlap
SMC3 4 datasets
ChIP HeLa GSE126990.SMC3.HeLa 143 bp overlap
ChIP HeLa-Kyoto GSE102884.SMC3.HeLa-Kyoto 143 bp overlap
ChIP HeLa-Kyoto GSE138405.SMC3.HeLa-Kyoto 143 bp overlap
ChIP HeLa-Kyoto_WAPL_PDS-depleted GSE102884.SMC3.HeLa-Kyoto_WAPL_PDS-depleted 286 bp overlap
SP8 5 datasets
Motif DE_12h DE_12h-SP8_MA0747.2 11 bp overlap
Motif DE_12h DE_12h-SP8_MA0747.2 11 bp overlap
Motif DE_60h DE_60h-SP8_MA0747.2 11 bp overlap
Motif ES_0h ES_0h-SP8_MA0747.2 11 bp overlap
Motif ES_0h ES_0h-SP8_MA0747.2 11 bp overlap
SRF 3 datasets
Motif DE_12h DE_12h-SRF_MA0083.3 16 bp overlap
Motif DE_48h DE_48h-SRF_MA0083.3 16 bp overlap
Motif ES_0h ES_0h-SRF_MA0083.3 16 bp overlap
STAG1 4 datasets
ChIP HeLa GSE126990.STAG1.HeLa 365 bp overlap
ChIP HeLa-Kyoto GSE138405.STAG1.HeLa-Kyoto 369 bp overlap
ChIP MCF-7 ERP000209.STAG1.MCF-7 220 bp overlap
ChIP MCF-7_E2 ERP000209.STAG1.MCF-7_E2 115 bp overlap
STAT1::STAT2 2 datasets
Motif DE_12h DE_12h-STAT1STAT2_MA0517.2 13 bp overlap
Motif ES_0h ES_0h-STAT1STAT2_MA0517.2 13 bp overlap
Smad4 2 datasets
Motif DE_12h DE_12h-Smad4_MA1153.2 7 bp overlap
Motif ES_0h ES_0h-Smad4_MA1153.2 7 bp overlap
Spz1 1 dataset
Motif DE_12h DE_12h-Spz1_MA0111.1 11 bp overlap
Stat2 2 datasets
Motif DE_12h DE_12h-Stat2_MA1623.2 10 bp overlap
Motif ES_0h ES_0h-Stat2_MA1623.2 10 bp overlap
TAL1::TCF3 3 datasets
Motif DE_12h DE_12h-TAL1TCF3_MA0091.2 10 bp overlap
Motif DE_36h DE_36h-TAL1TCF3_MA0091.2 10 bp overlap
Motif ES_0h ES_0h-TAL1TCF3_MA0091.2 10 bp overlap
TEAD4 1 dataset
ChIP hMSC-TERT4 GSE140782.TEAD4.hMSC-TERT4 220 bp overlap
TRIM25 1 dataset
ChIP MDA-MB-231 GSE79588.TRIM25.MDA-MB-231 178 bp overlap
YY1 2 datasets
ChIP H1 ENCFF524BTL 333 bp overlap
ChIP HEK293T_control GSE128106.YY1.HEK293T_control 262 bp overlap
ZBTB18 3 datasets
Motif DE_12h DE_12h-ZBTB18_MA0698.2 11 bp overlap
Motif DE_36h DE_36h-ZBTB18_MA0698.2 11 bp overlap
Motif ES_0h ES_0h-ZBTB18_MA0698.2 11 bp overlap
ZBTB7A 1 dataset
ChIP VCaP_FBS GSE123089.ZBTB7A.VCaP_FBS 215 bp overlap
ZIC4 3 datasets
Motif DE_12h DE_12h-ZIC4_MA0751.2 14 bp overlap
Motif DE_60h DE_60h-ZIC4_MA0751.2 14 bp overlap
Motif ES_0h ES_0h-ZIC4_MA0751.2 14 bp overlap
ZIM3 2 datasets
Motif DE_12h DE_12h-ZIM3_MA1709.2 11 bp overlap
Motif ES_0h ES_0h-ZIM3_MA1709.2 11 bp overlap
ZNF257 1 dataset
Motif DE_12h DE_12h-ZNF257_MA1710.2 10 bp overlap
ZNF282 1 dataset
Motif DE_12h DE_12h-ZNF282_MA1154.2 15 bp overlap
ZNF354A 3 datasets
Motif DE_12h DE_12h-ZNF354A_MA1978.2 20 bp overlap
Motif DE_36h DE_36h-ZNF354A_MA1978.2 20 bp overlap
Motif ES_0h ES_0h-ZNF354A_MA1978.2 20 bp overlap
ZNF418 3 datasets
Motif DE_12h DE_12h-ZNF418_MA1980.1 15 bp overlap
Motif DE_60h DE_60h-ZNF418_MA1980.1 15 bp overlap
Motif ES_0h ES_0h-ZNF418_MA1980.1 15 bp overlap
ZNF462 1 dataset
ChIP GM23338 ENCSR334UWP.ZNF462.GM23338 162 bp overlap
ZNF524 1 dataset
Motif DE_12h DE_12h-ZNF524_MA2096.1 9 bp overlap
ZNF654 2 datasets
ChIP HEK293 ENCFF636WIC 371 bp overlap
ChIP HEK293 ENCSR504VDV.ZNF654.HEK293 256 bp overlap
ZNF677 2 datasets
Motif DE_12h DE_12h-ZNF677_MA2101.1 12 bp overlap
Motif ES_0h ES_0h-ZNF677_MA2101.1 12 bp overlap
ZNF816 1 dataset
Motif DE_12h DE_12h-ZNF816_MA1719.2 15 bp overlap
ZSCAN16 3 datasets
Motif DE_12h DE_12h-ZSCAN16_MA2100.1 18 bp overlap
Motif DE_60h DE_60h-ZSCAN16_MA2100.1 18 bp overlap
Motif ES_0h ES_0h-ZSCAN16_MA2100.1 18 bp overlap
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