chr3 : 11,995,145 11,995,312
167 bp 90 TFs 1 linked gene
This 167 bp open chromatin element is linked to SYN2 and is bound by 90 transcription factors.
Linked Genes
1 gene
Gene Expression Dist. to TSS Distance Link type
SYN2 9.1 kb Proximal Proximity
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr3:11,990,145 – 12,000,312
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
90 transcription factors
Source
Cell type
AR 1 dataset
ChIP MCF-7 GSE48930.AR.MCF-7 86 bp overlap
ASCL1 6 datasets
ChIP G523NS_Dox GSE87618.ASCL1.G523NS_Dox 167 bp overlap
ChIP NCI-H128 GSE69394.ASCL1.NCI-H128 143 bp overlap
ChIP NCI-H2107 GSE69394.ASCL1.NCI-H2107 156 bp overlap
ChIP NCI-H82 GSE69394.ASCL1.NCI-H82 167 bp overlap
ChIP NCI-H889 GSE69394.ASCL1.NCI-H889 167 bp overlap
ChIP SCLC_ASCLP GSE61197.ASCL1.SCLC_ASCLP 167 bp overlap
ATRX 2 datasets
ChIP metastatic-neuroblastoma_SKNFI GSE100148.ATRX.metastatic-neuroblastoma_SKNFI 167 bp overlap
ChIP metastatic-neuroblastoma_SKNMM GSE100148.ATRX.metastatic-neuroblastoma_SKNMM 167 bp overlap
BRD4 10 datasets
ChIP Kelly_resistant GSE103030.BRD4.Kelly_resistant 136 bp overlap
ChIP Kelly_sensitive GSE103030.BRD4.Kelly_sensitive 167 bp overlap
ChIP NCI-H2171 GSE101821.BRD4.NCI-H2171 167 bp overlap
ChIP RH4_shCHD4 GSE140115.BRD4.RH4_shCHD4 167 bp overlap
ChIP RH4_shscr GSE140115.BRD4.RH4_shscr 167 bp overlap
ChIP SK-N-BE2-C_Naive-veh GSE107706.BRD4.SK-N-BE2-C_Naive-veh 53 bp overlap
ChIP SK-N-BE2-C_Resistant-veh GSE107706.BRD4.SK-N-BE2-C_Resistant-veh 167 bp overlap
ChIP cortical-interneuron_R133C_KCl-neg_JQ1-neg GSE117508.BRD4.cortical-interneuron_R133C_KCl-neg_JQ1-neg 151 bp overlap
ChIP cortical-interneuron_R133C_KCl-pos_JQ1-neg GSE117508.BRD4.cortical-interneuron_R133C_KCl-pos_JQ1-neg 158 bp overlap
ChIP hESC GSE33281.BRD4.hESC 61 bp overlap
CDK9 1 dataset
ChIP MM1-S_JQ1_500NM GSE42355.CDK9.MM1-S_JQ1_500NM 167 bp overlap
CHD2 1 dataset
ChIP SK-N-SH ENCSR274SLQ.CHD2.SK-N-SH 167 bp overlap
CHD4 2 datasets
ChIP RH5 GSE155861.CHD4.RH5 167 bp overlap
ChIP SCMC GSE155861.CHD4.SCMC 167 bp overlap
CREB1 1 dataset
ChIP WA01 ENCSR000BSN.CREB1.WA01 100 bp overlap
CTCF 31 datasets
ChIP HAP1 GSE152721.CTCF.HAP1 140 bp overlap
ChIP HAP1_clone21_TSS-CRISPR GSE152721.CTCF.HAP1_clone21_TSS-CRISPR 142 bp overlap
ChIP HeLa-Kyoto_PDS5-depleted GSE102884.CTCF.HeLa-Kyoto_PDS5-depleted 157 bp overlap
ChIP Kelly_resistant GSE103030.CTCF.Kelly_resistant 131 bp overlap
ChIP Kelly_sensitive GSE103030.CTCF.Kelly_sensitive 161 bp overlap
ChIP Loucy ENCSR464DKE.CTCF.Loucy 167 bp overlap
ChIP MCF-7 ENCFF414SZG 160 bp overlap
ChIP MCF-7 ENCFF424NQR 167 bp overlap
ChIP MCF-7 ENCFF844STM 167 bp overlap
ChIP MCF-7 ENCFF954TUV 161 bp overlap
ChIP MCF-7 ERP000380.CTCF.MCF-7 138 bp overlap
ChIP MCF-7 ENCSR000DMR.CTCF.MCF-7 158 bp overlap
ChIP MCF-7L_t24 GSE108787.CTCF.MCF-7L_t24 167 bp overlap
ChIP MCF-7_TAM ERP000380.CTCF.MCF-7_TAM 167 bp overlap
ChIP MCF-7_TAM ERP000783.CTCF.MCF-7_TAM 95 bp overlap
ChIP MM1-S GSE43743.CTCF.MM1-S 167 bp overlap
ChIP NB69 GSE101295.CTCF.NB69 167 bp overlap
ChIP RH4 GSE83726.CTCF.RH4 167 bp overlap
ChIP SH-SY5Y GSE101295.CTCF.SH-SY5Y 132 bp overlap
ChIP SK-N-SH ENCSR000DXQ.CTCF.SK-N-SH 167 bp overlap
ChIP VCaP ENCFF858YQT 167 bp overlap
ChIP WA01_NEUR GSE52457.CTCF.WA01_NEUR 167 bp overlap
ChIP cortical-interneuron_KCl-neg_JQ1-neg GSE117508.CTCF.cortical-interneuron_KCl-neg_JQ1-neg 143 bp overlap
ChIP cortical-interneuron_KCl-neg_JQ1-pos GSE117508.CTCF.cortical-interneuron_KCl-neg_JQ1-pos 154 bp overlap
ChIP cortical-interneuron_KCl-pos_JQ1-neg GSE117508.CTCF.cortical-interneuron_KCl-pos_JQ1-neg 126 bp overlap
ChIP cortical-interneuron_KCl-pos_JQ1-pos GSE117508.CTCF.cortical-interneuron_KCl-pos_JQ1-pos 134 bp overlap
ChIP neural ENCSR822CEA.CTCF.neural 167 bp overlap
ChIP neural cell ENCFF335ADI 167 bp overlap
ChIP parathyroid adenoma ENCFF173NJK 167 bp overlap
ChIP primary-epidermal-keratinocyte_diff_d0 GSE84657.CTCF.primary-epidermal-keratinocyte_diff_d0 90 bp overlap
ChIP primary-glioblastoma_G523 GSE121601.CTCF.primary-glioblastoma_G523 167 bp overlap
EP300 1 dataset
ChIP SK-N-SH ENCSR000EHV.EP300.SK-N-SH 167 bp overlap
ESR1 2 datasets
ChIP MCF-7_E2_45m GSE67295.ESR1.MCF-7_E2_45m 116 bp overlap
ChIP MCF-7_aldosterone GSE99626.ESR1.MCF-7_aldosterone 167 bp overlap
EZH2 2 datasets
ChIP RH30_DMSO GSE85169.EZH2.RH30_DMSO 167 bp overlap
ChIP THP-1 GSE135024.EZH2.THP-1 143 bp overlap
FOXA1 12 datasets
ChIP MCF-7 ERP000380.FOXA1.MCF-7 167 bp overlap
ChIP MCF-7 GSE59530.FOXA1.MCF-7 156 bp overlap
ChIP MCF-7 GSE60270.FOXA1.MCF-7 145 bp overlap
ChIP MCF-7 GSE72249.FOXA1.MCF-7 118 bp overlap
ChIP MCF-7_4OH-Tam GSE75201.FOXA1.MCF-7_4OH-Tam 119 bp overlap
ChIP MCF-7_E2 GSE60270.FOXA1.MCF-7_E2 157 bp overlap
ChIP MCF-7_E2 GSE23852.FOXA1.MCF-7_E2 110 bp overlap
ChIP MCF-7_ETOH GSE23852.FOXA1.MCF-7_ETOH 141 bp overlap
ChIP MCF-7_JC4691 GSE126004.FOXA1.MCF-7_JC4691 129 bp overlap
ChIP MCF-7_JC4697 GSE126004.FOXA1.MCF-7_JC4697 167 bp overlap
ChIP MCF-7_TNF GSE59530.FOXA1.MCF-7_TNF 154 bp overlap
ChIP MCF-7_estrogen_ab2 GSE112969.FOXA1.MCF-7_estrogen_ab2 167 bp overlap
FOXJ3 2 datasets
ChIP SK-N-SH ENCFF124KVL 167 bp overlap
ChIP SK-N-SH ENCFF124KVL 90 bp overlap
FOXO1-PAX3 2 datasets
ChIP RH4_DMSO-6H GSE116344.FOXO1-PAX3.RH4_DMSO-6H 167 bp overlap
ChIP RH4_Entinostat-6H GSE116344.FOXO1-PAX3.RH4_Entinostat-6H 159 bp overlap
GATA2 1 dataset
ChIP SH-SY5Y ENCSR000EYB.GATA2.SH-SY5Y 103 bp overlap
GLIS1 2 datasets
ChIP HEK293 ENCFF299RSE 167 bp overlap
ChIP HEK293 ENCSR482BBZ.GLIS1.HEK293 167 bp overlap
GTF3C2 1 dataset
ChIP H9 GSE94418.GTF3C2.H9 150 bp overlap
HAND2 3 datasets
ChIP CLB-Ga GSE90683.HAND2.CLB-Ga 167 bp overlap
ChIP Kelly GSE94822.HAND2.Kelly 167 bp overlap
ChIP SK-N-BE2-C GSE94822.HAND2.SK-N-BE2-C 167 bp overlap
HDAC2 5 datasets
ChIP MCF-7 ENCSR000BTP.HDAC2.MCF-7 140 bp overlap
ChIP RH4_DMSO-6H GSE116344.HDAC2.RH4_DMSO-6H 167 bp overlap
ChIP RH4_Entinostat-6H GSE116344.HDAC2.RH4_Entinostat-6H 167 bp overlap
ChIP RH4_Entinostat-6H_bioMerck GSE116344.HDAC2.RH4_Entinostat-6H_bioMerck 51 bp overlap
ChIP WA01 ENCSR000AVB.HDAC2.WA01 122 bp overlap
HDAC3 1 dataset
ChIP RH4_DMSO-6H GSE116344.HDAC3.RH4_DMSO-6H 167 bp overlap
HNF4A 1 dataset
ChIP Caco-2_DIFF GSE23436.HNF4A.Caco-2_DIFF 75 bp overlap
ISL1 1 dataset
ChIP SK-N-BE2-C GSE94822.ISL1.SK-N-BE2-C 144 bp overlap
KDM1A 2 datasets
ChIP SH-SY5Y_B3 GSE58258.KDM1A.SH-SY5Y_B3 150 bp overlap
ChIP pancreatic-progenitor_PP2-KDM1A-inh GSE104840.KDM1A.pancreatic-progenitor_PP2-KDM1A-inh 135 bp overlap
KDM5B 1 dataset
ChIP SUM185_SHCTCF GSE46055.KDM5B.SUM185_SHCTCF 123 bp overlap
KLF9 2 datasets
ChIP GBM1A GSE62211.KLF9.GBM1A 86 bp overlap
ChIP HEK293 ENCSR076EZB.KLF9.HEK293 167 bp overlap
LMO1 1 dataset
ChIP Jurkat GSE94391.LMO1.Jurkat 167 bp overlap
MAX 2 datasets
ChIP NCI-H2171 GSE41105.MAX.NCI-H2171 167 bp overlap
ChIP NCI-H2171 GSE36354.MAX.NCI-H2171 167 bp overlap
MAZ 1 dataset
ChIP HEK293 GSE76494.MAZ.HEK293 99 bp overlap
MED1 4 datasets
ChIP MCF-7_SHRARS GSE60270.MED1.MCF-7_SHRARS 167 bp overlap
ChIP MCF-7_SHRARS_E2 GSE60270.MED1.MCF-7_SHRARS_E2 124 bp overlap
ChIP NCI-H2171 GSE36354.MED1.NCI-H2171 167 bp overlap
ChIP RH4 GSE83726.MED1.RH4 137 bp overlap
MTA2 1 dataset
ChIP RH4 GSE155861.MTA2.RH4 167 bp overlap
MXI1 2 datasets
ChIP SK-N-SH ENCFF746HVJ 167 bp overlap
ChIP SK-N-SH ENCSR000EIA.MXI1.SK-N-SH 149 bp overlap
MYB 2 datasets
ChIP Jurkat GSE59657.MYB.Jurkat 167 bp overlap
ChIP PF-382 GSE94000.MYB.PF-382 167 bp overlap
MYC 2 datasets
ChIP NCI-H128 GSE41105.MYC.NCI-H128 167 bp overlap
ChIP NCI-H2171 GSE36354.MYC.NCI-H2171 167 bp overlap
MYCN 8 datasets
ChIP BE2C GSE80151.MYCN.BE2C 167 bp overlap
ChIP CHP-134 GSE129588.MYCN.CHP-134 78 bp overlap
ChIP COG-N-415 GSE138295.MYCN.COG-N-415 167 bp overlap
ChIP Kelly GSE94822.MYCN.Kelly 167 bp overlap
ChIP Kelly_sensitive GSE115249.MYCN.Kelly_sensitive 161 bp overlap
ChIP LA-N-5 GSE138295.MYCN.LA-N-5 167 bp overlap
ChIP SK-N-BE2-C GSE80151.MYCN.SK-N-BE2-C 167 bp overlap
ChIP SK-N-BE2-C GSE94822.MYCN.SK-N-BE2-C 110 bp overlap
MYF5 1 dataset
ChIP Rh18 GSE84628.MYF5.Rh18 167 bp overlap
MYOD1 8 datasets
ChIP IMR-90_GM GSE128527.MYOD1.IMR-90_GM 167 bp overlap
ChIP RD GSE137168.MYOD1.RD 167 bp overlap
ChIP RD_shSNAI2 GSE137168.MYOD1.RD_shSNAI2 167 bp overlap
ChIP RH30_DMSO GSE85169.MYOD1.RH30_DMSO 167 bp overlap
ChIP RH4 GSE83726.MYOD1.RH4 167 bp overlap
ChIP SMS-CTR GSE137168.MYOD1.SMS-CTR 167 bp overlap
ChIP myoblast GSE50413.MYOD1.myoblast 167 bp overlap
ChIP rhabdomyosarcoma GSE50413.MYOD1.rhabdomyosarcoma 167 bp overlap
MYOG 4 datasets
ChIP RH30_DMSO GSE85169.MYOG.RH30_DMSO 167 bp overlap
ChIP RH4 GSE83726.MYOG.RH4 167 bp overlap
ChIP RH4_DMSO-6H GSE116344.MYOG.RH4_DMSO-6H 167 bp overlap
ChIP RH4_Entinostat-6H GSE116344.MYOG.RH4_Entinostat-6H 167 bp overlap
NCOR1 1 dataset
ChIP LS180_125 GSE39277.NCOR1.LS180_125 101 bp overlap
NEUROD1 2 datasets
ChIP D341-Med_shGFP GSE92582.NEUROD1.D341-Med_shGFP 167 bp overlap
ChIP NCI-H524 GSE69394.NEUROD1.NCI-H524 167 bp overlap
NEUROG2 5 datasets
ChIP MRC-5_NFD_02DPT GSE75910.NEUROG2.MRC-5_NFD_02DPT 147 bp overlap
ChIP MRC-5_NFD_1DPT GSE75910.NEUROG2.MRC-5_NFD_1DPT 167 bp overlap
ChIP MRC-5_N_02DPT GSE75910.NEUROG2.MRC-5_N_02DPT 138 bp overlap
ChIP MRC-5_N_05DPT GSE75910.NEUROG2.MRC-5_N_05DPT 167 bp overlap
ChIP MRC-5_N_1DPT GSE75910.NEUROG2.MRC-5_N_1DPT 167 bp overlap
NKX2-1 1 dataset
ChIP H9_derived-cIN GSE99937.NKX2-1.H9_derived-cIN 88 bp overlap
NOTCH1 1 dataset
ChIP HPBALL GSE39263.NOTCH1.HPBALL 131 bp overlap
NR3C1 1 dataset
ChIP breast_tumor_Male_15 GSE104399.NR3C1.breast_tumor_Male_15 143 bp overlap
OSR2 1 dataset
ChIP HEK293 GSE76494.OSR2.HEK293 124 bp overlap
PAX3-FOXO1 2 datasets
ChIP RH3 GSE83726.PAX3-FOXO1.RH3 140 bp overlap
ChIP RH4 GSE140115.PAX3-FOXO1.RH4 167 bp overlap
PITX3 1 dataset
ChIP SH-SY5Y GSE93275.PITX3.SH-SY5Y 167 bp overlap
PRDM10 1 dataset
ChIP HEK293 ENCFF145WQQ 167 bp overlap
RAD21 5 datasets
ChIP IMR-5 GSE78957.RAD21.IMR-5 167 bp overlap
ChIP Ishikawa ENCSR000BTU.RAD21.Ishikawa 106 bp overlap
ChIP RH4 GSE83726.RAD21.RH4 167 bp overlap
ChIP SK-N-SH ENCFF747MAS 131 bp overlap
ChIP SK-N-SH ENCSR000EHX.RAD21.SK-N-SH 149 bp overlap
RBBP4 2 datasets
ChIP RH5 GSE155861.RBBP4.RH5 167 bp overlap
ChIP SCMC GSE155861.RBBP4.SCMC 167 bp overlap
RCOR1 2 datasets
ChIP SK-N-SH ENCFF518EXB 167 bp overlap
ChIP SK-N-SH ENCSR009TKN.RCOR1.SK-N-SH 161 bp overlap
RFX5 4 datasets
ChIP MCF-7 ENCFF983ILY 167 bp overlap
ChIP MCF-7 ENCSR924TVL.RFX5.MCF-7 167 bp overlap
ChIP SK-N-SH ENCFF755HLO 167 bp overlap
ChIP SK-N-SH ENCSR000EHY.RFX5.SK-N-SH 167 bp overlap
RUNX1 2 datasets
ChIP Jurkat GSE68976.RUNX1.Jurkat 123 bp overlap
ChIP Jurkat GSE76181.RUNX1.Jurkat 92 bp overlap
SAP30 1 dataset
ChIP WA01 ENCSR000ATR.SAP30.WA01 165 bp overlap
SIN3A 8 datasets
ChIP H1 ENCFF042ZSL 167 bp overlap
ChIP H1 ENCFF896IJG 167 bp overlap
ChIP MCF-7 ENCSR000BUM.SIN3A.MCF-7 131 bp overlap
ChIP WA01 ENCSR000EBO.SIN3A.WA01 167 bp overlap
ChIP WA01 ENCSR000BIS.SIN3A.WA01 144 bp overlap
ChIP hiPSC_IA12 GSE106870.SIN3A.hiPSC_IA12 167 bp overlap
ChIP hiPSC_IB12 GSE106870.SIN3A.hiPSC_IB12 144 bp overlap
ChIP hiPSC_IID12 GSE106870.SIN3A.hiPSC_IID12 79 bp overlap
SMARCA4 5 datasets
ChIP NGP GSE134626.SMARCA4.NGP 116 bp overlap
ChIP NGP_ARID1A-het GSE134626.SMARCA4.NGP_ARID1A-het 167 bp overlap
ChIP NGP_ARID1A-mut1 GSE134626.SMARCA4.NGP_ARID1A-mut1 167 bp overlap
ChIP NGP_ARID1A-mut2 GSE134626.SMARCA4.NGP_ARID1A-mut2 167 bp overlap
ChIP TTC-1240_delC GSE124903.SMARCA4.TTC-1240_delC 142 bp overlap
SMARCC1 1 dataset
ChIP TTC-1240_delC GSE124903.SMARCC1.TTC-1240_delC 167 bp overlap
SMC3 1 dataset
ChIP neural ENCSR404BPV.SMC3.neural 143 bp overlap
SNAI2 3 datasets
ChIP RD_shSNAI2 GSE137168.SNAI2.RD_shSNAI2 167 bp overlap
ChIP SMS-CTR_D48 GSE137168.SNAI2.SMS-CTR_D48 167 bp overlap
ChIP SMS-CTR_shSNAI2 GSE137168.SNAI2.SMS-CTR_shSNAI2 162 bp overlap
SOX8 2 datasets
ChIP RH4 GSE116344.SOX8.RH4 159 bp overlap
ChIP RH4_DMSO-6H GSE116344.SOX8.RH4_DMSO-6H 111 bp overlap
SPI1 1 dataset
ChIP macrophage_IFNG GSE47188.SPI1.macrophage_IFNG 167 bp overlap
TBL1X 1 dataset
ChIP HEK293T GSE35197.TBL1X.HEK293T 167 bp overlap
TBX2 2 datasets
ChIP Kelly GSE94822.TBX2.Kelly 167 bp overlap
ChIP SK-N-BE2-C GSE94822.TBX2.SK-N-BE2-C 134 bp overlap
TBX5 1 dataset
ChIP G296S_4 GSE85628.TBX5.G296S_4 139 bp overlap
TCF12 1 dataset
ChIP WA01 ENCSR000BIT.TCF12.WA01 154 bp overlap
TCF3 1 dataset
ChIP NPC GSE154479.TCF3.NPC 166 bp overlap
TFAP2A 1 dataset
ChIP MCF-7_E2 GSE60270.TFAP2A.MCF-7_E2 150 bp overlap
TFAP2B 1 dataset
ChIP SK-N-SH ENCFF869XXQ 167 bp overlap
TFAP2C 2 datasets
ChIP MCF-7_E2 GSE23852.TFAP2C.MCF-7_E2 133 bp overlap
ChIP MCF-7_ETOH GSE23852.TFAP2C.MCF-7_ETOH 154 bp overlap
TFAP4 1 dataset
ChIP Hep-G2 ENCSR103SZL.TFAP4.Hep-G2 135 bp overlap
TP53 1 dataset
ChIP Calu-1_MUT8-COMB GSE128673.TP53.Calu-1_MUT8-COMB 140 bp overlap
TWIST1 2 datasets
ChIP BE2C GSE80151.TWIST1.BE2C 167 bp overlap
ChIP SK-N-BE2-C GSE80151.TWIST1.SK-N-BE2-C 167 bp overlap
USF1 1 dataset
ChIP WA01 ENCSR000BIU.USF1.WA01 101 bp overlap
YY1 1 dataset
ChIP RH4_DMSO-6H GSE116344.YY1.RH4_DMSO-6H 167 bp overlap
ZBTB17 1 dataset
ChIP HEK293 ENCFF865LIO 167 bp overlap
ZBTB42 1 dataset
ChIP HEK293 GSE76494.ZBTB42.HEK293 167 bp overlap
ZBTB44 2 datasets
ChIP HEK293 ENCFF560VPN 162 bp overlap
ChIP HEK293 ENCSR076STQ.ZBTB44.HEK293 167 bp overlap
ZBTB7A 1 dataset
ChIP VCaP_FBS GSE123089.ZBTB7A.VCaP_FBS 167 bp overlap
ZEB1 1 dataset
ChIP neuron_bipolar_doxy_4d ENCSR418KUS.ZEB1.neuron_bipolar_doxy_4d 167 bp overlap
ZFP64 1 dataset
ChIP HEK293 GSE76494.ZFP64.HEK293 167 bp overlap
ZNF12 1 dataset
ChIP Hep-G2 ENCSR711KBM.ZNF12.Hep-G2 146 bp overlap
ZNF30 1 dataset
ChIP HEK293 GSE76494.ZNF30.HEK293 135 bp overlap
ZNF335 2 datasets
ChIP HEK293 ENCFF784SLD 167 bp overlap
ChIP HEK293 ENCSR328SUD.ZNF335.HEK293 167 bp overlap
ZNF341 3 datasets
ChIP HEK293 ENCFF944VMC 167 bp overlap
ChIP HEK293 ENCSR185FOY.ZNF341.HEK293 167 bp overlap
ChIP HEK293 GSE76494.ZNF341.HEK293 167 bp overlap
ZNF549 1 dataset
ChIP HEK293 GSE76494.ZNF549.HEK293 66 bp overlap
ZNF554 1 dataset
ChIP HEK293 GSE76494.ZNF554.HEK293 167 bp overlap
ZNF574 3 datasets
ChIP HEK293 GSE76494.ZNF574.HEK293 155 bp overlap
ChIP HepG2 ENCFF206MMY 167 bp overlap
ChIP HepG2 ENCFF206MMY 113 bp overlap