chr2 : 137,607,911 – 137,609,646
1,735 bp 126 TFs 0 linked genes
This 1.7 kb open chromatin element has no linked target genes and is bound by 126 transcription factors.
Linked Genes
No linked genes
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr2:137,602,911 – 137,614,646
This element Other elements ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
126 transcription factors
Source
Cell type
ARRB1 1 dataset
ChIP prostate GSE55615.ARRB1.prostate 176 bp overlap
Arid3a 2 datasets
Motif DE_48h DE_48h-Arid3a_MA0151.1 6 bp overlap
Motif DE_60h DE_60h-Arid3a_MA0151.1 6 bp overlap
BACH2 1 dataset
Motif DE_60h DE_60h-BACH2_MA1470.2 19 bp overlap
BARX2 2 datasets
Motif DE_60h DE_60h-BARX2_MA1471.2 9 bp overlap
Motif DE_60h DE_60h-BARX2_MA1471.2 9 bp overlap
CTCF 1 dataset
ChIP anterior-temporal-cortex_fetal GSE116825.CTCF.anterior-temporal-cortex_fetal 181 bp overlap
DMRTA2 1 dataset
Motif DE_60h DE_60h-DMRTA2_MA1478.2 6 bp overlap
DMRTC2 1 dataset
Motif DE_60h DE_60h-DMRTC2_MA1479.2 11 bp overlap
DUX4 2 datasets
Motif DE_60h DE_60h-DUX4_MA0468.1 11 bp overlap
ChIP WA01 GSE94322.DUX4.WA01 201 bp overlap
DUXA 1 dataset
Motif DE_60h DE_60h-DUXA_MA0884.2 13 bp overlap
EBF1 3 datasets
Motif DE_48h DE_48h-EBF1_MA0154.5 11 bp overlap
Motif DE_60h DE_60h-EBF1_MA0154.5 11 bp overlap
Motif DE_72h DE_72h-EBF1_MA0154.5 11 bp overlap
EBF3 3 datasets
Motif DE_48h DE_48h-EBF3_MA1637.2 9 bp overlap
Motif DE_60h DE_60h-EBF3_MA1637.2 9 bp overlap
Motif DE_72h DE_72h-EBF3_MA1637.2 9 bp overlap
ELK1::HOXA1 1 dataset
Motif DE_60h DE_60h-ELK1HOXA1_MA1931.1 14 bp overlap
EOMES 2 datasets
ChIP hESC GSE26097.EOMES.hESC 423 bp overlap
ChIP hESC GSE26097.EOMES.hESC 360 bp overlap
ERF::FIGLA 1 dataset
Motif DE_60h DE_60h-ERFFIGLA_MA1934.2 13 bp overlap
ESR2 3 datasets
Motif DE_48h DE_48h-ESR2_MA0258.2 15 bp overlap
Motif DE_60h DE_60h-ESR2_MA0258.2 15 bp overlap
Motif DE_72h DE_72h-ESR2_MA0258.2 15 bp overlap
EZH2 1 dataset
ChIP SU-DHL-5 GSE45982.EZH2.SU-DHL-5 89 bp overlap
Ebf4 3 datasets
Motif DE_48h DE_48h-Ebf4_MA2122.1 11 bp overlap
Motif DE_60h DE_60h-Ebf4_MA2122.1 11 bp overlap
Motif DE_72h DE_72h-Ebf4_MA2122.1 11 bp overlap
FOXA2 2 datasets
ChIP DE DE-FOXA2-1 1719 bp overlap
ChIP DE DE-FOXA2-2 1698 bp overlap
FOXE1 2 datasets
Motif DE_60h DE_60h-FOXE1_MA1487.3 12 bp overlap
Motif DE_72h DE_72h-FOXE1_MA1487.3 12 bp overlap
GATA2 2 datasets
ChIP LNCaP GSE38391.GATA2.LNCaP 139 bp overlap
ChIP LNCaP GSE52725.GATA2.LNCaP 139 bp overlap
GATA4 3 datasets
ChIP DE DE-GATA4-1 1735 bp overlap
ChIP DE DE-GATA4-2 1735 bp overlap
Motif DE_60h DE_60h-GATA4_MA0482.3 8 bp overlap
GATA5 1 dataset
Motif DE_60h DE_60h-GATA5_MA0766.3 8 bp overlap
GATA6 10 datasets
ChIP DE DE-GATA6-1 1735 bp overlap
ChIP DE DE-GATA6-2 1735 bp overlap
Motif DE_60h DE_60h-GATA6_MA1104.3 8 bp overlap
Motif DE_60h DE_60h-GATA6_MA1104.3 8 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 404 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 1579 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 507 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 358 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 585 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 779 bp overlap
GCM1 1 dataset
Motif DE_60h DE_60h-GCM1_MA0646.2 10 bp overlap
GLI3 1 dataset
Motif DE_60h DE_60h-GLI3_MA1491.3 15 bp overlap
GLIS3 1 dataset
Motif DE_60h DE_60h-GLIS3_MA0737.1 14 bp overlap
GMEB2 1 dataset
Motif DE_60h DE_60h-GMEB2_MA0862.1 8 bp overlap
GRHL1 2 datasets
Motif DE_60h DE_60h-GRHL1_MA0647.2 10 bp overlap
Motif DE_72h DE_72h-GRHL1_MA0647.2 10 bp overlap
GRHL2 2 datasets
Motif DE_60h DE_60h-GRHL2_MA1105.3 8 bp overlap
Motif DE_72h DE_72h-GRHL2_MA1105.3 8 bp overlap
Gata3 2 datasets
Motif DE_60h DE_60h-Gata3_MA0037.5 8 bp overlap
Motif DE_60h DE_60h-Gata3_MA0037.5 8 bp overlap
Gli1 1 dataset
Motif DE_60h DE_60h-Gli1_MA1990.2 10 bp overlap
Gli2 1 dataset
Motif DE_60h DE_60h-Gli2_MA0734.4 9 bp overlap
Gmeb1 1 dataset
Motif DE_60h DE_60h-Gmeb1_MA0615.2 6 bp overlap
HDAC2 1 dataset
ChIP Hep-G2 ENCSR337NWW.HDAC2.Hep-G2 210 bp overlap
HNF1A 3 datasets
Motif DE_48h DE_48h-HNF1A_MA0046.3 13 bp overlap
Motif DE_60h DE_60h-HNF1A_MA0046.3 13 bp overlap
Motif DE_72h DE_72h-HNF1A_MA0046.3 13 bp overlap
HNF1B 3 datasets
Motif DE_48h DE_48h-HNF1B_MA0153.2 13 bp overlap
Motif DE_60h DE_60h-HNF1B_MA0153.2 13 bp overlap
Motif DE_72h DE_72h-HNF1B_MA0153.2 13 bp overlap
HOXA4 1 dataset
Motif DE_60h DE_60h-HOXA4_MA1496.2 7 bp overlap
HOXB4 6 datasets
Motif DE_48h DE_48h-HOXB4_MA1499.2 6 bp overlap
Motif DE_48h DE_48h-HOXB4_MA1499.2 6 bp overlap
Motif DE_60h DE_60h-HOXB4_MA1499.2 6 bp overlap
Motif DE_60h DE_60h-HOXB4_MA1499.2 6 bp overlap
Motif DE_60h DE_60h-HOXB4_MA1499.2 6 bp overlap
Motif DE_60h DE_60h-HOXB4_MA1499.2 6 bp overlap
HOXB9 2 datasets
Motif DE_48h DE_48h-HOXB9_MA1503.2 9 bp overlap
Motif DE_60h DE_60h-HOXB9_MA1503.2 9 bp overlap
HOXC10 3 datasets
Motif DE_48h DE_48h-HOXC10_MA0905.2 9 bp overlap
Motif DE_60h DE_60h-HOXC10_MA0905.2 9 bp overlap
Motif DE_60h DE_60h-HOXC10_MA0905.2 9 bp overlap
HOXC13 3 datasets
Motif DE_48h DE_48h-HOXC13_MA0907.2 9 bp overlap
Motif DE_60h DE_60h-HOXC13_MA0907.2 9 bp overlap
Motif DE_60h DE_60h-HOXC13_MA0907.2 9 bp overlap
HOXC4 6 datasets
Motif DE_48h DE_48h-HOXC4_MA1504.2 6 bp overlap
Motif DE_48h DE_48h-HOXC4_MA1504.2 6 bp overlap
Motif DE_60h DE_60h-HOXC4_MA1504.2 6 bp overlap
Motif DE_60h DE_60h-HOXC4_MA1504.2 6 bp overlap
Motif DE_60h DE_60h-HOXC4_MA1504.2 6 bp overlap
Motif DE_60h DE_60h-HOXC4_MA1504.2 6 bp overlap
HOXD3 2 datasets
Motif DE_48h DE_48h-HOXD3_MA0912.2 8 bp overlap
Motif DE_60h DE_60h-HOXD3_MA0912.2 8 bp overlap
HOXD4 6 datasets
Motif DE_48h DE_48h-HOXD4_MA1507.2 6 bp overlap
Motif DE_48h DE_48h-HOXD4_MA1507.2 6 bp overlap
Motif DE_60h DE_60h-HOXD4_MA1507.2 6 bp overlap
Motif DE_60h DE_60h-HOXD4_MA1507.2 6 bp overlap
Motif DE_60h DE_60h-HOXD4_MA1507.2 6 bp overlap
Motif DE_60h DE_60h-HOXD4_MA1507.2 6 bp overlap
Hmga1 3 datasets
Motif DE_48h DE_48h-Hmga1_MA2124.1 8 bp overlap
Motif DE_60h DE_60h-Hmga1_MA2124.1 8 bp overlap
Motif DE_72h DE_72h-Hmga1_MA2124.1 8 bp overlap
Hmx2 1 dataset
Motif DE_60h DE_60h-Hmx2_MA0897.2 15 bp overlap
Hoxa11 2 datasets
Motif DE_48h DE_48h-Hoxa11_MA0911.2 9 bp overlap
Motif DE_60h DE_60h-Hoxa11_MA0911.2 9 bp overlap
Hoxa13 1 dataset
Motif DE_60h DE_60h-Hoxa13_MA0650.4 8 bp overlap
IKZF2 1 dataset
Motif DE_60h DE_60h-IKZF2_MA2326.1 6 bp overlap
ISL2 3 datasets
Motif DE_48h DE_48h-ISL2_MA0914.2 6 bp overlap
Motif DE_60h DE_60h-ISL2_MA0914.2 6 bp overlap
Motif DE_72h DE_72h-ISL2_MA0914.2 6 bp overlap
JUN 1 dataset
ChIP DE_D2 S02-DE-d2-JUN-exp1 554 bp overlap
Lhx3 1 dataset
Motif DE_60h DE_60h-Lhx3_MA0135.2 12 bp overlap
MAFF 2 datasets
ChIP HeLa-S3 ENCSR140DSL.MAFF.HeLa-S3 86 bp overlap
ChIP K-562 ENCSR000EGI.MAFF.K-562 72 bp overlap
MAFK 5 datasets
ChIP A549 ENCFF371EPR 195 bp overlap
ChIP Hep-G2 ENCSR000EDZ.MAFK.Hep-G2 109 bp overlap
ChIP HepG2 ENCFF743ZOF 51 bp overlap
ChIP HepG2 ENCFF767LDG 145 bp overlap
ChIP IMR-90 ENCFF336DHZ 155 bp overlap
MED1 1 dataset
ChIP VCaP_DHTTHZ1 GSE125245.MED1.VCaP_DHTTHZ1 159 bp overlap
MEIS1 1 dataset
Motif DE_60h DE_60h-MEIS1_MA1639.2 9 bp overlap
MEIS2 1 dataset
Motif DE_60h DE_60h-MEIS2_MA1640.2 9 bp overlap
MGA::EVX1 3 datasets
Motif DE_48h DE_48h-MGAEVX1_MA1960.2 11 bp overlap
Motif DE_60h DE_60h-MGAEVX1_MA1960.2 11 bp overlap
Motif DE_72h DE_72h-MGAEVX1_MA1960.2 11 bp overlap
NKX2-3 1 dataset
Motif DE_60h DE_60h-NKX2-3_MA0672.2 8 bp overlap
NKX2-4 1 dataset
Motif DE_60h DE_60h-NKX2-4_MA2003.2 8 bp overlap
NKX2-8 1 dataset
Motif DE_60h DE_60h-NKX2-8_MA0673.2 8 bp overlap
NKX6-1 4 datasets
Motif DE_48h DE_48h-NKX6-1_MA0674.2 7 bp overlap
Motif DE_60h DE_60h-NKX6-1_MA0674.2 7 bp overlap
Motif DE_60h DE_60h-NKX6-1_MA0674.2 7 bp overlap
Motif DE_60h DE_60h-NKX6-1_MA0674.2 7 bp overlap
NKX6-3 3 datasets
Motif DE_48h DE_48h-NKX6-3_MA1530.2 8 bp overlap
Motif DE_60h DE_60h-NKX6-3_MA1530.2 8 bp overlap
Motif DE_60h DE_60h-NKX6-3_MA1530.2 8 bp overlap
NR1D2 3 datasets
Motif DE_48h DE_48h-NR1D2_MA1532.2 15 bp overlap
Motif DE_60h DE_60h-NR1D2_MA1532.2 15 bp overlap
Motif DE_72h DE_72h-NR1D2_MA1532.2 15 bp overlap
NR4A1 3 datasets
Motif DE_48h DE_48h-NR4A1_MA1112.3 8 bp overlap
Motif DE_60h DE_60h-NR4A1_MA1112.3 8 bp overlap
Motif DE_72h DE_72h-NR4A1_MA1112.3 8 bp overlap
NR4A2 3 datasets
Motif DE_48h DE_48h-NR4A2_MA0160.3 8 bp overlap
Motif DE_60h DE_60h-NR4A2_MA0160.3 8 bp overlap
Motif DE_72h DE_72h-NR4A2_MA0160.3 8 bp overlap
Nkx2-1 1 dataset
Motif DE_60h DE_60h-Nkx2-1_MA1994.2 7 bp overlap
Nkx3-2 3 datasets
Motif DE_48h DE_48h-Nkx3-2_MA0122.4 10 bp overlap
Motif DE_60h DE_60h-Nkx3-2_MA0122.4 10 bp overlap
Motif DE_72h DE_72h-Nkx3-2_MA0122.4 10 bp overlap
Nr2e3 3 datasets
Motif DE_48h DE_48h-Nr2e3_MA0164.2 6 bp overlap
Motif DE_60h DE_60h-Nr2e3_MA0164.2 6 bp overlap
Motif DE_72h DE_72h-Nr2e3_MA0164.2 6 bp overlap
OSR2 1 dataset
Motif DE_60h DE_60h-OSR2_MA1646.2 8 bp overlap
PAX4 1 dataset
Motif DE_60h DE_60h-PAX4_MA0068.2 8 bp overlap
PGR 3 datasets
Motif DE_48h DE_48h-PGR_MA2327.1 9 bp overlap
Motif DE_60h DE_60h-PGR_MA2327.1 9 bp overlap
Motif DE_72h DE_72h-PGR_MA2327.1 9 bp overlap
PHOX2A 1 dataset
Motif DE_60h DE_60h-PHOX2A_MA0713.1 11 bp overlap
PHOX2B 1 dataset
Motif DE_60h DE_60h-PHOX2B_MA0681.3 12 bp overlap
PITX3 1 dataset
ChIP SH-SY5Y GSE93275.PITX3.SH-SY5Y 360 bp overlap
POU2F1 1 dataset
Motif DE_60h DE_60h-POU2F1_MA0785.2 9 bp overlap
POU2F3 1 dataset
Motif DE_60h DE_60h-POU2F3_MA0627.3 9 bp overlap
POU3F4 1 dataset
Motif DE_60h DE_60h-POU3F4_MA0789.1 9 bp overlap
POU4F1 1 dataset
Motif DE_60h DE_60h-POU4F1_MA0790.2 12 bp overlap
POU4F2 1 dataset
Motif DE_60h DE_60h-POU4F2_MA0683.2 15 bp overlap
POU4F3 1 dataset
Motif DE_60h DE_60h-POU4F3_MA0791.2 12 bp overlap
POU5F1 1 dataset
Motif DE_60h DE_60h-POU5F1_MA1115.2 7 bp overlap
POU5F1B 1 dataset
Motif DE_60h DE_60h-POU5F1B_MA0792.1 9 bp overlap
POU6F1 4 datasets
Motif DE_48h DE_48h-POU6F1_MA1549.2 7 bp overlap
Motif DE_60h DE_60h-POU6F1_MA1549.2 7 bp overlap
Motif DE_60h DE_60h-POU6F1_MA1549.2 7 bp overlap
Motif DE_60h DE_60h-POU6F1_MA1549.2 7 bp overlap
POU6F2 3 datasets
Motif DE_48h DE_48h-POU6F2_MA0793.2 9 bp overlap
Motif DE_60h DE_60h-POU6F2_MA0793.2 9 bp overlap
Motif DE_60h DE_60h-POU6F2_MA0793.2 9 bp overlap
PPARA::RXRA 1 dataset
Motif DE_60h DE_60h-PPARARXRA_MA1148.2 17 bp overlap
PROP1 1 dataset
Motif DE_60h DE_60h-PROP1_MA0715.1 11 bp overlap
REST 2 datasets
ChIP PFSK-1 ENCFF845VHA 294 bp overlap
ChIP PFSK1 ENCSR000BOX.REST.PFSK1 202 bp overlap
SMAD2 3 datasets
Motif DE_48h DE_48h-SMAD2_MA1964.2 6 bp overlap
Motif DE_60h DE_60h-SMAD2_MA1964.2 6 bp overlap
Motif DE_72h DE_72h-SMAD2_MA1964.2 6 bp overlap
SMAD2_3 3 datasets
ChIP DE_D2 S01-DE-d2-SMAD2_3-exp1 1360 bp overlap
ChIP DE_D2 S53-DE-d2-SMAD2_3-exp2 616 bp overlap
ChIP DE_D2 S53-DE-d2-SMAD2_3-exp2 420 bp overlap
SMARCA4 1 dataset
ChIP NGP GSE134626.SMARCA4.NGP 159 bp overlap
SOX15 3 datasets
Motif DE_48h DE_48h-SOX15_MA1152.2 7 bp overlap
Motif DE_60h DE_60h-SOX15_MA1152.2 7 bp overlap
Motif DE_72h DE_72h-SOX15_MA1152.2 7 bp overlap
SOX17 2 datasets
ChIP DE_D2 DED2-SOX17_Batch_II 437 bp overlap
ChIP DE_D2 DED2-SOX17_Batch_II 341 bp overlap
SOX17_M 1 dataset
ChIP DE_D2 DED2-SOX17-M_Batch_II 1671 bp overlap
SP5 4 datasets
Motif DE_48h DE_48h-SP5_MA1965.2 6 bp overlap
Motif DE_60h DE_60h-SP5_MA1965.2 6 bp overlap
Motif DE_60h DE_60h-SP5_MA1965.2 6 bp overlap
Motif DE_72h DE_72h-SP5_MA1965.2 6 bp overlap
STAT1::STAT2 2 datasets
Motif DE_60h DE_60h-STAT1STAT2_MA0517.2 13 bp overlap
Motif DE_72h DE_72h-STAT1STAT2_MA0517.2 13 bp overlap
STAT3 1 dataset
ChIP HCC1187 GSE152203.STAT3.HCC1187 179 bp overlap
Stat5a::Stat5b 1 dataset
Motif DE_60h DE_60h-Stat5aStat5b_MA0519.2 9 bp overlap
T 1 dataset
ChIP H9_ENDODERM GSE60606.T.H9_ENDODERM 179 bp overlap
TCF7L1 1 dataset
Motif DE_60h DE_60h-TCF7L1_MA1421.1 12 bp overlap
TFAP2A 1 dataset
ChIP WA09 GSE105081.TFAP2A.WA09 146 bp overlap
TFAP2C 1 dataset
ChIP HCC1954 GSE126897.TFAP2C.HCC1954 153 bp overlap
TFCP2 2 datasets
Motif DE_60h DE_60h-TFCP2_MA1968.2 9 bp overlap
Motif DE_72h DE_72h-TFCP2_MA1968.2 9 bp overlap
TLE3 1 dataset
ChIP LNCaP GSE94682.TLE3.LNCaP 165 bp overlap
TRPS1 2 datasets
Motif DE_60h DE_60h-TRPS1_MA1970.2 8 bp overlap
Motif DE_60h DE_60h-TRPS1_MA1970.2 8 bp overlap
Thap11 1 dataset
Motif DE_60h DE_60h-Thap11_MA1573.2 14 bp overlap
YBX1 1 dataset
ChIP Hep-G2 ENCSR799GOY.YBX1.Hep-G2 208 bp overlap
ZIC1 3 datasets
Motif DE_48h DE_48h-ZIC1_MA0696.1 14 bp overlap
Motif DE_60h DE_60h-ZIC1_MA0696.1 14 bp overlap
Motif DE_72h DE_72h-ZIC1_MA0696.1 14 bp overlap
ZIC4 3 datasets
Motif DE_48h DE_48h-ZIC4_MA0751.2 14 bp overlap
Motif DE_60h DE_60h-ZIC4_MA0751.2 14 bp overlap
Motif DE_72h DE_72h-ZIC4_MA0751.2 14 bp overlap
ZNF135 2 datasets
Motif DE_48h DE_48h-ZNF135_MA1587.1 14 bp overlap
Motif DE_60h DE_60h-ZNF135_MA1587.1 14 bp overlap
ZNF189 1 dataset
Motif DE_60h DE_60h-ZNF189_MA1725.2 9 bp overlap
ZNF24 2 datasets
Motif DE_60h DE_60h-ZNF24_MA1124.1 13 bp overlap
Motif DE_60h DE_60h-ZNF24_MA1124.1 13 bp overlap
ZNF282 2 datasets
Motif DE_60h DE_60h-ZNF282_MA1154.2 15 bp overlap
ChIP Hep-G2 ENCSR752NDX.ZNF282.Hep-G2 148 bp overlap
ZNF384 2 datasets
Motif DE_60h DE_60h-ZNF384_MA1125.2 8 bp overlap
Motif DE_72h DE_72h-ZNF384_MA1125.2 8 bp overlap
ZNF410 1 dataset
Motif DE_60h DE_60h-ZNF410_MA0752.2 16 bp overlap
ZNF418 1 dataset
Motif DE_60h DE_60h-ZNF418_MA1980.1 15 bp overlap
ZNF454 2 datasets
Motif DE_48h DE_48h-ZNF454_MA1712.2 17 bp overlap
Motif DE_60h DE_60h-ZNF454_MA1712.2 17 bp overlap
ZNF460 3 datasets
Motif DE_48h DE_48h-ZNF460_MA1596.1 16 bp overlap
Motif DE_60h DE_60h-ZNF460_MA1596.1 16 bp overlap
Motif DE_72h DE_72h-ZNF460_MA1596.1 16 bp overlap
ZNF528 1 dataset
Motif DE_60h DE_60h-ZNF528_MA1597.1 17 bp overlap
ZNF558 1 dataset
Motif DE_60h DE_60h-ZNF558_MA2335.1 29 bp overlap
ZNF582 1 dataset
Motif DE_60h DE_60h-ZNF582_MA1983.2 19 bp overlap
ZNF701 2 datasets
Motif DE_60h DE_60h-ZNF701_MA1987.2 17 bp overlap
Motif DE_60h DE_60h-ZNF701_MA1987.2 17 bp overlap
ZNF750 1 dataset
ChIP keratinocyte_diff GSE57702.ZNF750.keratinocyte_diff 225 bp overlap
ZNF768 1 dataset
Motif DE_60h DE_60h-ZNF768_MA1731.2 9 bp overlap
ZSCAN21 1 dataset
Motif DE_60h DE_60h-ZSCAN21_MA2336.1 7 bp overlap
Zfp335 1 dataset
Motif DE_60h DE_60h-Zfp335_MA2002.2 7 bp overlap
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