chr4 : 140,640,083 140,640,873
790 bp 115 TFs 5 linked genes
This 790 bp open chromatin element is linked to 5 target genes and is bound by 115 transcription factors.
Linked Genes
5 genes
Distance
Link type
Gene Expression Dist. to TSS Distance Link type
TNRC18P1 4.6 kb Proximal Proximity
TBC1D9 115.9 kb Distal Multiome
ELMOD2 116.3 kb Distal Multiome
CLGN 212.8 kb Distal Multiome
SCOC 266.8 kb Distal Multiome
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr4:140,635,083 – 140,645,873
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
115 transcription factors
Source
Cell type
ASH2L 1 dataset
ChIP Hep-G2 ENCSR508LMH.ASH2L.Hep-G2 327 bp overlap
ATF2 1 dataset
Motif DE_36h DE_36h-ATF2_MA1632.2 10 bp overlap
ATF4 1 dataset
ChIP HepG2 ENCFF903ADR 129 bp overlap
BACH2 1 dataset
Motif DE_36h DE_36h-BACH2_MA1470.2 19 bp overlap
BCL6 4 datasets
Motif DE_36h DE_36h-BCL6_MA0463.3 13 bp overlap
Motif DE_48h DE_48h-BCL6_MA0463.3 13 bp overlap
Motif DE_60h DE_60h-BCL6_MA0463.3 13 bp overlap
ChIP HepG2 ENCFF423EJH 162 bp overlap
CCAR2 1 dataset
ChIP Hep-G2 ENCSR247XFV.CCAR2.Hep-G2 164 bp overlap
CEBPD 2 datasets
Motif DE_36h DE_36h-CEBPD_MA0836.3 8 bp overlap
Motif DE_60h DE_60h-CEBPD_MA0836.3 8 bp overlap
CTCFL 3 datasets
Motif DE_36h DE_36h-CTCFL_MA1102.3 8 bp overlap
Motif DE_48h DE_48h-CTCFL_MA1102.3 8 bp overlap
Motif DE_60h DE_60h-CTCFL_MA1102.3 8 bp overlap
DMAP1 1 dataset
ChIP Hep-G2 ENCSR670YPQ.DMAP1.Hep-G2 221 bp overlap
Dmrt1 3 datasets
Motif DE_36h DE_36h-Dmrt1_MA1603.2 9 bp overlap
Motif DE_48h DE_48h-Dmrt1_MA1603.2 9 bp overlap
Motif DE_60h DE_60h-Dmrt1_MA1603.2 9 bp overlap
EBF1 3 datasets
Motif DE_36h DE_36h-EBF1_MA0154.5 11 bp overlap
Motif DE_48h DE_48h-EBF1_MA0154.5 11 bp overlap
Motif DE_60h DE_60h-EBF1_MA0154.5 11 bp overlap
EBF3 3 datasets
Motif DE_36h DE_36h-EBF3_MA1637.2 9 bp overlap
Motif DE_48h DE_48h-EBF3_MA1637.2 9 bp overlap
Motif DE_60h DE_60h-EBF3_MA1637.2 9 bp overlap
EOMES 1 dataset
ChIP hESC GSE26097.EOMES.hESC 378 bp overlap
EP300 1 dataset
ChIP Hep-G2 ENCSR000BLW.EP300.Hep-G2 262 bp overlap
ERF::FOXO1 2 datasets
Motif DE_36h DE_36h-ERFFOXO1_MA1936.2 12 bp overlap
Motif DE_60h DE_60h-ERFFOXO1_MA1936.2 12 bp overlap
ESR1_pS118 2 datasets
ChIP MCF-7_E2_sc GSE117569.ESR1_pS118.MCF-7_E2_sc 176 bp overlap
ChIP MCF-7_Veh_sc GSE117569.ESR1_pS118.MCF-7_Veh_sc 183 bp overlap
Ebf2 3 datasets
Motif DE_36h DE_36h-Ebf2_MA1604.2 9 bp overlap
Motif DE_48h DE_48h-Ebf2_MA1604.2 9 bp overlap
Motif DE_60h DE_60h-Ebf2_MA1604.2 9 bp overlap
Ebf4 3 datasets
Motif DE_36h DE_36h-Ebf4_MA2122.1 11 bp overlap
Motif DE_48h DE_48h-Ebf4_MA2122.1 11 bp overlap
Motif DE_60h DE_60h-Ebf4_MA2122.1 11 bp overlap
FOXA1 2 datasets
ChIP breast-cancer_Veh-2856 GSE128018.FOXA1.breast-cancer_Veh-2856 264 bp overlap
ChIP primary-breast-cancer_B2_DSG GSE114737.FOXA1.primary-breast-cancer_B2_DSG 310 bp overlap
FOXA2 3 datasets
ChIP DE DE-FOXA2-1 790 bp overlap
ChIP DE DE-FOXA2-2 790 bp overlap
ChIP HepG2 ENCFF533COJ 297 bp overlap
FOXB1 3 datasets
Motif DE_36h DE_36h-FOXB1_MA0845.1 11 bp overlap
Motif DE_48h DE_48h-FOXB1_MA0845.1 11 bp overlap
Motif DE_60h DE_60h-FOXB1_MA0845.1 11 bp overlap
FOXD2 3 datasets
Motif DE_36h DE_36h-FOXD2_MA0847.4 11 bp overlap
Motif DE_48h DE_48h-FOXD2_MA0847.4 11 bp overlap
Motif DE_60h DE_60h-FOXD2_MA0847.4 11 bp overlap
FOXD3 3 datasets
Motif DE_36h DE_36h-FOXD3_MA0041.3 14 bp overlap
Motif DE_48h DE_48h-FOXD3_MA0041.3 14 bp overlap
Motif DE_60h DE_60h-FOXD3_MA0041.3 14 bp overlap
FOXO1::ELK1 2 datasets
Motif DE_36h DE_36h-FOXO1ELK1_MA1954.2 13 bp overlap
Motif DE_60h DE_60h-FOXO1ELK1_MA1954.2 13 bp overlap
Foxl2 3 datasets
Motif DE_36h DE_36h-Foxl2_MA1607.2 10 bp overlap
Motif DE_48h DE_48h-Foxl2_MA1607.2 10 bp overlap
Motif DE_60h DE_60h-Foxl2_MA1607.2 10 bp overlap
GATA2 3 datasets
Motif DE_36h DE_36h-GATA2_MA0036.4 7 bp overlap
Motif DE_48h DE_48h-GATA2_MA0036.4 7 bp overlap
ChIP Hep-G2 ENCSR897LDT.GATA2.Hep-G2 260 bp overlap
GATA3 1 dataset
ChIP breast_tumor_Male_21 GSE104399.GATA3.breast_tumor_Male_21 413 bp overlap
GATA4 5 datasets
ChIP DE DE-GATA4-1 724 bp overlap
ChIP DE DE-GATA4-2 731 bp overlap
Motif DE_36h DE_36h-GATA4_MA0482.3 8 bp overlap
Motif DE_48h DE_48h-GATA4_MA0482.3 8 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA4.pancreatic-progenitor 762 bp overlap
GATA5 2 datasets
Motif DE_36h DE_36h-GATA5_MA0766.3 8 bp overlap
Motif DE_48h DE_48h-GATA5_MA0766.3 8 bp overlap
GATA6 12 datasets
ChIP DE DE-GATA6-1 758 bp overlap
ChIP DE DE-GATA6-2 730 bp overlap
Motif DE_36h DE_36h-GATA6_MA1104.3 8 bp overlap
Motif DE_48h DE_48h-GATA6_MA1104.3 8 bp overlap
ChIP DE_D1 S09-DE-d1-GATA6-exp1 560 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 590 bp overlap
ChIP DE_D1 S41-DE-d1-GATA6-exp2 481 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 651 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 656 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 566 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 605 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA6.pancreatic-progenitor 414 bp overlap
GFI1 1 dataset
ChIP HepG2 ENCFF472INF 557 bp overlap
Gata3 2 datasets
Motif DE_36h DE_36h-Gata3_MA0037.5 8 bp overlap
Motif DE_48h DE_48h-Gata3_MA0037.5 8 bp overlap
HNRNPK 3 datasets
ChIP Hep-G2 GSE120104.HNRNPK.Hep-G2 561 bp overlap
ChIP Hep-G2 ENCSR519QAA.HNRNPK.Hep-G2 510 bp overlap
ChIP HepG2 ENCFF826MXP 505 bp overlap
IKZF1 2 datasets
Motif DE_36h DE_36h-IKZF1_MA1508.2 8 bp overlap
Motif DE_60h DE_60h-IKZF1_MA1508.2 8 bp overlap
ISL2 1 dataset
ChIP Hep-G2 ENCSR492IHH.ISL2.Hep-G2 252 bp overlap
JUN 1 dataset
ChIP DE_D2 S02-DE-d2-JUN-exp1 381 bp overlap
KLF1 2 datasets
Motif DE_36h DE_36h-KLF1_MA0493.3 8 bp overlap
Motif DE_48h DE_48h-KLF1_MA0493.3 8 bp overlap
KLF10 2 datasets
Motif DE_36h DE_36h-KLF10_MA1511.2 9 bp overlap
Motif DE_48h DE_48h-KLF10_MA1511.2 9 bp overlap
KLF14 2 datasets
Motif DE_36h DE_36h-KLF14_MA0740.2 9 bp overlap
Motif DE_48h DE_48h-KLF14_MA0740.2 9 bp overlap
KLF2 2 datasets
Motif DE_36h DE_36h-KLF2_MA1515.2 8 bp overlap
Motif DE_48h DE_48h-KLF2_MA1515.2 8 bp overlap
KLF4 2 datasets
Motif DE_36h DE_36h-KLF4_MA0039.5 8 bp overlap
Motif DE_48h DE_48h-KLF4_MA0039.5 8 bp overlap
KLF5 2 datasets
Motif DE_36h DE_36h-KLF5_MA0599.1 10 bp overlap
Motif DE_48h DE_48h-KLF5_MA0599.1 10 bp overlap
MAX 1 dataset
ChIP Hep-G2 ENCSR168DYA.MAX.Hep-G2 282 bp overlap
MEIS1 4 datasets
Motif DE_36h DE_36h-MEIS1_MA0498.3 5 bp overlap
Motif DE_36h DE_36h-MEIS1_MA0498.3 5 bp overlap
Motif DE_48h DE_48h-MEIS1_MA0498.3 5 bp overlap
Motif DE_60h DE_60h-MEIS1_MA0498.3 5 bp overlap
MGA::EVX1 3 datasets
Motif DE_36h DE_36h-MGAEVX1_MA1960.2 11 bp overlap
Motif DE_48h DE_48h-MGAEVX1_MA1960.2 11 bp overlap
Motif DE_60h DE_60h-MGAEVX1_MA1960.2 11 bp overlap
MSC 3 datasets
Motif DE_36h DE_36h-MSC_MA0665.1 10 bp overlap
Motif DE_48h DE_48h-MSC_MA0665.1 10 bp overlap
Motif DE_60h DE_60h-MSC_MA0665.1 10 bp overlap
MXI1 1 dataset
Motif DE_36h DE_36h-MXI1_MA1108.3 6 bp overlap
MYBL2 1 dataset
ChIP Hep-G2 ENCSR581KCO.MYBL2.Hep-G2 184 bp overlap
MYF6 3 datasets
Motif DE_36h DE_36h-MYF6_MA0667.1 10 bp overlap
Motif DE_48h DE_48h-MYF6_MA0667.1 10 bp overlap
Motif DE_60h DE_60h-MYF6_MA0667.1 10 bp overlap
NR2C1 1 dataset
Motif DE_36h DE_36h-NR2C1_MA1535.2 6 bp overlap
NR2C2 1 dataset
Motif DE_36h DE_36h-NR2C2_MA1536.2 6 bp overlap
NR2F2 1 dataset
Motif DE_36h DE_36h-NR2F2_MA1111.2 7 bp overlap
NR2F6 1 dataset
ChIP Hep-G2 ENCSR518WPL.NR2F6.Hep-G2 292 bp overlap
NR4A2 1 dataset
Motif DE_36h DE_36h-NR4A2_MA0160.3 8 bp overlap
Nfatc2 2 datasets
Motif DE_36h DE_36h-Nfatc2_MA0152.3 8 bp overlap
Motif DE_60h DE_60h-Nfatc2_MA0152.3 8 bp overlap
Nr1H2 1 dataset
Motif DE_36h DE_36h-Nr1H2_MA1996.2 6 bp overlap
Nr1H4 1 dataset
Motif DE_36h DE_36h-Nr1H4_MA1110.3 6 bp overlap
Nr1h3 1 dataset
Motif DE_36h DE_36h-Nr1h3_MA2337.1 6 bp overlap
PATZ1 4 datasets
Motif DE_36h DE_36h-PATZ1_MA1961.2 11 bp overlap
Motif DE_48h DE_48h-PATZ1_MA1961.2 11 bp overlap
ChIP HepG2 ENCFF723PFC 155 bp overlap
ChIP HepG2 ENCFF723PFC 345 bp overlap
PCBP2 3 datasets
ChIP Hep-G2 ENCSR945NSF.PCBP2.Hep-G2 258 bp overlap
ChIP Hep-G2 GSE120104.PCBP2.Hep-G2 258 bp overlap
ChIP HepG2 ENCFF033VWK 402 bp overlap
PLAG1 3 datasets
Motif DE_36h DE_36h-PLAG1_MA0163.1 14 bp overlap
Motif DE_48h DE_48h-PLAG1_MA0163.1 14 bp overlap
Motif DE_60h DE_60h-PLAG1_MA0163.1 14 bp overlap
POU1F1 3 datasets
Motif DE_36h DE_36h-POU1F1_MA0784.3 14 bp overlap
Motif DE_48h DE_48h-POU1F1_MA0784.3 14 bp overlap
Motif DE_60h DE_60h-POU1F1_MA0784.3 14 bp overlap
POU2F1 3 datasets
Motif DE_36h DE_36h-POU2F1_MA0785.2 9 bp overlap
Motif DE_48h DE_48h-POU2F1_MA0785.2 9 bp overlap
Motif DE_60h DE_60h-POU2F1_MA0785.2 9 bp overlap
POU2F2 3 datasets
Motif DE_36h DE_36h-POU2F2_MA0507.3 13 bp overlap
Motif DE_48h DE_48h-POU2F2_MA0507.3 13 bp overlap
Motif DE_60h DE_60h-POU2F2_MA0507.3 13 bp overlap
POU2F3 3 datasets
Motif DE_36h DE_36h-POU2F3_MA0627.3 9 bp overlap
Motif DE_48h DE_48h-POU2F3_MA0627.3 9 bp overlap
Motif DE_60h DE_60h-POU2F3_MA0627.3 9 bp overlap
POU3F2 3 datasets
Motif DE_36h DE_36h-POU3F2_MA0787.1 12 bp overlap
Motif DE_48h DE_48h-POU3F2_MA0787.1 12 bp overlap
Motif DE_60h DE_60h-POU3F2_MA0787.1 12 bp overlap
POU3F3 3 datasets
Motif DE_36h DE_36h-POU3F3_MA0788.1 13 bp overlap
Motif DE_48h DE_48h-POU3F3_MA0788.1 13 bp overlap
Motif DE_60h DE_60h-POU3F3_MA0788.1 13 bp overlap
POU3F4 3 datasets
Motif DE_36h DE_36h-POU3F4_MA0789.1 9 bp overlap
Motif DE_48h DE_48h-POU3F4_MA0789.1 9 bp overlap
Motif DE_60h DE_60h-POU3F4_MA0789.1 9 bp overlap
POU5F1 4 datasets
Motif DE_36h DE_36h-POU5F1_MA1115.2 7 bp overlap
Motif DE_48h DE_48h-POU5F1_MA1115.2 7 bp overlap
Motif DE_60h DE_60h-POU5F1_MA1115.2 7 bp overlap
ChIP DE_D1 DED1-OCT4_Batch_II 408 bp overlap
POU5F1B 3 datasets
Motif DE_36h DE_36h-POU5F1B_MA0792.1 9 bp overlap
Motif DE_48h DE_48h-POU5F1B_MA0792.1 9 bp overlap
Motif DE_60h DE_60h-POU5F1B_MA0792.1 9 bp overlap
POU5F1_M 1 dataset
ChIP DE_D1 DED1-OCT4-M_Batch_II 363 bp overlap
PPARG 1 dataset
ChIP Hep-G2 ENCSR130VQL.PPARG.Hep-G2 227 bp overlap
PRDM9 3 datasets
Motif DE_36h DE_36h-PRDM9_MA1723.2 20 bp overlap
Motif DE_36h DE_36h-PRDM9_MA1723.2 20 bp overlap
Motif DE_48h DE_48h-PRDM9_MA1723.2 20 bp overlap
Ppara 1 dataset
Motif DE_36h DE_36h-Ppara_MA2338.1 7 bp overlap
RREB1 2 datasets
Motif DE_36h DE_36h-RREB1_MA0073.2 19 bp overlap
Motif DE_48h DE_48h-RREB1_MA0073.2 19 bp overlap
Rhox11 3 datasets
Motif DE_36h DE_36h-Rhox11_MA0629.2 9 bp overlap
Motif DE_48h DE_48h-Rhox11_MA0629.2 9 bp overlap
Motif DE_60h DE_60h-Rhox11_MA0629.2 9 bp overlap
Runx1 3 datasets
Motif DE_36h DE_36h-Runx1_MA0002.3 9 bp overlap
Motif DE_48h DE_48h-Runx1_MA0002.3 9 bp overlap
Motif DE_60h DE_60h-Runx1_MA0002.3 9 bp overlap
SMAD2 1 dataset
ChIP endoderm GSE29422.SMAD2.endoderm 157 bp overlap
SMAD2-3 3 datasets
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 790 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.SMAD2-3.HUES-8_DE_JNKi 718 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.SMAD2-3.HUES-8_DE_ctrl 277 bp overlap
SMAD2_3 3 datasets
ChIP DE_D1 S12-DE-d1-SMAD2_3-exp1 663 bp overlap
ChIP DE_D2 S01-DE-d2-SMAD2_3-exp1 790 bp overlap
ChIP DE_D2 S53-DE-d2-SMAD2_3-exp2 733 bp overlap
SMAD4 1 dataset
ChIP endoderm GSE29422.SMAD4.endoderm 176 bp overlap
SMARCB1 1 dataset
ChIP Hep-G2 GSE69566.SMARCB1.Hep-G2 411 bp overlap
SOX17 1 dataset
ChIP DE_D2 DED2-SOX17_Batch_II 444 bp overlap
SOX17_M 1 dataset
ChIP DE_D2 DED2-SOX17-M_Batch_II 510 bp overlap
SOX6 1 dataset
ChIP Hep-G2 ENCSR766TSU.SOX6.Hep-G2 225 bp overlap
SP5 3 datasets
Motif DE_36h DE_36h-SP5_MA1965.2 6 bp overlap
Motif DE_48h DE_48h-SP5_MA1965.2 6 bp overlap
Motif DE_60h DE_60h-SP5_MA1965.2 6 bp overlap
SP8 2 datasets
Motif DE_36h DE_36h-SP8_MA0747.2 11 bp overlap
Motif DE_48h DE_48h-SP8_MA0747.2 11 bp overlap
Smad4 3 datasets
Motif DE_36h DE_36h-Smad4_MA1153.2 7 bp overlap
Motif DE_48h DE_48h-Smad4_MA1153.2 7 bp overlap
Motif DE_60h DE_60h-Smad4_MA1153.2 7 bp overlap
Sox11 2 datasets
Motif DE_36h DE_36h-Sox11_MA0869.3 8 bp overlap
Motif DE_60h DE_60h-Sox11_MA0869.3 8 bp overlap
Sox6 2 datasets
Motif DE_36h DE_36h-Sox6_MA0515.1 10 bp overlap
Motif DE_60h DE_60h-Sox6_MA0515.1 10 bp overlap
TCF7 1 dataset
ChIP HepG2 ENCFF628OFQ 357 bp overlap
TFAP4 2 datasets
Motif DE_36h DE_36h-TFAP4_MA1570.1 10 bp overlap
ChIP HepG2 ENCFF932XOY 397 bp overlap
TFDP2 1 dataset
ChIP Hep-G2 ENCSR069JKP.TFDP2.Hep-G2 117 bp overlap
TFE3 1 dataset
ChIP Hep-G2 ENCSR589SNT.TFE3.Hep-G2 173 bp overlap
TRPS1 2 datasets
Motif DE_36h DE_36h-TRPS1_MA1970.2 8 bp overlap
Motif DE_48h DE_48h-TRPS1_MA1970.2 8 bp overlap
Tcf21 3 datasets
Motif DE_36h DE_36h-Tcf21_MA0832.2 10 bp overlap
Motif DE_48h DE_48h-Tcf21_MA0832.2 10 bp overlap
Motif DE_60h DE_60h-Tcf21_MA0832.2 10 bp overlap
VEZF1 2 datasets
Motif DE_36h DE_36h-VEZF1_MA1578.2 6 bp overlap
Motif DE_48h DE_48h-VEZF1_MA1578.2 6 bp overlap
Wt1 5 datasets
Motif DE_36h DE_36h-Wt1_MA1627.2 10 bp overlap
Motif DE_36h DE_36h-Wt1_MA1627.2 10 bp overlap
Motif DE_48h DE_48h-Wt1_MA1627.2 10 bp overlap
Motif DE_48h DE_48h-Wt1_MA1627.2 10 bp overlap
Motif DE_60h DE_60h-Wt1_MA1627.2 10 bp overlap
ZGPAT 1 dataset
ChIP Hep-G2 ENCSR704IGU.ZGPAT.Hep-G2 173 bp overlap
ZNF219 1 dataset
ChIP HepG2 ENCFF266JIR 431 bp overlap
ZNF232 3 datasets
ChIP Hep-G2 ENCSR837GLU.ZNF232.Hep-G2 431 bp overlap
ChIP HepG2 ENCFF905UTT 441 bp overlap
ChIP HepG2 ENCFF905UTT 441 bp overlap
ZNF282 1 dataset
Motif DE_36h DE_36h-ZNF282_MA1154.2 15 bp overlap
ZNF320 1 dataset
Motif DE_36h DE_36h-ZNF320_MA1976.2 20 bp overlap
ZNF532 1 dataset
ChIP NMC24335 GSE96775.ZNF532.NMC24335 293 bp overlap
ZNF614 1 dataset
ChIP HepG2 ENCFF677IUD 161 bp overlap
ZNF680 3 datasets
Motif DE_36h DE_36h-ZNF680_MA1729.2 11 bp overlap
Motif DE_48h DE_48h-ZNF680_MA1729.2 11 bp overlap
Motif DE_60h DE_60h-ZNF680_MA1729.2 11 bp overlap
ZNF740 2 datasets
Motif DE_36h DE_36h-ZNF740_MA0753.3 10 bp overlap
Motif DE_48h DE_48h-ZNF740_MA0753.3 10 bp overlap
ZNF75A 1 dataset
Motif DE_36h DE_36h-ZNF75A_MA2097.1 12 bp overlap
ZNF766 1 dataset
ChIP K562 ENCFF348LDO 382 bp overlap
ZNF770 3 datasets
Motif DE_36h DE_36h-ZNF770_MA2099.1 8 bp overlap
Motif DE_48h DE_48h-ZNF770_MA2099.1 8 bp overlap
Motif DE_60h DE_60h-ZNF770_MA2099.1 8 bp overlap
ZSCAN4 3 datasets
Motif DE_36h DE_36h-ZSCAN4_MA1155.1 15 bp overlap
Motif DE_48h DE_48h-ZSCAN4_MA1155.1 15 bp overlap
Motif DE_60h DE_60h-ZSCAN4_MA1155.1 15 bp overlap