chr4 : 104,493,172 104,494,092
920 bp 96 TFs 2 linked genes
This 920 bp open chromatin element is linked to CXXC4 and CXXC4-AS1 and is bound by 96 transcription factors.
Linked Genes
2 genes
Distance
Gene Expression Dist. to TSS Distance Link type
CXXC4 808 bp At TSS Proximity
CXXC4-AS1 2.2 kb Proximal Proximity
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr4:104,488,172 – 104,499,092
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
96 transcription factors
Source
Cell type
ATF2 1 dataset
ChIP GM12878 ENCFF521LQJ 61 bp overlap
BCOR 4 datasets
ChIP WA01 GSE104690.BCOR.WA01 348 bp overlap
ChIP WA01_KDM2B_KO GSE104690.BCOR.WA01_KDM2B_KO 228 bp overlap
ChIP WA01_RNF2-R GSE104690.BCOR.WA01_RNF2-R 265 bp overlap
ChIP WA01_RNF2-R_Doxycyclin GSE104690.BCOR.WA01_RNF2-R_Doxycyclin 400 bp overlap
BRD4 11 datasets
ChIP Kelly_resistant GSE103030.BRD4.Kelly_resistant 177 bp overlap
ChIP Kelly_resistant_JQ1 GSE103030.BRD4.Kelly_resistant_JQ1 176 bp overlap
ChIP Kelly_sensitive GSE103030.BRD4.Kelly_sensitive 475 bp overlap
ChIP MM1-S GSE45984.BRD4.MM1-S 712 bp overlap
ChIP MM1-S GSE43743.BRD4.MM1-S 292 bp overlap
ChIP MM1-S GSE43743.BRD4.MM1-S 162 bp overlap
ChIP SK-N-BE2-C_Naive-veh GSE107706.BRD4.SK-N-BE2-C_Naive-veh 830 bp overlap
ChIP SK-N-BE2-C_Resistant-veh GSE107706.BRD4.SK-N-BE2-C_Resistant-veh 920 bp overlap
ChIP SUM159PT_shMYC_dox_48h GSE87418.BRD4.SUM159PT_shMYC_dox_48h 204 bp overlap
ChIP retina_AB1-FW20 GSE86981.BRD4.retina_AB1-FW20 243 bp overlap
ChIP retina_AB1-FW23 GSE86981.BRD4.retina_AB1-FW23 348 bp overlap
CRX 1 dataset
ChIP retina_Hu6 GSE137311.CRX.retina_Hu6 425 bp overlap
Crx 2 datasets
Motif DE_12h DE_12h-Crx_MA0467.3 6 bp overlap
Motif DE_24h DE_24h-Crx_MA0467.3 6 bp overlap
DUX4 1 dataset
ChIP WA01 GSE94322.DUX4.WA01 310 bp overlap
EOMES 1 dataset
ChIP hESC GSE26097.EOMES.hESC 271 bp overlap
EP300 2 datasets
ChIP neural ENCSR843ZUP.EP300.neural 471 bp overlap
ChIP neural cell ENCFF442QNK 517 bp overlap
ETS1 3 datasets
ChIP cardiomyocyte_D2 GSE129986.ETS1.cardiomyocyte_D2 168 bp overlap
ChIP cardiomyocyte_D9 GSE129986.ETS1.cardiomyocyte_D9 129 bp overlap
ChIP hESC ENCSR534VHI.ETS1.hESC 175 bp overlap
FEZF2 1 dataset
Motif DE_12h DE_12h-FEZF2_MA2341.1 8 bp overlap
FOXA1 2 datasets
ChIP PDAC GSE64557.FOXA1.PDAC 67 bp overlap
ChIP VCaP_DHT24H GSE58428.FOXA1.VCaP_DHT24H 247 bp overlap
FOXA2 3 datasets
ChIP DE DE-FOXA2-1 496 bp overlap
ChIP DE DE-FOXA2-2 570 bp overlap
ChIP HepG2 ENCFF570ABM 405 bp overlap
GATA2 2 datasets
ChIP SH-SY5Y ENCSR000EYB.GATA2.SH-SY5Y 225 bp overlap
ChIP SK-N-SH ENCSR706VOO.GATA2.SK-N-SH 144 bp overlap
GATA4 8 datasets
ChIP DE DE-GATA4-1 439 bp overlap
ChIP DE DE-GATA4-2 521 bp overlap
ChIP G296S GSE85628.GATA4.G296S 266 bp overlap
ChIP G296S_2 GSE85628.GATA4.G296S_2 266 bp overlap
ChIP cardiomyocyte_5 GSE85628.GATA4.cardiomyocyte_5 199 bp overlap
ChIP foregut GSE117136.GATA4.foregut 366 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA4.pancreatic-progenitor 544 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA4.pancreatic-progenitor_PP1 477 bp overlap
GATA6 13 datasets
ChIP DE DE-GATA6-1 376 bp overlap
ChIP DE DE-GATA6-2 556 bp overlap
ChIP DE_D1 S09-DE-d1-GATA6-exp1 688 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 671 bp overlap
ChIP DE_D1 S41-DE-d1-GATA6-exp2 727 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 448 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 757 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 773 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 402 bp overlap
ChIP foregut GSE117136.GATA6.foregut 300 bp overlap
ChIP foregut_KO GSE117136.GATA6.foregut_KO 436 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA6.pancreatic-progenitor 364 bp overlap
ChIP pancreatic-progenitor_KO GSE117136.GATA6.pancreatic-progenitor_KO 431 bp overlap
GFI1 3 datasets
Motif DE_12h DE_12h-GFI1_MA0038.3 11 bp overlap
Motif DE_24h DE_24h-GFI1_MA0038.3 11 bp overlap
Motif ES_0h ES_0h-GFI1_MA0038.3 11 bp overlap
GSC 2 datasets
Motif DE_12h DE_12h-GSC_MA0648.2 6 bp overlap
Motif DE_24h DE_24h-GSC_MA0648.2 6 bp overlap
GSC2 2 datasets
Motif DE_12h DE_12h-GSC2_MA0891.2 6 bp overlap
Motif DE_24h DE_24h-GSC2_MA0891.2 6 bp overlap
Gfi1B 3 datasets
Motif DE_12h DE_12h-Gfi1B_MA0483.2 10 bp overlap
Motif DE_24h DE_24h-Gfi1B_MA0483.2 10 bp overlap
Motif ES_0h ES_0h-Gfi1B_MA0483.2 10 bp overlap
HOXA9 1 dataset
ChIP HEK293-FT GSE62586.HOXA9.HEK293-FT 117 bp overlap
HOXB13 1 dataset
ChIP G-401 GSE65381.HOXB13.G-401 357 bp overlap
HSF1 1 dataset
Motif DE_12h DE_12h-HSF1_MA0486.2 13 bp overlap
HSF2 1 dataset
Motif DE_12h DE_12h-HSF2_MA0770.1 13 bp overlap
HSF4 1 dataset
Motif DE_12h DE_12h-HSF4_MA0771.1 13 bp overlap
IKZF1 1 dataset
ChIP GM12878 ENCFF824TGK 627 bp overlap
JUN 2 datasets
ChIP DE_D1 S40-DE-d1-JUN-exp2 358 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.JUN.HUES-8_DE_ctrl 359 bp overlap
KDM1A 1 dataset
ChIP pancreatic-progenitor_GT GSE104840.KDM1A.pancreatic-progenitor_GT 246 bp overlap
KLF5 1 dataset
ChIP HEK293_E419K GSE88976.KLF5.HEK293_E419K 182 bp overlap
KMT2A 1 dataset
ChIP HEK293T_N-term_C104 GSE90762.KMT2A.HEK293T_N-term_C104 175 bp overlap
MED1 3 datasets
ChIP G296S_4 GSE85628.MED1.G296S_4 320 bp overlap
ChIP G296S_4 GSE85628.MED1.G296S_4 181 bp overlap
ChIP MM1-S_JQ1 GSE42161.MED1.MM1-S_JQ1 784 bp overlap
MXI1 2 datasets
ChIP neural ENCSR934NHU.MXI1.neural 353 bp overlap
ChIP neural cell ENCFF623HQN 577 bp overlap
MYCN 1 dataset
ChIP COG-N-415 GSE138295.MYCN.COG-N-415 326 bp overlap
NANOG 8 datasets
ChIP GM23338 ENCFF065NZG 311 bp overlap
ChIP GM23338 ENCSR061DGF.NANOG.GM23338 155 bp overlap
ChIP HUES-8 GSE109524.NANOG.HUES-8 778 bp overlap
ChIP WA01 ERP004238.NANOG.WA01 231 bp overlap
ChIP WA01_3IL ERP004238.NANOG.WA01_3IL 449 bp overlap
ChIP WA09 GSE105028.NANOG.WA09 383 bp overlap
ChIP WA09_heat-shock GSE105028.NANOG.WA09_heat-shock 305 bp overlap
ChIP hESC GSE18292.NANOG.hESC 107 bp overlap
NR1I3 1 dataset
Motif DE_12h DE_12h-NR1I3_MA1534.2 8 bp overlap
NRIP1 1 dataset
ChIP MCF-7_E2 ERP005838.NRIP1.MCF-7_E2 127 bp overlap
Nr2e3 3 datasets
Motif DE_12h DE_12h-Nr2e3_MA0164.2 6 bp overlap
Motif DE_24h DE_24h-Nr2e3_MA0164.2 6 bp overlap
Motif ES_0h ES_0h-Nr2e3_MA0164.2 6 bp overlap
OTX1 2 datasets
Motif DE_12h DE_12h-OTX1_MA0711.2 6 bp overlap
Motif DE_24h DE_24h-OTX1_MA0711.2 6 bp overlap
OTX2 1 dataset
ChIP retina_Hu7 GSE137311.OTX2.retina_Hu7 375 bp overlap
PHOX2B 2 datasets
ChIP CLB-Ga GSE90683.PHOX2B.CLB-Ga 403 bp overlap
ChIP SK-N-BE2-C GSE94822.PHOX2B.SK-N-BE2-C 244 bp overlap
PITX1 2 datasets
Motif DE_12h DE_12h-PITX1_MA0682.3 6 bp overlap
Motif DE_24h DE_24h-PITX1_MA0682.3 6 bp overlap
PITX3 3 datasets
Motif DE_12h DE_12h-PITX3_MA0714.2 6 bp overlap
Motif DE_24h DE_24h-PITX3_MA0714.2 6 bp overlap
ChIP SH-SY5Y GSE93275.PITX3.SH-SY5Y 920 bp overlap
PKNOX1 3 datasets
Motif DE_12h DE_12h-PKNOX1_MA0782.3 10 bp overlap
Motif DE_24h DE_24h-PKNOX1_MA0782.3 10 bp overlap
Motif ES_0h ES_0h-PKNOX1_MA0782.3 10 bp overlap
POLR2A 2 datasets
ChIP GM23338 ENCFF450WCS 491 bp overlap
ChIP neural cell ENCFF604SPB 385 bp overlap
POU2F1 2 datasets
ChIP 22Rv1_DHT GSE123565.POU2F1.22Rv1_DHT 293 bp overlap
ChIP 22Rv1_DHT GSE123565.POU2F1.22Rv1_DHT 84 bp overlap
POU2F2 2 datasets
ChIP GM12878 ENCFF207RKY 321 bp overlap
ChIP GM12891 ENCSR000BII.POU2F2.GM12891 163 bp overlap
POU5F1 2 datasets
ChIP DE_D1 DED1-OCT4_Batch_II 382 bp overlap
ChIP HUES-8 GSE109524.POU5F1.HUES-8 621 bp overlap
POU5F1_M 1 dataset
ChIP DE_D1 DED1-OCT4-M_Batch_II 308 bp overlap
PRDM14 2 datasets
ChIP hESC GSE22767.PRDM14.hESC 410 bp overlap
ChIP primordial-germ-cell-like-cell GSE138674.PRDM14.primordial-germ-cell-like-cell 360 bp overlap
PSIP1 1 dataset
ChIP T-REx-293 GSE117155.PSIP1.T-REx-293 145 bp overlap
RAD21 2 datasets
ChIP neural cell ENCFF564MOT 203 bp overlap
ChIP neural cell ENCFF564MOT 334 bp overlap
RARA 2 datasets
ChIP hiPSC_D2 GSE132532.RARA.hiPSC_D2 329 bp overlap
ChIP hiPSC_D5 GSE132532.RARA.hiPSC_D5 485 bp overlap
RB1 1 dataset
ChIP GM12878 ENCSR785OKZ.RB1.GM12878 127 bp overlap
RBBP4 1 dataset
ChIP SCMC GSE155861.RBBP4.SCMC 277 bp overlap
RELB 1 dataset
ChIP GM12878 ENCSR387QUV.RELB.GM12878 246 bp overlap
RHOXF1 2 datasets
Motif DE_12h DE_12h-RHOXF1_MA0719.2 6 bp overlap
Motif DE_24h DE_24h-RHOXF1_MA0719.2 6 bp overlap
RNF2 2 datasets
ChIP SYO-1_shCt GSE139053.RNF2.SYO-1_shCt 580 bp overlap
ChIP fibroblast GSE139053.RNF2.fibroblast 170 bp overlap
RUNX3 1 dataset
ChIP GM12878 ENCFF395WHA 371 bp overlap
SMAD2-3 2 datasets
ChIP HUES-8_DE_JNKi GSE109524.SMAD2-3.HUES-8_DE_JNKi 273 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.SMAD2-3.HUES-8_DE_ctrl 313 bp overlap
SMARCA4 5 datasets
ChIP NGP_ARID1A-mut2 GSE134626.SMARCA4.NGP_ARID1A-mut2 58 bp overlap
ChIP NSC GSE125033.SMARCA4.NSC 630 bp overlap
ChIP hiPSC GSE124903.SMARCA4.hiPSC 364 bp overlap
ChIP hiPSC_WT GSE124903.SMARCA4.hiPSC_WT 97 bp overlap
ChIP hiPSC_WT GSE124903.SMARCA4.hiPSC_WT 281 bp overlap
SMARCB1 1 dataset
ChIP hiPSC_WT GSE124903.SMARCB1.hiPSC_WT 240 bp overlap
SMARCC1 2 datasets
ChIP HS-SY-2 GSE108025.SMARCC1.HS-SY-2 308 bp overlap
ChIP hiPSC_WT GSE124903.SMARCC1.hiPSC_WT 182 bp overlap
SMC3 1 dataset
ChIP neural cell ENCFF795YGY 531 bp overlap
SOX2 1 dataset
ChIP HNSC GSE69479.SOX2.HNSC 272 bp overlap
SREBF1 1 dataset
Motif DE_12h DE_12h-SREBF1_MA0595.1 10 bp overlap
SREBF2 1 dataset
Motif DE_12h DE_12h-SREBF2_MA0596.1 10 bp overlap
SS18 1 dataset
ChIP Aska-SS_BAF47KO1 GSE108025.SS18.Aska-SS_BAF47KO1 855 bp overlap
STAT1 1 dataset
Motif DE_12h DE_12h-STAT1_MA0137.4 9 bp overlap
STAT3 1 dataset
Motif DE_12h DE_12h-STAT3_MA0144.3 9 bp overlap
Stat4 1 dataset
Motif DE_12h DE_12h-Stat4_MA0518.2 10 bp overlap
Stat5b 1 dataset
Motif DE_12h DE_12h-Stat5b_MA1625.2 9 bp overlap
T 1 dataset
ChIP H9_ENDODERM GSE60606.T.H9_ENDODERM 277 bp overlap
TAF1 1 dataset
ChIP neural ENCSR000BTX.TAF1.neural 254 bp overlap
TBX2 1 dataset
ChIP Kelly GSE94822.TBX2.Kelly 205 bp overlap
TBX5 1 dataset
ChIP cardiomyocyte_5 GSE85628.TBX5.cardiomyocyte_5 130 bp overlap
TCF12 1 dataset
ChIP WA01 ENCSR000BIT.TCF12.WA01 211 bp overlap
TP53 1 dataset
ChIP H9_ectoderm GSE142050.TP53.H9_ectoderm 254 bp overlap
Vdr 1 dataset
Motif DE_12h DE_12h-Vdr_MA0693.4 7 bp overlap
ZFP42 3 datasets
Motif DE_12h DE_12h-ZFP42_MA1651.2 13 bp overlap
Motif DE_24h DE_24h-ZFP42_MA1651.2 13 bp overlap
Motif ES_0h ES_0h-ZFP42_MA1651.2 13 bp overlap
ZFP64 1 dataset
ChIP HEK293 GSE76494.ZFP64.HEK293 156 bp overlap
ZKSCAN3 1 dataset
Motif DE_12h DE_12h-ZKSCAN3_MA1973.2 14 bp overlap
ZNF184 3 datasets
Motif DE_12h DE_12h-ZNF184_MA2120.1 13 bp overlap
Motif DE_24h DE_24h-ZNF184_MA2120.1 13 bp overlap
Motif ES_0h ES_0h-ZNF184_MA2120.1 13 bp overlap
ZNF213 1 dataset
Motif DE_12h DE_12h-ZNF213_MA2121.1 12 bp overlap
ZNF214 3 datasets
Motif DE_12h DE_12h-ZNF214_MA1975.2 13 bp overlap
Motif DE_24h DE_24h-ZNF214_MA1975.2 13 bp overlap
Motif ES_0h ES_0h-ZNF214_MA1975.2 13 bp overlap
ZNF35 1 dataset
Motif DE_12h DE_12h-ZNF35_MA2333.1 7 bp overlap
ZNF547 1 dataset
Motif DE_12h DE_12h-ZNF547_MA2334.1 13 bp overlap
ZNF816 3 datasets
Motif DE_12h DE_12h-ZNF816_MA1719.2 15 bp overlap
Motif DE_24h DE_24h-ZNF816_MA1719.2 15 bp overlap
Motif ES_0h ES_0h-ZNF816_MA1719.2 15 bp overlap
ZNF85 3 datasets
Motif DE_12h DE_12h-ZNF85_MA1720.2 12 bp overlap
Motif DE_24h DE_24h-ZNF85_MA1720.2 12 bp overlap
Motif ES_0h ES_0h-ZNF85_MA1720.2 12 bp overlap
Zfp335 3 datasets
Motif DE_12h DE_12h-Zfp335_MA2002.2 7 bp overlap
Motif DE_24h DE_24h-Zfp335_MA2002.2 7 bp overlap
Motif ES_0h ES_0h-Zfp335_MA2002.2 7 bp overlap
Zic2 3 datasets
Motif DE_12h DE_12h-Zic2_MA1629.2 9 bp overlap
Motif DE_24h DE_24h-Zic2_MA1629.2 9 bp overlap
Motif ES_0h ES_0h-Zic2_MA1629.2 9 bp overlap