chr4 : 53,923,917 53,924,791
874 bp 114 TFs 1 linked gene
This 874 bp open chromatin element is linked to CHIC2 and is bound by 114 transcription factors.
Linked Genes
1 gene
Gene Expression Dist. to TSS Distance Link type
CHIC2 140.3 kb Distal Multiome
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr4:53,918,917 – 53,929,791
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
114 transcription factors
Source
Cell type
AR 2 datasets
ChIP MCF-7 GSE48930.AR.MCF-7 380 bp overlap
ChIP prostate_fetal-fibroblast GSE90772.AR.prostate_fetal-fibroblast 237 bp overlap
ATF3 1 dataset
ChIP HCT-116_CAMP GSE74355.ATF3.HCT-116_CAMP 106 bp overlap
Atf3 1 dataset
Motif DE_48h DE_48h-Atf3_MA1988.2 7 bp overlap
BACH1 1 dataset
Motif DE_48h DE_48h-BACH1_MA1633.2 9 bp overlap
BATF 1 dataset
Motif DE_48h DE_48h-BATF_MA1634.2 7 bp overlap
BATF3 1 dataset
Motif DE_48h DE_48h-BATF3_MA0835.3 7 bp overlap
BNC2 1 dataset
Motif DE_48h DE_48h-BNC2_MA1928.2 7 bp overlap
BRD1 1 dataset
ChIP HUES-64 GSE104059.BRD1.HUES-64 309 bp overlap
CLOCK 1 dataset
ChIP U2OS GSE44236.CLOCK.U2OS 133 bp overlap
CRY1 1 dataset
ChIP U2OS GSE44236.CRY1.U2OS 201 bp overlap
CTCF 13 datasets
ChIP GSC23 GSE139416.CTCF.GSC23 170 bp overlap
ChIP RWPE-2 ENCSR856JJB.CTCF.RWPE-2 329 bp overlap
ChIP RWPE2 ENCFF911IEE 693 bp overlap
ChIP bonchial_epithelial ENCSR000DXI.CTCF.bonchial_epithelial 129 bp overlap
ChIP epithelial cell of esophagus ENCFF946GGT 305 bp overlap
ChIP epithelial_esophagus ENCSR000DTR.CTCF.epithelial_esophagus 113 bp overlap
ChIP epithelial_mammary ENCSR000DUS.CTCF.epithelial_mammary 147 bp overlap
ChIP foreskin keratinocyte ENCFF980OWR 267 bp overlap
ChIP keratinocyte ENCFF046PBT 185 bp overlap
ChIP keratinocyte ENCFF291YDC 185 bp overlap
ChIP keratinocyte ENCSR000DNC.CTCF.keratinocyte 117 bp overlap
ChIP mammary epithelial cell ENCFF164SPU 265 bp overlap
ChIP primary-epidermal-keratinocyte_diff_d6 GSE84657.CTCF.primary-epidermal-keratinocyte_diff_d6 190 bp overlap
EOMES 3 datasets
Motif DE_36h DE_36h-EOMES_MA0800.2 9 bp overlap
Motif DE_48h DE_48h-EOMES_MA0800.2 9 bp overlap
ChIP hESC GSE26097.EOMES.hESC 296 bp overlap
ERF::FOXI1 2 datasets
Motif DE_36h DE_36h-ERFFOXI1_MA1935.2 10 bp overlap
Motif DE_48h DE_48h-ERFFOXI1_MA1935.2 10 bp overlap
ERG 1 dataset
ChIP AMLPZ12 GSE23730.ERG.AMLPZ12 203 bp overlap
ESR1 3 datasets
ChIP Ishikawa_ETV4-KO1 GSE129803.ESR1.Ishikawa_ETV4-KO1 184 bp overlap
ChIP Ishikawa_siNTC-96h GSE129803.ESR1.Ishikawa_siNTC-96h 213 bp overlap
ChIP MCF-7_jc5856 GSE126004.ESR1.MCF-7_jc5856 57 bp overlap
ETS1 1 dataset
ChIP cardiomyocyte_D2 GSE129986.ETS1.cardiomyocyte_D2 304 bp overlap
ETV2::FOXI1 2 datasets
Motif DE_36h DE_36h-ETV2FOXI1_MA1942.2 11 bp overlap
Motif DE_48h DE_48h-ETV2FOXI1_MA1942.2 11 bp overlap
ETV5::FOXO1 2 datasets
Motif DE_36h DE_36h-ETV5FOXO1_MA1947.2 10 bp overlap
Motif DE_48h DE_48h-ETV5FOXO1_MA1947.2 10 bp overlap
FIGLA 1 dataset
Motif DE_48h DE_48h-FIGLA_MA0820.2 6 bp overlap
FLI1::FOXI1 2 datasets
Motif DE_36h DE_36h-FLI1FOXI1_MA1950.2 11 bp overlap
Motif DE_48h DE_48h-FLI1FOXI1_MA1950.2 11 bp overlap
FOS 1 dataset
ChIP MCF-10A ENCSR000DON.FOS.MCF-10A 165 bp overlap
FOXA2 2 datasets
ChIP DE DE-FOXA2-1 406 bp overlap
ChIP DE DE-FOXA2-2 339 bp overlap
FOXO1::ELF1 2 datasets
Motif DE_36h DE_36h-FOXO1ELF1_MA1953.2 13 bp overlap
Motif DE_48h DE_48h-FOXO1ELF1_MA1953.2 13 bp overlap
FOXO1::ELK1 2 datasets
Motif DE_36h DE_36h-FOXO1ELK1_MA1954.2 13 bp overlap
Motif DE_48h DE_48h-FOXO1ELK1_MA1954.2 13 bp overlap
FOXO1::ELK3 2 datasets
Motif DE_36h DE_36h-FOXO1ELK3_MA1955.2 13 bp overlap
Motif DE_48h DE_48h-FOXO1ELK3_MA1955.2 13 bp overlap
GATA1::TAL1 2 datasets
Motif DE_36h DE_36h-GATA1TAL1_MA0140.3 17 bp overlap
Motif DE_48h DE_48h-GATA1TAL1_MA0140.3 17 bp overlap
GATA2 3 datasets
Motif DE_36h DE_36h-GATA2_MA0036.4 7 bp overlap
Motif DE_48h DE_48h-GATA2_MA0036.4 7 bp overlap
ChIP ME-1_CBFB-MYH11-KD GSE117138.GATA2.ME-1_CBFB-MYH11-KD 225 bp overlap
GATA3 5 datasets
ChIP Kelly GSE65664.GATA3.Kelly 64 bp overlap
ChIP MCF-7 ENCFF178GBS 108 bp overlap
ChIP MCF-7 ENCFF352QVM 141 bp overlap
ChIP MCF-7 ENCFF437NQS 73 bp overlap
ChIP MCF-7 ENCSR000EWS.GATA3.MCF-7 67 bp overlap
GATA4 8 datasets
ChIP BJ1-hTERT_FOXA2_GATA4_Coexp GSE92491.GATA4.BJ1-hTERT_FOXA2_GATA4_Coexp 243 bp overlap
ChIP DE DE-GATA4-1 498 bp overlap
ChIP DE DE-GATA4-2 547 bp overlap
Motif DE_36h DE_36h-GATA4_MA0482.3 8 bp overlap
Motif DE_48h DE_48h-GATA4_MA0482.3 8 bp overlap
ChIP foregut GSE117136.GATA4.foregut 471 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA4.pancreatic-progenitor 386 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA4.pancreatic-progenitor_PP1 263 bp overlap
GATA5 2 datasets
Motif DE_36h DE_36h-GATA5_MA0766.3 8 bp overlap
Motif DE_48h DE_48h-GATA5_MA0766.3 8 bp overlap
GATA6 14 datasets
ChIP AGS GSE51705.GATA6.AGS 289 bp overlap
ChIP DE DE-GATA6-1 537 bp overlap
ChIP DE DE-GATA6-2 508 bp overlap
Motif DE_36h DE_36h-GATA6_MA1104.3 8 bp overlap
Motif DE_48h DE_48h-GATA6_MA1104.3 8 bp overlap
ChIP DE_D1 S09-DE-d1-GATA6-exp1 761 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 633 bp overlap
ChIP DE_D1 S41-DE-d1-GATA6-exp2 612 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 874 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 874 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 851 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 464 bp overlap
ChIP foregut GSE117136.GATA6.foregut 455 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA6.pancreatic-progenitor 333 bp overlap
GRHL2 2 datasets
Motif DE_36h DE_36h-GRHL2_MA1105.3 8 bp overlap
Motif DE_48h DE_48h-GRHL2_MA1105.3 8 bp overlap
Gata3 2 datasets
Motif DE_36h DE_36h-Gata3_MA0037.5 8 bp overlap
Motif DE_48h DE_48h-Gata3_MA0037.5 8 bp overlap
HAND2 1 dataset
Motif DE_48h DE_48h-HAND2_MA1638.2 6 bp overlap
HOXB9 3 datasets
Motif DE_36h DE_36h-HOXB9_MA1503.2 9 bp overlap
Motif DE_48h DE_48h-HOXB9_MA1503.2 9 bp overlap
Motif DE_60h DE_60h-HOXB9_MA1503.2 9 bp overlap
HOXC10 3 datasets
Motif DE_36h DE_36h-HOXC10_MA0905.2 9 bp overlap
Motif DE_48h DE_48h-HOXC10_MA0905.2 9 bp overlap
Motif DE_60h DE_60h-HOXC10_MA0905.2 9 bp overlap
HOXC12 3 datasets
Motif DE_36h DE_36h-HOXC12_MA0906.2 10 bp overlap
Motif DE_48h DE_48h-HOXC12_MA0906.2 10 bp overlap
Motif DE_60h DE_60h-HOXC12_MA0906.2 10 bp overlap
HOXC9 3 datasets
Motif DE_36h DE_36h-HOXC9_MA0485.3 9 bp overlap
Motif DE_48h DE_48h-HOXC9_MA0485.3 9 bp overlap
Motif DE_60h DE_60h-HOXC9_MA0485.3 9 bp overlap
HOXD10 3 datasets
Motif DE_36h DE_36h-HOXD10_MA1506.2 10 bp overlap
Motif DE_48h DE_48h-HOXD10_MA1506.2 10 bp overlap
Motif DE_60h DE_60h-HOXD10_MA1506.2 10 bp overlap
HOXD11 3 datasets
Motif DE_36h DE_36h-HOXD11_MA0908.2 9 bp overlap
Motif DE_48h DE_48h-HOXD11_MA0908.2 9 bp overlap
Motif DE_60h DE_60h-HOXD11_MA0908.2 9 bp overlap
HOXD12::ELK1 3 datasets
Motif DE_36h DE_36h-HOXD12ELK1_MA1958.2 13 bp overlap
Motif DE_48h DE_48h-HOXD12ELK1_MA1958.2 13 bp overlap
Motif DE_60h DE_60h-HOXD12ELK1_MA1958.2 13 bp overlap
Hoxa11 3 datasets
Motif DE_36h DE_36h-Hoxa11_MA0911.2 9 bp overlap
Motif DE_48h DE_48h-Hoxa11_MA0911.2 9 bp overlap
Motif DE_60h DE_60h-Hoxa11_MA0911.2 9 bp overlap
JUN 2 datasets
ChIP DE_D1 S40-DE-d1-JUN-exp2 331 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.JUN.HUES-8_DE_ctrl 287 bp overlap
Jun 1 dataset
Motif DE_48h DE_48h-Jun_MA0489.3 8 bp overlap
KLF11 3 datasets
Motif DE_36h DE_36h-KLF11_MA1512.2 10 bp overlap
Motif DE_48h DE_48h-KLF11_MA1512.2 10 bp overlap
Motif DE_60h DE_60h-KLF11_MA1512.2 10 bp overlap
KLF16 3 datasets
Motif DE_36h DE_36h-KLF16_MA0741.1 11 bp overlap
Motif DE_48h DE_48h-KLF16_MA0741.1 11 bp overlap
Motif DE_60h DE_60h-KLF16_MA0741.1 11 bp overlap
KLF3 1 dataset
Motif DE_48h DE_48h-KLF3_MA1516.2 10 bp overlap
KLF4 2 datasets
ChIP BJ_INDUCED GSE36570.KLF4.BJ_INDUCED 105 bp overlap
ChIP keratinocyte_diff GSE57702.KLF4.keratinocyte_diff 133 bp overlap
KLF5 2 datasets
Motif DE_36h DE_36h-KLF5_MA0599.1 10 bp overlap
Motif DE_48h DE_48h-KLF5_MA0599.1 10 bp overlap
KLF9 3 datasets
Motif DE_36h DE_36h-KLF9_MA1107.3 11 bp overlap
Motif DE_48h DE_48h-KLF9_MA1107.3 11 bp overlap
Motif DE_60h DE_60h-KLF9_MA1107.3 11 bp overlap
LMO2 1 dataset
ChIP Kasumi-1_SIRUNX1ETO GSE60130.LMO2.Kasumi-1_SIRUNX1ETO 84 bp overlap
MAZ 2 datasets
Motif DE_36h DE_36h-MAZ_MA1522.2 8 bp overlap
Motif DE_48h DE_48h-MAZ_MA1522.2 8 bp overlap
MEIS1 2 datasets
Motif DE_36h DE_36h-MEIS1_MA0498.3 5 bp overlap
Motif DE_48h DE_48h-MEIS1_MA0498.3 5 bp overlap
MGA 2 datasets
Motif DE_36h DE_36h-MGA_MA0801.1 8 bp overlap
Motif DE_48h DE_48h-MGA_MA0801.1 8 bp overlap
MITF 1 dataset
ChIP melanocyte_BRAFV600E GSE50681.MITF.melanocyte_BRAFV600E 210 bp overlap
MYB 1 dataset
Motif DE_48h DE_48h-MYB_MA0100.4 6 bp overlap
NANOG 1 dataset
ChIP hESC GSE18292.NANOG.hESC 93 bp overlap
NFE2L1 1 dataset
ChIP WTC11 ENCFF644BPU 377 bp overlap
NR3C1 1 dataset
ChIP A-549 ENCSR000BJR.NR3C1.A-549 149 bp overlap
PATZ1 1 dataset
Motif DE_48h DE_48h-PATZ1_MA1961.2 11 bp overlap
PGR 2 datasets
ChIP MCF-7_R5020 GSE68355.PGR.MCF-7_R5020 94 bp overlap
ChIP T-47D_R5020-A1 GSE126859.PGR.T-47D_R5020-A1 75 bp overlap
POU5F1 1 dataset
ChIP hESC_NAIVE GSE69646.POU5F1.hESC_NAIVE 430 bp overlap
RBPJ 1 dataset
Motif DE_48h DE_48h-RBPJ_MA1116.2 6 bp overlap
RUNX1 2 datasets
ChIP AML_age-50-70 GSE111821.RUNX1.AML_age-50-70 219 bp overlap
ChIP epididymis_HEE GSE109061.RUNX1.epididymis_HEE 255 bp overlap
SMAD2 1 dataset
Motif DE_48h DE_48h-SMAD2_MA1964.2 6 bp overlap
SMAD2-3 3 datasets
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 336 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.SMAD2-3.HUES-8_DE_JNKi 647 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.SMAD2-3.HUES-8_DE_ctrl 444 bp overlap
SMAD2_3 5 datasets
ChIP DE_D1 S07-DE-d1-SMAD2_3-exp1 372 bp overlap
ChIP DE_D1 S12-DE-d1-SMAD2_3-exp1 610 bp overlap
ChIP DE_D1 S39-DE-d1-SMAD2_3-exp2 412 bp overlap
ChIP DE_D2 S01-DE-d2-SMAD2_3-exp1 498 bp overlap
ChIP DE_D2 S53-DE-d2-SMAD2_3-exp2 577 bp overlap
SMAD3 3 datasets
ChIP HCC1954_TGFb GSE104760.SMAD3.HCC1954_TGFb 308 bp overlap
ChIP HMLE_Doxycicline_TGFb GSE104760.SMAD3.HMLE_Doxycicline_TGFb 239 bp overlap
ChIP NCI-H441_SIITTF1 GSE51509.SMAD3.NCI-H441_SIITTF1 158 bp overlap
SMAD4 1 dataset
ChIP Caco-2 GSE112946.SMAD4.Caco-2 248 bp overlap
SMARCC1 1 dataset
ChIP DE_D1 S15-DE-d1-BAF155-exp1 270 bp overlap
SOX17_M 1 dataset
ChIP DE_D2 DED2-SOX17-M_Batch_II 760 bp overlap
SP3 1 dataset
Motif DE_48h DE_48h-SP3_MA0746.3 11 bp overlap
SP4 2 datasets
Motif DE_36h DE_36h-SP4_MA0685.2 9 bp overlap
Motif DE_48h DE_48h-SP4_MA0685.2 9 bp overlap
SP5 2 datasets
Motif DE_36h DE_36h-SP5_MA1965.2 6 bp overlap
Motif DE_48h DE_48h-SP5_MA1965.2 6 bp overlap
SP8 3 datasets
Motif DE_36h DE_36h-SP8_MA0747.2 11 bp overlap
Motif DE_48h DE_48h-SP8_MA0747.2 11 bp overlap
Motif DE_60h DE_60h-SP8_MA0747.2 11 bp overlap
SPI1 1 dataset
ChIP primary-monocyte_donorA_ncMO GSE128834.SPI1.primary-monocyte_donorA_ncMO 62 bp overlap
STAG1 1 dataset
ChIP MCF-7_E2_SHCTCF ERP000209.STAG1.MCF-7_E2_SHCTCF 54 bp overlap
STAG2 1 dataset
ChIP HMEC-1 GSE101921.STAG2.HMEC-1 142 bp overlap
STAT3 6 datasets
ChIP HCC1187 GSE152203.STAT3.HCC1187 149 bp overlap
ChIP HCC70 GSE152203.STAT3.HCC70 153 bp overlap
ChIP HCC70_EtOH GSE85579.STAT3.HCC70_EtOH 239 bp overlap
ChIP MCF-10A ENCSR000DOQ.STAT3.MCF-10A 167 bp overlap
ChIP MCF10A-Er-Src_TAM GSE115597.STAT3.MCF10A-Er-Src_TAM 177 bp overlap
ChIP T-47D_JC5056 GSE126004.STAT3.T-47D_JC5056 147 bp overlap
Six3 3 datasets
Motif DE_36h DE_36h-Six3_MA0631.2 11 bp overlap
Motif DE_48h DE_48h-Six3_MA0631.2 11 bp overlap
Motif DE_60h DE_60h-Six3_MA0631.2 11 bp overlap
Spi1 1 dataset
Motif DE_48h DE_48h-Spi1_MA0080.7 13 bp overlap
TBL1X 1 dataset
ChIP HEK293T GSE35197.TBL1X.HEK293T 191 bp overlap
TBR1 2 datasets
Motif DE_36h DE_36h-TBR1_MA0802.2 9 bp overlap
Motif DE_48h DE_48h-TBR1_MA0802.2 9 bp overlap
TBX1 2 datasets
Motif DE_36h DE_36h-TBX1_MA0805.1 8 bp overlap
Motif DE_48h DE_48h-TBX1_MA0805.1 8 bp overlap
TBX15 2 datasets
Motif DE_36h DE_36h-TBX15_MA0803.1 8 bp overlap
Motif DE_48h DE_48h-TBX15_MA0803.1 8 bp overlap
TBX18 3 datasets
Motif DE_36h DE_36h-TBX18_MA1565.2 9 bp overlap
Motif DE_48h DE_48h-TBX18_MA1565.2 9 bp overlap
Motif DE_60h DE_60h-TBX18_MA1565.2 9 bp overlap
TBX19 2 datasets
Motif DE_36h DE_36h-TBX19_MA0804.2 17 bp overlap
Motif DE_48h DE_48h-TBX19_MA0804.2 17 bp overlap
TBX20 1 dataset
Motif DE_48h DE_48h-TBX20_MA0689.1 11 bp overlap
TBX4 2 datasets
Motif DE_36h DE_36h-TBX4_MA0806.1 8 bp overlap
Motif DE_48h DE_48h-TBX4_MA0806.1 8 bp overlap
TFAP2B 1 dataset
ChIP SK-N-SH ENCFF869XXQ 101 bp overlap
TFAP2C 1 dataset
ChIP HCC1954 GSE126897.TFAP2C.HCC1954 50 bp overlap
TFCP2 1 dataset
Motif DE_48h DE_48h-TFCP2_MA1968.2 9 bp overlap
TRIM28 1 dataset
ChIP WIBR3_NAIVE GSE84382.TRIM28.WIBR3_NAIVE 380 bp overlap
TRPS1 2 datasets
Motif DE_36h DE_36h-TRPS1_MA1970.2 8 bp overlap
Motif DE_48h DE_48h-TRPS1_MA1970.2 8 bp overlap
TWIST1 1 dataset
Motif DE_48h DE_48h-TWIST1_MA1123.3 8 bp overlap
USF1 1 dataset
ChIP WA01 ENCSR000BIU.USF1.WA01 392 bp overlap
ZBTB26 2 datasets
Motif DE_36h DE_36h-ZBTB26_MA1579.2 8 bp overlap
Motif DE_48h DE_48h-ZBTB26_MA1579.2 8 bp overlap
ZEB1 1 dataset
Motif DE_48h DE_48h-ZEB1_MA0103.4 6 bp overlap
ZNF148 2 datasets
Motif DE_36h DE_36h-ZNF148_MA1653.2 10 bp overlap
Motif DE_48h DE_48h-ZNF148_MA1653.2 10 bp overlap
ZNF213 1 dataset
Motif DE_48h DE_48h-ZNF213_MA2121.1 12 bp overlap
ZNF257 1 dataset
Motif DE_48h DE_48h-ZNF257_MA1710.2 10 bp overlap
ZNF281 1 dataset
Motif DE_48h DE_48h-ZNF281_MA1630.3 10 bp overlap
ZNF416 1 dataset
Motif DE_48h DE_48h-ZNF416_MA1979.2 10 bp overlap
ZNF534 1 dataset
ChIP HEK293T GSE78099.ZNF534.HEK293T 88 bp overlap
ZNF582 2 datasets
Motif DE_36h DE_36h-ZNF582_MA1983.2 19 bp overlap
Motif DE_48h DE_48h-ZNF582_MA1983.2 19 bp overlap
Zfp961 1 dataset
Motif DE_48h DE_48h-Zfp961_MA2126.1 8 bp overlap
Zic1::Zic2 3 datasets
Motif DE_36h DE_36h-Zic1Zic2_MA1628.2 7 bp overlap
Motif DE_48h DE_48h-Zic1Zic2_MA1628.2 7 bp overlap
Motif DE_60h DE_60h-Zic1Zic2_MA1628.2 7 bp overlap
Zic3 3 datasets
Motif DE_36h DE_36h-Zic3_MA0697.3 7 bp overlap
Motif DE_48h DE_48h-Zic3_MA0697.3 7 bp overlap
Motif DE_60h DE_60h-Zic3_MA0697.3 7 bp overlap