chr4 : 188,109,275 188,109,719
444 bp 75 TFs 1 linked gene
This 444 bp open chromatin element is linked to TRIML2 and is bound by 75 transcription factors.
Linked Genes
1 gene
Gene Expression Dist. to TSS Distance Link type
TRIML2 at TSS At TSS Proximity
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr4:188,104,275 – 188,114,719
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
75 transcription factors
Source
Cell type
AHR 1 dataset
ChIP MCF-7_TCDD_45min GSE90550.AHR.MCF-7_TCDD_45min 134 bp overlap
AR 3 datasets
ChIP LNCaP_ETOH GSE69043.AR.LNCaP_ETOH 119 bp overlap
ChIP LNCaP_R1881 GSE69043.AR.LNCaP_R1881 141 bp overlap
ChIP LNCaP_SHGATA2_R1881 GSE69043.AR.LNCaP_SHGATA2_R1881 173 bp overlap
ARID1A 3 datasets
ChIP 12Z GSE129781.ARID1A.12Z 175 bp overlap
ChIP MCF-7_4-hydroxytamoxifen GSE123284.ARID1A.MCF-7_4-hydroxytamoxifen 330 bp overlap
ChIP MCF-7_Fulvestrant GSE123284.ARID1A.MCF-7_Fulvestrant 118 bp overlap
ASH2L 1 dataset
ChIP WA01 ENCSR850KIP.ASH2L.WA01 206 bp overlap
ATF3 1 dataset
ChIP HCT-116_CAMP GSE74355.ATF3.HCT-116_CAMP 102 bp overlap
BICRA 1 dataset
ChIP Mel270_K700E GSE124720.BICRA.Mel270_K700E 153 bp overlap
BRD2 2 datasets
ChIP MDA-MB-231_MGSK2801 GSE116879.BRD2.MDA-MB-231_MGSK2801 174 bp overlap
ChIP MDA-MB-231_MGSK2801_rDNA GSE116879.BRD2.MDA-MB-231_MGSK2801_rDNA 174 bp overlap
BRD4 17 datasets
ChIP HAP1 GSE108387.BRD4.HAP1 194 bp overlap
ChIP HCC1806_100nMtrametinib_24h GSE87418.BRD4.HCC1806_100nMtrametinib_24h 358 bp overlap
ChIP HCC1937 GSE124748.BRD4.HCC1937 126 bp overlap
ChIP MDA-MB-231_DMSO GSE116879.BRD4.MDA-MB-231_DMSO 436 bp overlap
ChIP MDA-MB-231_DMSO_rDNA GSE116879.BRD4.MDA-MB-231_DMSO_rDNA 436 bp overlap
ChIP MDA-MB-231_MGSK2801 GSE116879.BRD4.MDA-MB-231_MGSK2801 444 bp overlap
ChIP MDA-MB-231_MGSK2801_rDNA GSE116879.BRD4.MDA-MB-231_MGSK2801_rDNA 444 bp overlap
ChIP SUM159PT_100nMtrametinib_4h GSE87418.BRD4.SUM159PT_100nMtrametinib_4h 297 bp overlap
ChIP SUM159PT_30nMbortezomib_8h GSE87418.BRD4.SUM159PT_30nMbortezomib_8h 214 bp overlap
ChIP SUM159PT_DMSO_48h GSE87418.BRD4.SUM159PT_DMSO_48h 362 bp overlap
ChIP SUM159PT_DMSO_8h GSE87418.BRD4.SUM159PT_DMSO_8h 353 bp overlap
ChIP SUM159PT_Dox_48h GSE87418.BRD4.SUM159PT_Dox_48h 444 bp overlap
ChIP SUM159PT_scramble GSE131097.BRD4.SUM159PT_scramble 444 bp overlap
ChIP SUM159PT_shMYC_DMSO_48h GSE87418.BRD4.SUM159PT_shMYC_DMSO_48h 359 bp overlap
ChIP SUM159PT_shMYC_dox_48h GSE87418.BRD4.SUM159PT_shMYC_dox_48h 123 bp overlap
ChIP SUM159PT_shMYC_dox_48h GSE87418.BRD4.SUM159PT_shMYC_dox_48h 234 bp overlap
ChIP keratinocyte GSE140992.BRD4.keratinocyte 364 bp overlap
BRD9 3 datasets
ChIP MDA-MB-231_JQ1_MGSK2801 GSE116879.BRD9.MDA-MB-231_JQ1_MGSK2801 340 bp overlap
ChIP MDA-MB-231_MGSK2801 GSE116879.BRD9.MDA-MB-231_MGSK2801 56 bp overlap
ChIP MDA-MB-231_MGSK2801 GSE116879.BRD9.MDA-MB-231_MGSK2801 188 bp overlap
CBX3 2 datasets
ChIP HCT-116 ENCSR000BUH.CBX3.HCT-116 151 bp overlap
ChIP HCT116 ENCFF947BOL 364 bp overlap
CEBPB 7 datasets
ChIP A-549 ENCSR000BUB.CEBPB.A-549 119 bp overlap
ChIP A549 ENCFF235AIY 200 bp overlap
ChIP A549 ENCFF781RLJ 226 bp overlap
ChIP A549 ENCFF797MXZ 262 bp overlap
ChIP H1 ENCFF871PTR 205 bp overlap
ChIP HCT116 ENCFF097OLY 302 bp overlap
ChIP WA01 ENCSR000EBV.CEBPB.WA01 79 bp overlap
CHD1 1 dataset
ChIP H1 ENCFF998XEK 106 bp overlap
CHD4 1 dataset
ChIP HaCaT GSE139685.CHD4.HaCaT 172 bp overlap
CREBBP 1 dataset
ChIP MCF-7 ERP000901.CREBBP.MCF-7 164 bp overlap
Cebpa 1 dataset
ChIP BLaER1 ENCFF460KDD 233 bp overlap
ELL2 1 dataset
ChIP HeLa_DOX_EGF GSE40632.ELL2.HeLa_DOX_EGF 91 bp overlap
EP300 5 datasets
ChIP A-549 ENCSR000BPW.EP300.A-549 77 bp overlap
ChIP A-549 ENCSR000BPW.EP300.A-549 263 bp overlap
ChIP HeLa-S3 ENCFF245KNK 279 bp overlap
ChIP HeLa-S3 ENCSR000ECV.EP300.HeLa-S3 221 bp overlap
ChIP HeLa-S3 ENCSR680OFU.EP300.HeLa-S3 216 bp overlap
ESR1 1 dataset
ChIP MCF-7_RAD001 GSE103023.ESR1.MCF-7_RAD001 149 bp overlap
FOS 4 datasets
ChIP HeLa-S3 ENCSR000EZE.FOS.HeLa-S3 205 bp overlap
ChIP MCF-10A ENCSR000DON.FOS.MCF-10A 137 bp overlap
ChIP MCF-7 ENCFF282FWZ 183 bp overlap
ChIP MCF-7 ENCSR569XNP.FOS.MCF-7 348 bp overlap
FOSL1 4 datasets
ChIP BT-549 GSE46166.FOSL1.BT-549 292 bp overlap
ChIP BT-549 GSE112961.FOSL1.BT-549 217 bp overlap
ChIP HCT-116 ENCSR000BTE.FOSL1.HCT-116 224 bp overlap
ChIP HCT116 ENCFF540ZXN 369 bp overlap
FOSL2 6 datasets
ChIP A-549 ENCSR448TVS.FOSL2.A-549 423 bp overlap
ChIP A-549 ENCSR000BQO.FOSL2.A-549 255 bp overlap
ChIP A549 ENCFF195CES 210 bp overlap
ChIP A549 ENCFF651PDH 352 bp overlap
ChIP MCF-7 ENCFF716UWP 291 bp overlap
ChIP MCF-7 ENCSR000BUI.FOSL2.MCF-7 156 bp overlap
GATA2 6 datasets
ChIP ESF GSE108408.GATA2.ESF 183 bp overlap
ChIP LNCaP_CSFCS GSE69043.GATA2.LNCaP_CSFCS 221 bp overlap
ChIP LNCaP_FBS GSE69043.GATA2.LNCaP_FBS 175 bp overlap
ChIP WA09 GSE105081.GATA2.WA09 149 bp overlap
ChIP primary-endometrial-stromal-cell_IVD_P2 GSE145673.GATA2.primary-endometrial-stromal-cell_IVD_P2 168 bp overlap
ChIP primary-endometrial-stromal-cell_Veh_P1 GSE145673.GATA2.primary-endometrial-stromal-cell_Veh_P1 212 bp overlap
GATA3 7 datasets
ChIP A-549 ENCSR000BTI.GATA3.A-549 253 bp overlap
ChIP A549 ENCFF226FVV 236 bp overlap
ChIP MCF-7 ENCFF352QVM 186 bp overlap
ChIP MCF-7 GSE128445.GATA3.MCF-7 241 bp overlap
ChIP MCF-7 GSE117492.GATA3.MCF-7 172 bp overlap
ChIP MCF-7 GSE122847.GATA3.MCF-7 157 bp overlap
ChIP MCF-7 GSE51274.GATA3.MCF-7 156 bp overlap
GATA3_Nter 2 datasets
ChIP T-47D_CR3flp GSE99479.GATA3_Nter.T-47D_CR3flp 183 bp overlap
ChIP T-47D_flp-ctrl GSE99479.GATA3_Nter.T-47D_flp-ctrl 158 bp overlap
GATA4 1 dataset
ChIP DE DE-GATA4-2 239 bp overlap
GATA6 8 datasets
ChIP DE DE-GATA6-1 116 bp overlap
ChIP DE DE-GATA6-2 222 bp overlap
ChIP DE_D1 S09-DE-d1-GATA6-exp1 331 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 365 bp overlap
ChIP DE_D1 S41-DE-d1-GATA6-exp2 221 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 444 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 344 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 264 bp overlap
GATAD1 1 dataset
ChIP HeLa GSE20303.GATAD1.HeLa 219 bp overlap
GTF3C2 2 datasets
ChIP HeLa-S3 ENCFF144RTV 321 bp overlap
ChIP HeLa-S3 ENCSR000DNY.GTF3C2.HeLa-S3 183 bp overlap
HSF1 8 datasets
ChIP BPE_HEAT GSE38901.HSF1.BPE_HEAT 197 bp overlap
ChIP HCT-116_A10_43 GSE152144.HSF1.HCT-116_A10_43 311 bp overlap
ChIP HCT-116_A8_43 GSE152144.HSF1.HCT-116_A8_43 305 bp overlap
ChIP HCT-116_A9_43 GSE152144.HSF1.HCT-116_A9_43 202 bp overlap
ChIP MCF-10A_HEAT GSE38901.HSF1.MCF-10A_HEAT 144 bp overlap
ChIP U2OS_HEAT GSE60984.HSF1.U2OS_HEAT 198 bp overlap
ChIP U2OS_HEAT_20 GSE60984.HSF1.U2OS_HEAT_20 179 bp overlap
ChIP hTERT-HME1_HEAT GSE38901.HSF1.hTERT-HME1_HEAT 200 bp overlap
JUN 7 datasets
ChIP BT-549 GSE46166.JUN.BT-549 257 bp overlap
ChIP DE_D1 S08-DE-d1-JUN-exp1 178 bp overlap
ChIP DE_D2 S02-DE-d2-JUN-exp1 426 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.JUN.HUES-8_DE_ctrl 268 bp overlap
ChIP HeLa-S3 ENCFF668QVP 335 bp overlap
ChIP MCF-7_E2 GSE102410.JUN.MCF-7_E2 180 bp overlap
ChIP MDA-BoM-1833 GSE112444.JUN.MDA-BoM-1833 233 bp overlap
JUNB 1 dataset
ChIP keratinocyte_CTR GSE139685.JUNB.keratinocyte_CTR 159 bp overlap
JUND 7 datasets
ChIP A-549 ENCSR000BRF.JUND.A-549 108 bp overlap
ChIP H1 ENCFF010YXS 304 bp overlap
ChIP HCT-116 ENCSR000BSA.JUND.HCT-116 197 bp overlap
ChIP HCT116 ENCFF748ZQX 352 bp overlap
ChIP HeLa-S3 ENCFF642OHL 321 bp overlap
ChIP HeLa-S3 ENCSR000EDH.JUND.HeLa-S3 235 bp overlap
ChIP WA01 ENCSR000EBZ.JUND.WA01 160 bp overlap
KDM5B 2 datasets
ChIP SUM185 GSE46055.KDM5B.SUM185 80 bp overlap
ChIP SUM185_SHCTCF GSE46055.KDM5B.SUM185_SHCTCF 119 bp overlap
KLF3 1 dataset
ChIP keratinocyte GSE140991.KLF3.keratinocyte 188 bp overlap
MAFF 1 dataset
ChIP HeLa-S3 ENCSR140DSL.MAFF.HeLa-S3 120 bp overlap
MAML1 1 dataset
ChIP SCC_4h GSE156486.MAML1.SCC_4h 303 bp overlap
MAX 1 dataset
ChIP A-549 ENCSR000BTJ.MAX.A-549 154 bp overlap
MED26 2 datasets
ChIP HCT-116 GSE121355.MED26.HCT-116 168 bp overlap
ChIP HCT-116_aux GSE121355.MED26.HCT-116_aux 444 bp overlap
MORC2 1 dataset
ChIP H9 GSE95374.MORC2.H9 183 bp overlap
MYC 1 dataset
ChIP HeLa-S3 ENCSR000EZD.MYC.HeLa-S3 208 bp overlap
MYCN 1 dataset
ChIP SH-EP_pRRL-MYCN GSE111905.MYCN.SH-EP_pRRL-MYCN 79 bp overlap
NANOG 2 datasets
ChIP WA01_3IL ERP004238.NANOG.WA01_3IL 327 bp overlap
ChIP WA09_heat-shock GSE105028.NANOG.WA09_heat-shock 165 bp overlap
NELFE 1 dataset
ChIP HeLa GSE125534.NELFE.HeLa 130 bp overlap
NIPBL 3 datasets
ChIP HCT-116_RAD21-mAC GSE104888.NIPBL.HCT-116_RAD21-mAC 209 bp overlap
ChIP HCT-116_RAD21-mAC_500uM_auxin GSE104888.NIPBL.HCT-116_RAD21-mAC_500uM_auxin 169 bp overlap
ChIP HCT-116_RAD21-mAC_500uM_auxin GSE104888.NIPBL.HCT-116_RAD21-mAC_500uM_auxin 226 bp overlap
NR3C1 3 datasets
ChIP A-549 ENCSR000BHF.NR3C1.A-549 189 bp overlap
ChIP BEAS-2B_TNF_IA1 GSE125623.NR3C1.BEAS-2B_TNF_IA1 246 bp overlap
ChIP HCC1937 GSE152203.NR3C1.HCC1937 158 bp overlap
NR5A2 2 datasets
ChIP A-549 ENCSR190GIW.NR5A2.A-549 244 bp overlap
ChIP A549 ENCFF834RVE 130 bp overlap
NRF1 1 dataset
ChIP HeLa-S3 GSE108856.NRF1.HeLa-S3 226 bp overlap
OTX2 1 dataset
ChIP WTC11 ENCFF634NAO 245 bp overlap
PAF1 1 dataset
ChIP HCT-116_ab20662 GSE97527.PAF1.HCT-116_ab20662 215 bp overlap
PHIP 3 datasets
ChIP HCT-116_MLL1-KO__ab833 GSE101646.PHIP.HCT-116_MLL1-KO__ab833 165 bp overlap
ChIP HCT-116_MLL4-SET-KO_ab834 GSE101646.PHIP.HCT-116_MLL4-SET-KO_ab834 117 bp overlap
ChIP HCT-116_ab834 GSE101646.PHIP.HCT-116_ab834 114 bp overlap
POLR2A 2 datasets
ChIP H1 ENCFF566JSR 444 bp overlap
ChIP H1 ENCFF833NJP 314 bp overlap
RAD21 1 dataset
ChIP A-549 ENCSR000BUC.RAD21.A-549 114 bp overlap
RBPJ 1 dataset
ChIP SCC_4h GSE156486.RBPJ.SCC_4h 289 bp overlap
SIN3A 2 datasets
ChIP HCT116 ENCFF203YBB 444 bp overlap
ChIP HCT116 ENCFF203YBB 112 bp overlap
SMAD2_3 2 datasets
ChIP DE_D2 S01-DE-d2-SMAD2_3-exp1 332 bp overlap
ChIP DE_D2 S53-DE-d2-SMAD2_3-exp2 312 bp overlap
SMAD3 1 dataset
ChIP HCC1954_TGFb GSE104760.SMAD3.HCC1954_TGFb 243 bp overlap
SMARCA4 10 datasets
ChIP A-549_AG15677 GSE132290.SMARCA4.A-549_AG15677 61 bp overlap
ChIP A-549_AG15720 GSE132290.SMARCA4.A-549_AG15720 81 bp overlap
ChIP A-549_AG15721 GSE132290.SMARCA4.A-549_AG15721 63 bp overlap
ChIP A-549_AG15723 GSE132290.SMARCA4.A-549_AG15723 120 bp overlap
ChIP A-549_AG15724 GSE132290.SMARCA4.A-549_AG15724 66 bp overlap
ChIP BIN-67_lentivirus-SMARCA4 GSE117734.SMARCA4.BIN-67_lentivirus-SMARCA4 215 bp overlap
ChIP TTC-1240 GSE124903.SMARCA4.TTC-1240 198 bp overlap
ChIP TTC-1240_delC GSE124903.SMARCA4.TTC-1240_delC 173 bp overlap
ChIP hiPSC GSE124903.SMARCA4.hiPSC 202 bp overlap
ChIP hiPSC_WT GSE124903.SMARCA4.hiPSC_WT 259 bp overlap
SMARCB1 3 datasets
ChIP MCF-7_estrogen GSE123284.SMARCB1.MCF-7_estrogen 61 bp overlap
ChIP hiPSC GSE124903.SMARCB1.hiPSC 172 bp overlap
ChIP hiPSC_WT GSE124903.SMARCB1.hiPSC_WT 153 bp overlap
SMARCC1 3 datasets
ChIP DE_D1 S15-DE-d1-BAF155-exp1 391 bp overlap
ChIP hiPSC GSE124903.SMARCC1.hiPSC 274 bp overlap
ChIP hiPSC_WT GSE124903.SMARCC1.hiPSC_WT 384 bp overlap
SMC1 1 dataset
ChIP HCT-116 GSE131606.SMC1.HCT-116 194 bp overlap
SMC3 1 dataset
ChIP HeLa-Kyoto_PDS5-depleted GSE102884.SMC3.HeLa-Kyoto_PDS5-depleted 134 bp overlap
SP1 3 datasets
ChIP A-549 ENCSR000BPE.SP1.A-549 244 bp overlap
ChIP HCT-116 ENCSR000BSF.SP1.HCT-116 155 bp overlap
ChIP HCT116 ENCFF800LBN 334 bp overlap
SPDEF 1 dataset
ChIP MCF-7 ENCSR042GSX.SPDEF.MCF-7 179 bp overlap
SS18 1 dataset
ChIP BIN-67_lentivirus-SMARCA4 GSE117734.SS18.BIN-67_lentivirus-SMARCA4 207 bp overlap
STAT3 4 datasets
ChIP MCF-7_jc5836 GSE126004.STAT3.MCF-7_jc5836 173 bp overlap
ChIP T-47D_JC5054 GSE126004.STAT3.T-47D_JC5054 321 bp overlap
ChIP T-47D_JC5056 GSE126004.STAT3.T-47D_JC5056 260 bp overlap
ChIP T-47D_JC5062 GSE126004.STAT3.T-47D_JC5062 192 bp overlap
SUPT5H 2 datasets
ChIP HCT-116_pThr806 GSE138548.SUPT5H.HCT-116_pThr806 274 bp overlap
ChIP HeLa_DOX GSE125534.SUPT5H.HeLa_DOX 310 bp overlap
TBP 5 datasets
ChIP HeLa-S3 ENCSR000EDD.TBP.HeLa-S3 174 bp overlap
ChIP hESC GSE122298.TBP.hESC 154 bp overlap
ChIP hESC_10h GSE122298.TBP.hESC_10h 204 bp overlap
ChIP hESC_2h GSE122298.TBP.hESC_2h 230 bp overlap
ChIP hESC_8h GSE122298.TBP.hESC_8h 153 bp overlap
TCF12 1 dataset
ChIP A-549 ENCSR000BQQ.TCF12.A-549 231 bp overlap
TCF7L2 3 datasets
ChIP HCT-116 ENCSR000EUV.TCF7L2.HCT-116 221 bp overlap
ChIP HeLa-S3 ENCSR000EVE.TCF7L2.HeLa-S3 214 bp overlap
ChIP LNCaP GSE51621.TCF7L2.LNCaP 244 bp overlap
TEAD1 4 datasets
ChIP HUCCT1 GSE68296.TEAD1.HUCCT1 121 bp overlap
ChIP HUCCT1 GSE68296.TEAD1.HUCCT1 261 bp overlap
ChIP MSTO GSE68170.TEAD1.MSTO 338 bp overlap
ChIP keratinocyte GSE138727.TEAD1.keratinocyte 438 bp overlap
TEAD4 6 datasets
ChIP A-549 ENCSR000BUD.TEAD4.A-549 133 bp overlap
ChIP BJ_fibroblast_CD13-neg GSE114367.TEAD4.BJ_fibroblast_CD13-neg 211 bp overlap
ChIP ESC S26-ESC-d0-TEAD4-exp1 288 bp overlap
ChIP H1 ENCFF778PAX 206 bp overlap
ChIP HCT116 ENCFF526YYD 201 bp overlap
ChIP SNU-216 GSE44416.TEAD4.SNU-216 426 bp overlap
YY1 2 datasets
ChIP HCT-116 ENCSR000BNX.YY1.HCT-116 172 bp overlap
ChIP Hep-G2_RBM25-KD GSE120104.YY1.Hep-G2_RBM25-KD 285 bp overlap
YY1AP1 1 dataset
ChIP MCF-10A_DOX GSE115787.YY1AP1.MCF-10A_DOX 318 bp overlap
ZHX1 1 dataset
ChIP HeLa-S3 ENCSR855WKQ.ZHX1.HeLa-S3 159 bp overlap