chr8 : 116,352,118 116,352,437
319 bp 83 TFs 0 linked genes
This 319 bp open chromatin element has no linked target genes and is bound by 83 transcription factors.
Linked Genes
No linked genes
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr8:116,347,118 – 116,357,437
This element Other elements ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
83 transcription factors
Source
Cell type
AR 2 datasets
ChIP MCF-7 GSE48930.AR.MCF-7 106 bp overlap
ChIP THP-1_R1881 GSE131381.AR.THP-1_R1881 264 bp overlap
ARID1A 3 datasets
ChIP MCF-7 GSE123284.ARID1A.MCF-7 145 bp overlap
ChIP MCF-7_Fulvestrant GSE123284.ARID1A.MCF-7_Fulvestrant 319 bp overlap
ChIP MCF-7_JQ1 GSE123284.ARID1A.MCF-7_JQ1 208 bp overlap
ARNTL 1 dataset
ChIP U2OS_trough_DMSO GSE85096.ARNTL.U2OS_trough_DMSO 178 bp overlap
Atoh1 1 dataset
Motif DE_12h DE_12h-Atoh1_MA0461.3 8 bp overlap
BRD2 1 dataset
ChIP MDA-MB-231_MGSK2801 GSE116879.BRD2.MDA-MB-231_MGSK2801 153 bp overlap
BRD3 1 dataset
ChIP HEK293T GSE39579.BRD3.HEK293T 67 bp overlap
BRD4 10 datasets
ChIP CD4_Th1_DMSO GSE62482.BRD4.CD4_Th1_DMSO 287 bp overlap
ChIP HCC1806_100nMtrametinib_24h GSE87418.BRD4.HCC1806_100nMtrametinib_24h 88 bp overlap
ChIP HFOB_DIFF GSE82295.BRD4.HFOB_DIFF 175 bp overlap
ChIP IMR-90_QUIES GSE74238.BRD4.IMR-90_QUIES 138 bp overlap
ChIP IMR-90_SENES GSE74238.BRD4.IMR-90_SENES 171 bp overlap
ChIP MCF-7_parental_4-hydroxytamoxifen GSE123284.BRD4.MCF-7_parental_4-hydroxytamoxifen 319 bp overlap
ChIP SGBS_TNF GSE64233.BRD4.SGBS_TNF 235 bp overlap
ChIP SUM159PT_DMSO_48h GSE87418.BRD4.SUM159PT_DMSO_48h 162 bp overlap
ChIP SUM159PT_Dox_48h GSE87418.BRD4.SUM159PT_Dox_48h 96 bp overlap
ChIP SUM159PT_scramble GSE131097.BRD4.SUM159PT_scramble 319 bp overlap
CBX1 1 dataset
ChIP Hep-G2 ENCSR819WZE.CBX1.Hep-G2 170 bp overlap
CDK8 3 datasets
ChIP leiomyoma_PT886 GSE128230.CDK8.leiomyoma_PT886 61 bp overlap
ChIP monocyte_IFNg GSE120943.CDK8.monocyte_IFNg 181 bp overlap
ChIP monocyte_IFNg-LPS GSE120943.CDK8.monocyte_IFNg-LPS 185 bp overlap
CEBPA 6 datasets
ChIP Kasumi-1_CEBPA-ER_E2 GSE102697.CEBPA.Kasumi-1_CEBPA-ER_E2 245 bp overlap
ChIP SGBS GSE41629.CEBPA.SGBS 209 bp overlap
ChIP SKH1_CEBPA-ER_E2 GSE102697.CEBPA.SKH1_CEBPA-ER_E2 163 bp overlap
ChIP THP-1_1-25D_24h GSE124032.CEBPA.THP-1_1-25D_24h 151 bp overlap
ChIP THP-1_1-25D_8h GSE124032.CEBPA.THP-1_1-25D_8h 92 bp overlap
ChIP THP-1_EtOH_24h GSE124032.CEBPA.THP-1_EtOH_24h 136 bp overlap
CEBPB 7 datasets
ChIP IMR-90 ENCFF468UGY 251 bp overlap
ChIP THP-1_NS1-Pam3csk-4h GSE103477.CEBPB.THP-1_NS1-Pam3csk-4h 236 bp overlap
ChIP THP-1_eGFP-Pam3csk-4h GSE103477.CEBPB.THP-1_eGFP-Pam3csk-4h 319 bp overlap
ChIP monocyte GSE98367.CEBPB.monocyte 198 bp overlap
ChIP monocyte_IFNg-LPS GSE120943.CEBPB.monocyte_IFNg-LPS 319 bp overlap
ChIP monocyte_INFg GSE98367.CEBPB.monocyte_INFg 319 bp overlap
ChIP monocyte_MACROPHAGE GSE31621.CEBPB.monocyte_MACROPHAGE 169 bp overlap
CRY1 2 datasets
ChIP U2OS GSE130602.CRY1.U2OS 254 bp overlap
ChIP U2OS_DMSO GSE130506.CRY1.U2OS_DMSO 254 bp overlap
CTCF 1 dataset
ChIP THP-1_macrophage_PMA GSE96800.CTCF.THP-1_macrophage_PMA 235 bp overlap
Cebpa 10 datasets
ChIP BLaER1 ENCFF205PMS 319 bp overlap
ChIP BLaER1 ENCFF262VBH 319 bp overlap
ChIP BLaER1 ENCFF374ODN 319 bp overlap
ChIP BLaER1 ENCFF374ODN 230 bp overlap
ChIP BLaER1 ENCFF419EBE 319 bp overlap
ChIP BLaER1 ENCFF460KDD 255 bp overlap
ChIP BLaER1 ENCFF538NOH 319 bp overlap
ChIP BLaER1 ENCFF538NOH 181 bp overlap
ChIP BLaER1 ENCFF680YXW 319 bp overlap
ChIP BLaER1 ENCFF896HSY 319 bp overlap
DPF2 1 dataset
ChIP SCCOHT-1 GSE117734.DPF2.SCCOHT-1 244 bp overlap
EGR1 2 datasets
ChIP macrophage_D3 GSE136216.EGR1.macrophage_D3 277 bp overlap
ChIP macrophage_D4 GSE136216.EGR1.macrophage_D4 235 bp overlap
ELF1 1 dataset
ChIP SK-N-MC_SHFLI_96H GSE61944.ELF1.SK-N-MC_SHFLI_96H 231 bp overlap
ELF3 2 datasets
ChIP PDAC GSE64557.ELF3.PDAC 319 bp overlap
ChIP PDAC_SHCTR GSE64557.ELF3.PDAC_SHCTR 319 bp overlap
EP300 1 dataset
ChIP tibial nerve ENCFF346AYA 319 bp overlap
ESR1 9 datasets
ChIP Ishikawa_E2_D538G_clone2 GSE132426.ESR1.Ishikawa_E2_D538G_clone2 190 bp overlap
ChIP Ishikawa_ETV4-KO2_Mili GSE129803.ESR1.Ishikawa_ETV4-KO2_Mili 201 bp overlap
ChIP MCF-7 GSE76893.ESR1.MCF-7 90 bp overlap
ChIP MCF-7 GSE128445.ESR1.MCF-7 72 bp overlap
ChIP MCF-7_G6274 GSE117941.ESR1.MCF-7_G6274 318 bp overlap
ChIP MCF-7_TamR GSE128445.ESR1.MCF-7_TamR 167 bp overlap
ChIP ZR751 ERP000783.ESR1.ZR751 158 bp overlap
ChIP breast_tumor_Male_10 GSE104399.ESR1.breast_tumor_Male_10 299 bp overlap
ChIP breast_tumor_Male_14 GSE104399.ESR1.breast_tumor_Male_14 209 bp overlap
ESRRA 1 dataset
ChIP BT-474 GSE81651.ESRRA.BT-474 55 bp overlap
ETV1 1 dataset
ChIP GIST GSE22441.ETV1.GIST 121 bp overlap
FLI1 1 dataset
ChIP A-673_1_KRAB_GFP GSE106914.FLI1.A-673_1_KRAB_GFP 130 bp overlap
FOS 2 datasets
ChIP IMR-90 ENCFF179EDA 203 bp overlap
ChIP IMR-90 ENCSR124AIG.FOS.IMR-90 144 bp overlap
FOXA1 27 datasets
ChIP CFPAC-1_FOXA1-KO GSE119930.FOXA1.CFPAC-1_FOXA1-KO 180 bp overlap
ChIP MCF-7 GSE72249.FOXA1.MCF-7 102 bp overlap
ChIP MCF-7 GSE128445.FOXA1.MCF-7 190 bp overlap
ChIP MCF-7 GSE59530.FOXA1.MCF-7 72 bp overlap
ChIP MCF-7 ERP001226.FOXA1.MCF-7 100 bp overlap
ChIP MCF-7_DEX GSE72249.FOXA1.MCF-7_DEX 262 bp overlap
ChIP MCF-7_DOX-neg GSE124654.FOXA1.MCF-7_DOX-neg 143 bp overlap
ChIP MCF-7_E2 GSE72249.FOXA1.MCF-7_E2 298 bp overlap
ChIP MCF-7_E2_TAM ERP000380.FOXA1.MCF-7_E2_TAM 90 bp overlap
ChIP MCF-7_TAMR_E2_TAM ERP000380.FOXA1.MCF-7_TAMR_E2_TAM 103 bp overlap
ChIP MCF-7_TNF GSE59530.FOXA1.MCF-7_TNF 90 bp overlap
ChIP MCF-7_TamR GSE128445.FOXA1.MCF-7_TamR 143 bp overlap
ChIP MCF-7_estrogen_ab2 GSE112969.FOXA1.MCF-7_estrogen_ab2 82 bp overlap
ChIP MCF-7_vehicle_ab2 GSE112969.FOXA1.MCF-7_vehicle_ab2 165 bp overlap
ChIP NCI-H3122 GSE39998.FOXA1.NCI-H3122 175 bp overlap
ChIP PDAC GSE64557.FOXA1.PDAC 319 bp overlap
ChIP ZR-75-1_estrogen_ab1 GSE112969.FOXA1.ZR-75-1_estrogen_ab1 131 bp overlap
ChIP ZR-75-1_estrogen_ab2 GSE112969.FOXA1.ZR-75-1_estrogen_ab2 319 bp overlap
ChIP ZR-75-1_vehicle_ab1 GSE112969.FOXA1.ZR-75-1_vehicle_ab1 319 bp overlap
ChIP ZR-75-1_vehicle_ab2 GSE112969.FOXA1.ZR-75-1_vehicle_ab2 319 bp overlap
ChIP ZR751 GSE72249.FOXA1.ZR751 228 bp overlap
ChIP ZR751_DEX GSE72249.FOXA1.ZR751_DEX 319 bp overlap
ChIP ZR751_E2 GSE72249.FOXA1.ZR751_E2 209 bp overlap
ChIP breast-cancer_Veh-2856 GSE128018.FOXA1.breast-cancer_Veh-2856 72 bp overlap
ChIP breast_tumor_Female_1 GSE104399.FOXA1.breast_tumor_Female_1 67 bp overlap
ChIP primary-breast-cancer_B1_DSG GSE114737.FOXA1.primary-breast-cancer_B1_DSG 65 bp overlap
ChIP primary-breast-cancer_B1_DSG GSE114737.FOXA1.primary-breast-cancer_B1_DSG 181 bp overlap
FOXA2 3 datasets
ChIP CFPAC-1_EV GSE119930.FOXA2.CFPAC-1_EV 104 bp overlap
ChIP CFPAC-1_HOXB8-OE GSE119930.FOXA2.CFPAC-1_HOXB8-OE 244 bp overlap
ChIP DE DE-FOXA2-1 121 bp overlap
FOXF1 3 datasets
ChIP GIST-T1 GSE106624.FOXF1.GIST-T1 110 bp overlap
ChIP GIST48 GSE106624.FOXF1.GIST48 131 bp overlap
ChIP GIST48_siETV1 GSE106624.FOXF1.GIST48_siETV1 79 bp overlap
FOXS1 1 dataset
Motif DE_12h DE_12h-FOXS1_MA2118.1 8 bp overlap
GATA2 1 dataset
ChIP dermal-fibroblast_alone GSE51025.GATA2.dermal-fibroblast_alone 209 bp overlap
GATA3 1 dataset
ChIP MCF-7 GSE51274.GATA3.MCF-7 192 bp overlap
HIF1A 2 datasets
ChIP U2OS_DMSO GSE85096.HIF1A.U2OS_DMSO 176 bp overlap
ChIP U2OS_trough_DMOG GSE85096.HIF1A.U2OS_trough_DMOG 212 bp overlap
IRF1 1 dataset
ChIP CD14_LPS GSE43036.IRF1.CD14_LPS 171 bp overlap
JUN 2 datasets
ChIP MCF-7_DMSO GSE157385.JUN.MCF-7_DMSO 217 bp overlap
ChIP MCF-7_TamR_BD610326 GSE128445.JUN.MCF-7_TamR_BD610326 124 bp overlap
JUNB 1 dataset
ChIP CFPAC-1 GSE119930.JUNB.CFPAC-1 134 bp overlap
KDM5B 1 dataset
ChIP HCC2157 GSE46055.KDM5B.HCC2157 249 bp overlap
KMT2A 1 dataset
ChIP blood_cord GSE83671.KMT2A.blood_cord 258 bp overlap
MED1 6 datasets
ChIP CD4_Th1_DMSO GSE62482.MED1.CD4_Th1_DMSO 111 bp overlap
ChIP SGBS GSE64233.MED1.SGBS 180 bp overlap
ChIP SGBS_TNF GSE64233.MED1.SGBS_TNF 179 bp overlap
ChIP U-87MG GSE36354.MED1.U-87MG 319 bp overlap
ChIP hMSC-TERT4_D7 GSE104537.MED1.hMSC-TERT4_D7 204 bp overlap
ChIP hMSC-TERT4_adipocyte-D14 GSE113253.MED1.hMSC-TERT4_adipocyte-D14 186 bp overlap
MED12 1 dataset
ChIP leiomyoma_PT886 GSE128230.MED12.leiomyoma_PT886 82 bp overlap
MXI1 2 datasets
Motif DE_12h DE_12h-MXI1_MA1108.3 6 bp overlap
ChIP IMR-90 ENCSR000EFE.MXI1.IMR-90 134 bp overlap
MYB 2 datasets
ChIP THP-1 GSE90769.MYB.THP-1 277 bp overlap
ChIP THP-1_OG86 GSE90769.MYB.THP-1_OG86 241 bp overlap
MYC 2 datasets
ChIP SK-N-AS GSE138295.MYC.SK-N-AS 111 bp overlap
ChIP SK-N-AS GSE138295.MYC.SK-N-AS 98 bp overlap
MYOD1 1 dataset
ChIP RD_shSNAI2 GSE137168.MYOD1.RD_shSNAI2 168 bp overlap
NANOG 2 datasets
ChIP HUES-8 GSE109524.NANOG.HUES-8 179 bp overlap
ChIP WA01_3IL ERP004238.NANOG.WA01_3IL 196 bp overlap
NCAPH2 4 datasets
ChIP IMR-90_Triptolide-30min GSE118494.NCAPH2.IMR-90_Triptolide-30min 180 bp overlap
ChIP IMR-90_alpha-amanitin-1H GSE118494.NCAPH2.IMR-90_alpha-amanitin-1H 269 bp overlap
ChIP IMR-90_alpha-amanitin-30min GSE118494.NCAPH2.IMR-90_alpha-amanitin-30min 319 bp overlap
ChIP IMR-90_ctrl_OIS GSE118494.NCAPH2.IMR-90_ctrl_OIS 319 bp overlap
NFATC3 1 dataset
Motif DE_12h DE_12h-NFATC3_MA0625.3 6 bp overlap
NFATC4 1 dataset
Motif DE_12h DE_12h-NFATC4_MA1525.3 9 bp overlap
NFE2 1 dataset
ChIP ProEs GSE59087.NFE2.ProEs 167 bp overlap
NR3C1 1 dataset
ChIP SUM159PT GSE152203.NR3C1.SUM159PT 234 bp overlap
Nfat5 1 dataset
Motif DE_12h DE_12h-Nfat5_MA0606.3 8 bp overlap
Nfatc1 1 dataset
Motif DE_12h DE_12h-Nfatc1_MA0624.3 6 bp overlap
POLR2A 4 datasets
ChIP sigmoid colon ENCFF101ILL 122 bp overlap
ChIP sigmoid colon ENCFF725QFT 90 bp overlap
ChIP sigmoid colon ENCFF748YVT 90 bp overlap
ChIP sigmoid colon ENCFF754JQR 198 bp overlap
POU2F2 1 dataset
Motif DE_12h DE_12h-POU2F2_MA0507.3 13 bp overlap
POU2F3 1 dataset
Motif DE_12h DE_12h-POU2F3_MA0627.3 9 bp overlap
POU3F1 1 dataset
Motif DE_12h DE_12h-POU3F1_MA0786.2 10 bp overlap
POU3F4 1 dataset
Motif DE_12h DE_12h-POU3F4_MA0789.1 9 bp overlap
POU5F1 1 dataset
Motif DE_12h DE_12h-POU5F1_MA1115.2 7 bp overlap
PRDM6 1 dataset
ChIP HEK293 ENCFF283AJL 121 bp overlap
RAD21 16 datasets
ChIP IMR-90 ENCFF752PTH 118 bp overlap
ChIP MDM GSE103477.RAD21.MDM 319 bp overlap
ChIP MDM_-dNS1 GSE103477.RAD21.MDM_-dNS1 297 bp overlap
ChIP MDM_H5N1 GSE103477.RAD21.MDM_H5N1 311 bp overlap
ChIP MDM_IFNb GSE103477.RAD21.MDM_IFNb 307 bp overlap
ChIP THP-1_PMA_Dex-0h GSE103477.RAD21.THP-1_PMA_Dex-0h 319 bp overlap
ChIP THP-1_PMA_Dex-6h GSE103477.RAD21.THP-1_PMA_Dex-6h 319 bp overlap
ChIP THP-1_PMA_IFNb-6h GSE103477.RAD21.THP-1_PMA_IFNb-6h 164 bp overlap
ChIP THP-1_Pam3csk-000m-Flavo-000m GSE103477.RAD21.THP-1_Pam3csk-000m-Flavo-000m 275 bp overlap
ChIP THP-1_Pam3csk-000m-Flavo-240m GSE103477.RAD21.THP-1_Pam3csk-000m-Flavo-240m 238 bp overlap
ChIP THP-1_Pam3csk-020m-Flavo-000m GSE103477.RAD21.THP-1_Pam3csk-020m-Flavo-000m 277 bp overlap
ChIP THP-1_Pam3csk-025m-Flavo-000m GSE103477.RAD21.THP-1_Pam3csk-025m-Flavo-000m 288 bp overlap
ChIP THP-1_Pam3csk-150m-Flavo-030m GSE103477.RAD21.THP-1_Pam3csk-150m-Flavo-030m 261 bp overlap
ChIP THP-1_Pam3csk-360m-Flavo-240m GSE103477.RAD21.THP-1_Pam3csk-360m-Flavo-240m 285 bp overlap
ChIP THP-1_siCtrl-eGFP-Pam3csk-4h GSE103477.RAD21.THP-1_siCtrl-eGFP-Pam3csk-4h 225 bp overlap
ChIP peripheral-blood-neutrophil_US-1 GSE126755.RAD21.peripheral-blood-neutrophil_US-1 315 bp overlap
RELA 6 datasets
ChIP CD4_Th1_DMSO GSE62482.RELA.CD4_Th1_DMSO 319 bp overlap
ChIP HAEC_IL1b_4h GSE89970.RELA.HAEC_IL1b_4h 91 bp overlap
ChIP IMR-90_TNF GSE43070.RELA.IMR-90_TNF 66 bp overlap
ChIP SGBS GSE64233.RELA.SGBS 172 bp overlap
ChIP SGBS_TNF GSE64233.RELA.SGBS_TNF 317 bp overlap
ChIP THP-1_eGFP-Pam3csk-4h GSE103477.RELA.THP-1_eGFP-Pam3csk-4h 203 bp overlap
REST 1 dataset
ChIP hippocampus GSE144226.REST.hippocampus 281 bp overlap
RXR 1 dataset
ChIP macrophage ERP008801.RXR.macrophage 263 bp overlap
SIX2 2 datasets
ChIP HEK GSE73865.SIX2.HEK 110 bp overlap
ChIP kidney_fetal_17w GSE75948.SIX2.kidney_fetal_17w 90 bp overlap
SMAD3 1 dataset
ChIP HCC1954 GSE104760.SMAD3.HCC1954 281 bp overlap
SMARCC1 3 datasets
ChIP Aska-SS GSE108025.SMARCC1.Aska-SS 104 bp overlap
ChIP Aska-SS_shSSX GSE108025.SMARCC1.Aska-SS_shSSX 319 bp overlap
ChIP SCCOHT-1 GSE117734.SMARCC1.SCCOHT-1 319 bp overlap
SMC1A 1 dataset
ChIP monocyte_INFg GSE98367.SMC1A.monocyte_INFg 238 bp overlap
SMC3 1 dataset
ChIP IMR-90 ENCSR000HPG.SMC3.IMR-90 150 bp overlap
SNAI2 1 dataset
ChIP RD GSE137168.SNAI2.RD 212 bp overlap
SPI1 8 datasets
ChIP DC_96h_donorM GSE128834.SPI1.DC_96h_donorM 110 bp overlap
ChIP macrophage_D7_donorP GSE128834.SPI1.macrophage_D7_donorP 229 bp overlap
ChIP monocyte GSE31621.SPI1.monocyte 208 bp overlap
ChIP monocyte_MACROPHAGE GSE31621.SPI1.monocyte_MACROPHAGE 154 bp overlap
ChIP primary-monocyte_18h_donorO GSE128834.SPI1.primary-monocyte_18h_donorO 319 bp overlap
ChIP primary-monocyte_4h_donorO GSE128834.SPI1.primary-monocyte_4h_donorO 218 bp overlap
ChIP primary-monocyte_LPS-4h_donorO GSE128834.SPI1.primary-monocyte_LPS-4h_donorO 314 bp overlap
ChIP primary-monocyte_donorA_ncMO GSE128834.SPI1.primary-monocyte_donorA_ncMO 129 bp overlap
SREBP2 2 datasets
ChIP monocyte GSE129202.SREBP2.monocyte 278 bp overlap
ChIP monocyte_TNF GSE129202.SREBP2.monocyte_TNF 319 bp overlap
SRF 1 dataset
Motif DE_12h DE_12h-SRF_MA0083.3 16 bp overlap
STAG1 1 dataset
ChIP MCF-7 ERP000209.STAG1.MCF-7 162 bp overlap
STAT1 3 datasets
ChIP CD14_INFG GSE43036.STAT1.CD14_INFG 191 bp overlap
ChIP CD14_INFG_LPS GSE43036.STAT1.CD14_INFG_LPS 215 bp overlap
ChIP CD14_LPS GSE43036.STAT1.CD14_LPS 220 bp overlap
STAT3 9 datasets
ChIP HCC1187 GSE152203.STAT3.HCC1187 199 bp overlap
ChIP MDA-MB-468_EtOH GSE85579.STAT3.MDA-MB-468_EtOH 319 bp overlap
ChIP SUM159PT GSE152203.STAT3.SUM159PT 195 bp overlap
ChIP breast-cancer_3487 GSE126004.STAT3.breast-cancer_3487 191 bp overlap
ChIP monocyte_IFNg GSE120943.STAT3.monocyte_IFNg 187 bp overlap
ChIP monocyte_IFNg-LPS GSE120943.STAT3.monocyte_IFNg-LPS 210 bp overlap
ChIP monocyte_IL10 GSE120943.STAT3.monocyte_IL10 317 bp overlap
ChIP monocyte_LPS GSE120943.STAT3.monocyte_LPS 307 bp overlap
ChIP monocyte_resting GSE120943.STAT3.monocyte_resting 141 bp overlap
TBX21 1 dataset
ChIP Th1_CD3-CD28_donor2 GSE81881.TBX21.Th1_CD3-CD28_donor2 173 bp overlap
TCF12 2 datasets
ChIP Kasumi-1 GSE23730.TCF12.Kasumi-1 178 bp overlap
ChIP SK-N-SH ENCFF147AHB 183 bp overlap
TEAD1 1 dataset
ChIP HUCCT1 GSE68296.TEAD1.HUCCT1 114 bp overlap
ZBTB40 1 dataset
ChIP MCF-7 ENCFF044DWL 123 bp overlap