chr2 : 9,218,015 9,218,554
539 bp 107 TFs 5 linked genes
This 539 bp open chromatin element is linked to 5 target genes and is bound by 107 transcription factors.
Linked Genes
5 genes
Gene Expression Dist. to TSS Distance Link type
ASAP2 11.4 kb Distal Multiome
ITGB1BP1 205.4 kb Distal Multiome
CPSF3 205.5 kb Distal Multiome
MBOAT2 214.5 kb Distal Multiome
IAH1 255.6 kb Distal Multiome
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr2:9,213,015 – 9,223,554
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
107 transcription factors
Source
Cell type
AR 1 dataset
ChIP LNCaP-abl GSE39459.AR.LNCaP-abl 117 bp overlap
ASCL1 2 datasets
Motif DE_12h DE_12h-ASCL1_MA1631.2 9 bp overlap
ChIP SCLC_ASCLP GSE61197.ASCL1.SCLC_ASCLP 129 bp overlap
ATF2 1 dataset
ChIP WA01 ENCSR000BQU.ATF2.WA01 172 bp overlap
Arid3a 1 dataset
Motif DE_24h DE_24h-Arid3a_MA0151.1 6 bp overlap
BCL6B 1 dataset
Motif DE_24h DE_24h-BCL6B_MA0731.1 17 bp overlap
BCOR 3 datasets
ChIP WA01 GSE104690.BCOR.WA01 381 bp overlap
ChIP WA01_KDM2B_KO GSE104690.BCOR.WA01_KDM2B_KO 272 bp overlap
ChIP WA01_RNF2-R_Doxycyclin GSE104690.BCOR.WA01_RNF2-R_Doxycyclin 156 bp overlap
CDKN1B 1 dataset
ChIP MDA-BoM-1833_shp27 GSE112444.CDKN1B.MDA-BoM-1833_shp27 173 bp overlap
CREBBP 1 dataset
ChIP keratinocyte_KLF3i GSE140991.CREBBP.keratinocyte_KLF3i 137 bp overlap
Cebpa 2 datasets
ChIP BLaER1 ENCFF093OYK 140 bp overlap
ChIP BLaER1 ENCFF262VBH 203 bp overlap
EOMES 1 dataset
ChIP hESC GSE26097.EOMES.hESC 364 bp overlap
EP300 3 datasets
ChIP neural ENCSR843ZUP.EP300.neural 413 bp overlap
ChIP neural cell ENCFF442QNK 441 bp overlap
ChIP neural cell ENCFF442QNK 414 bp overlap
ESR1 10 datasets
ChIP Ishikawa ENCSR000BIY.ESR1.Ishikawa 190 bp overlap
ChIP Ishikawa ENCSR000BKL.ESR1.Ishikawa 145 bp overlap
ChIP Ishikawa_DMSO_D538G_clone3 GSE132426.ESR1.Ishikawa_DMSO_D538G_clone3 391 bp overlap
ChIP Ishikawa_E2 GSE132426.ESR1.Ishikawa_E2 234 bp overlap
ChIP Ishikawa_E2_D538G GSE132426.ESR1.Ishikawa_E2_D538G 395 bp overlap
ChIP Ishikawa_E2_D538G_clone2 GSE132426.ESR1.Ishikawa_E2_D538G_clone2 410 bp overlap
ChIP Ishikawa_ETV4-KO1 GSE129803.ESR1.Ishikawa_ETV4-KO1 230 bp overlap
ChIP Ishikawa_ETV4-KO2_Mili GSE129803.ESR1.Ishikawa_ETV4-KO2_Mili 274 bp overlap
ChIP Ishikawa_M2-5 GSE147141.ESR1.Ishikawa_M2-5 211 bp overlap
ChIP U2OS_E2 GSE26110.ESR1.U2OS_E2 183 bp overlap
ETV5::HOXA2 2 datasets
Motif DE_12h DE_12h-ETV5HOXA2_MA1948.2 12 bp overlap
Motif DE_24h DE_24h-ETV5HOXA2_MA1948.2 12 bp overlap
FIGLA 2 datasets
Motif DE_12h DE_12h-FIGLA_MA0820.2 6 bp overlap
Motif DE_24h DE_24h-FIGLA_MA0820.2 6 bp overlap
FOSL2 1 dataset
ChIP LPS141 GSE111253.FOSL2.LPS141 296 bp overlap
FOXA2 2 datasets
ChIP DE DE-FOXA2-1 390 bp overlap
ChIP DE DE-FOXA2-2 303 bp overlap
Foxj2 1 dataset
Motif DE_12h DE_12h-Foxj2_MA0614.1 8 bp overlap
GATA4 3 datasets
ChIP DE DE-GATA4-1 465 bp overlap
ChIP DE DE-GATA4-2 434 bp overlap
ChIP foregut GSE117136.GATA4.foregut 283 bp overlap
GATA6 11 datasets
ChIP AGS GSE51936.GATA6.AGS 68 bp overlap
ChIP DE DE-GATA6-1 377 bp overlap
ChIP DE DE-GATA6-2 393 bp overlap
ChIP DE_D1 S09-DE-d1-GATA6-exp1 419 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 439 bp overlap
ChIP DE_D1 S41-DE-d1-GATA6-exp2 529 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 381 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 453 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 520 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 370 bp overlap
ChIP foregut GSE117136.GATA6.foregut 349 bp overlap
GLIS1 1 dataset
ChIP HEK293 ENCSR482BBZ.GLIS1.HEK293 284 bp overlap
GLIS3 1 dataset
ChIP H9_plus GSE109562.GLIS3.H9_plus 448 bp overlap
HDAC2 1 dataset
ChIP H1 ENCFF939VKA 237 bp overlap
HDAC3 1 dataset
ChIP RH4_DMSO-6H GSE116344.HDAC3.RH4_DMSO-6H 210 bp overlap
HIC2 1 dataset
Motif DE_24h DE_24h-HIC2_MA0738.2 6 bp overlap
JUN 4 datasets
ChIP DE_D1 S08-DE-d1-JUN-exp1 341 bp overlap
ChIP DE_D1 S40-DE-d1-JUN-exp2 484 bp overlap
ChIP DE_D2 S02-DE-d2-JUN-exp1 328 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.JUN.HUES-8_DE_ctrl 449 bp overlap
MAFF 3 datasets
ChIP HeLa-S3 ENCSR140DSL.MAFF.HeLa-S3 61 bp overlap
ChIP HepG2 ENCFF452YUT 89 bp overlap
ChIP K562 ENCFF071YKK 79 bp overlap
MAFG 1 dataset
ChIP K562 ENCFF455EEO 136 bp overlap
MAFK 3 datasets
ChIP Hep-G2 ENCSR000EDZ.MAFK.Hep-G2 107 bp overlap
ChIP HepG2 ENCFF743ZOF 58 bp overlap
ChIP HepG2 ENCFF767LDG 60 bp overlap
MED1 1 dataset
ChIP U-87MG GSE36354.MED1.U-87MG 110 bp overlap
MED26 1 dataset
ChIP HEK293T GSE121024.MED26.HEK293T 179 bp overlap
MEIS1 5 datasets
Motif DE_12h DE_12h-MEIS1_MA0498.3 5 bp overlap
Motif DE_12h DE_12h-MEIS1_MA0498.3 5 bp overlap
Motif DE_12h DE_12h-MEIS1_MA0498.3 5 bp overlap
Motif DE_24h DE_24h-MEIS1_MA0498.3 5 bp overlap
Motif DE_24h DE_24h-MEIS1_MA0498.3 5 bp overlap
MORC2 1 dataset
ChIP H9 GSE95374.MORC2.H9 250 bp overlap
MYOD1 1 dataset
Motif DE_24h DE_24h-MYOD1_MA0499.3 9 bp overlap
MYOG 1 dataset
ChIP RH30_DMSO GSE85169.MYOG.RH30_DMSO 223 bp overlap
NANOG 11 datasets
ChIP GM23338 ENCFF065NZG 311 bp overlap
ChIP GM23338 ENCSR061DGF.NANOG.GM23338 271 bp overlap
ChIP H1 ENCFF747ZPQ 80 bp overlap
ChIP HUES-8 GSE109524.NANOG.HUES-8 497 bp overlap
ChIP WA01 ERP004238.NANOG.WA01 380 bp overlap
ChIP WA01 ENCSR000BMT.NANOG.WA01 315 bp overlap
ChIP WA01_3IL ERP004238.NANOG.WA01_3IL 539 bp overlap
ChIP WA09 GSE105028.NANOG.WA09 454 bp overlap
ChIP WA09_heat-shock GSE105028.NANOG.WA09_heat-shock 443 bp overlap
ChIP hESC GSE18292.NANOG.hESC 290 bp overlap
ChIP hESC GSE20650.NANOG.hESC 314 bp overlap
NIPBL 1 dataset
ChIP WA09_heat-shock GSE105028.NIPBL.WA09_heat-shock 260 bp overlap
NR2C1 2 datasets
Motif DE_12h DE_12h-NR2C1_MA1535.2 6 bp overlap
Motif DE_24h DE_24h-NR2C1_MA1535.2 6 bp overlap
NR2C2 2 datasets
Motif DE_12h DE_12h-NR2C2_MA1536.2 6 bp overlap
Motif DE_24h DE_24h-NR2C2_MA1536.2 6 bp overlap
NR2F1 2 datasets
Motif DE_12h DE_12h-NR2F1_MA0017.3 12 bp overlap
Motif DE_24h DE_24h-NR2F1_MA0017.3 12 bp overlap
Nanog 2 datasets
Motif DE_12h DE_12h-Nanog_MA2339.1 7 bp overlap
Motif DE_24h DE_24h-Nanog_MA2339.1 7 bp overlap
Nr1H2 2 datasets
Motif DE_12h DE_12h-Nr1H2_MA1996.2 6 bp overlap
Motif DE_24h DE_24h-Nr1H2_MA1996.2 6 bp overlap
Nr1H4 2 datasets
Motif DE_12h DE_12h-Nr1H4_MA1110.3 6 bp overlap
Motif DE_24h DE_24h-Nr1H4_MA1110.3 6 bp overlap
Nr1h3 2 datasets
Motif DE_12h DE_12h-Nr1h3_MA2337.1 6 bp overlap
Motif DE_24h DE_24h-Nr1h3_MA2337.1 6 bp overlap
OTX2 1 dataset
ChIP WTC11 ENCFF634NAO 245 bp overlap
PBX3 2 datasets
Motif DE_12h DE_12h-PBX3_MA1114.2 11 bp overlap
Motif DE_24h DE_24h-PBX3_MA1114.2 11 bp overlap
PKNOX1 1 dataset
Motif DE_24h DE_24h-PKNOX1_MA0782.3 10 bp overlap
POLR2A 1 dataset
ChIP neural cell ENCFF604SPB 154 bp overlap
POU5F1 5 datasets
ChIP DE_D1 DED1-OCT4_Batch_II 369 bp overlap
ChIP NCCIT_SNF5 GSE36134.POU5F1.NCCIT_SNF5 289 bp overlap
ChIP WA01_3IL ERP004238.POU5F1.WA01_3IL 169 bp overlap
ChIP hESC_PRIMED GSE69646.POU5F1.hESC_PRIMED 254 bp overlap
ChIP hiPSC_3s2 GSE81899.POU5F1.hiPSC_3s2 146 bp overlap
POU5F1_M 1 dataset
ChIP DE_D1 DED1-OCT4-M_Batch_II 459 bp overlap
PRDM14 1 dataset
ChIP NCCIT GSE71675.PRDM14.NCCIT 373 bp overlap
Plagl1 1 dataset
Motif DE_12h DE_12h-Plagl1_MA1615.2 8 bp overlap
Ptf1A 1 dataset
Motif DE_24h DE_24h-Ptf1A_MA1620.2 8 bp overlap
RARA 1 dataset
Motif DE_24h DE_24h-RARA_MA0730.1 17 bp overlap
RBM22 1 dataset
ChIP HepG2 ENCFF561IAJ 113 bp overlap
RELA 1 dataset
ChIP LNCaP_SIFOXA1_TNFA GSE83860.RELA.LNCaP_SIFOXA1_TNFA 129 bp overlap
REST 1 dataset
ChIP neural ENCSR000BTV.REST.neural 209 bp overlap
RNF2 1 dataset
ChIP WA09 GSE105028.RNF2.WA09 236 bp overlap
RREB1 1 dataset
Motif DE_24h DE_24h-RREB1_MA0073.2 19 bp overlap
RXRA 1 dataset
ChIP H1 ENCFF570NHK 201 bp overlap
Rarb 1 dataset
Motif DE_24h DE_24h-Rarb_MA0858.1 17 bp overlap
SALL3 1 dataset
ChIP hiPSC GSE104863.SALL3.hiPSC 204 bp overlap
SMAD2 1 dataset
ChIP endoderm GSE29422.SMAD2.endoderm 177 bp overlap
SMAD2-3 3 datasets
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 539 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.SMAD2-3.HUES-8_DE_JNKi 451 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.SMAD2-3.HUES-8_DE_ctrl 480 bp overlap
SMAD2_3 5 datasets
ChIP DE_D1 S07-DE-d1-SMAD2_3-exp1 455 bp overlap
ChIP DE_D1 S12-DE-d1-SMAD2_3-exp1 429 bp overlap
ChIP DE_D1 S39-DE-d1-SMAD2_3-exp2 457 bp overlap
ChIP DE_D2 S01-DE-d2-SMAD2_3-exp1 286 bp overlap
ChIP DE_D2 S53-DE-d2-SMAD2_3-exp2 404 bp overlap
SMAD3 6 datasets
ChIP BG03_DIFF_2H GSE36578.SMAD3.BG03_DIFF_2H 156 bp overlap
ChIP BG03_DIFF_48H GSE36578.SMAD3.BG03_DIFF_48H 200 bp overlap
ChIP HMLE_Doxycicline_TGFb GSE104760.SMAD3.HMLE_Doxycicline_TGFb 282 bp overlap
ChIP HMLE_TGFb GSE104760.SMAD3.HMLE_TGFb 221 bp overlap
ChIP NCI-H441_SIITTF1 GSE51509.SMAD3.NCI-H441_SIITTF1 137 bp overlap
ChIP endoderm GSE29422.SMAD3.endoderm 224 bp overlap
SMAD4 1 dataset
ChIP endoderm GSE29422.SMAD4.endoderm 292 bp overlap
SMARCA4 2 datasets
ChIP hiPSC GSE124903.SMARCA4.hiPSC 164 bp overlap
ChIP hiPSC_WT GSE124903.SMARCA4.hiPSC_WT 209 bp overlap
SMARCC1 2 datasets
ChIP DE_D1 S15-DE-d1-BAF155-exp1 258 bp overlap
ChIP hiPSC_WT GSE124903.SMARCC1.hiPSC_WT 280 bp overlap
SMC1A 1 dataset
ChIP primary-epidermal-keratinocyte GSE85526.SMC1A.primary-epidermal-keratinocyte 172 bp overlap
SNAI1 1 dataset
Motif DE_12h DE_12h-SNAI1_MA1558.2 7 bp overlap
SNAI2 3 datasets
Motif DE_24h DE_24h-SNAI2_MA0745.3 8 bp overlap
ChIP RD GSE137168.SNAI2.RD 157 bp overlap
ChIP keratinocyte_LacZ GSE55421.SNAI2.keratinocyte_LacZ 235 bp overlap
SNAI3 1 dataset
Motif DE_24h DE_24h-SNAI3_MA1559.2 9 bp overlap
SOX13 2 datasets
Motif DE_12h DE_12h-SOX13_MA1120.2 7 bp overlap
Motif DE_24h DE_24h-SOX13_MA1120.2 7 bp overlap
SOX17_M 1 dataset
ChIP DE_D2 DED2-SOX17-M_Batch_II 351 bp overlap
SOX2 6 datasets
Motif DE_12h DE_12h-SOX2_MA0143.5 7 bp overlap
Motif DE_24h DE_24h-SOX2_MA0143.5 7 bp overlap
ChIP H9 GSE46837.SOX2.H9 301 bp overlap
ChIP hESC GSE18292.SOX2.hESC 95 bp overlap
ChIP hiPSC GSE56567.SOX2.hiPSC 366 bp overlap
ChIP hiPSC_3s2 GSE81899.SOX2.hiPSC_3s2 290 bp overlap
SOX21 1 dataset
Motif DE_12h DE_12h-SOX21_MA0866.1 15 bp overlap
SP1 2 datasets
ChIP H1 ENCFF263FUH 321 bp overlap
ChIP WA01 ENCSR000BIR.SP1.WA01 170 bp overlap
STAT3 1 dataset
ChIP SUM159PT GSE152203.STAT3.SUM159PT 170 bp overlap
Sox1 1 dataset
Motif DE_12h DE_12h-Sox1_MA0870.1 15 bp overlap
Sox17 2 datasets
Motif DE_12h DE_12h-Sox17_MA0078.3 10 bp overlap
Motif DE_24h DE_24h-Sox17_MA0078.3 10 bp overlap
Sox3 2 datasets
Motif DE_12h DE_12h-Sox3_MA0514.3 7 bp overlap
Motif DE_24h DE_24h-Sox3_MA0514.3 7 bp overlap
Sox5 2 datasets
Motif DE_12h DE_12h-Sox5_MA0087.3 8 bp overlap
Motif DE_24h DE_24h-Sox5_MA0087.3 8 bp overlap
T 2 datasets
ChIP H9_ENDODERM GSE60606.T.H9_ENDODERM 268 bp overlap
ChIP H9_MESODERM GSE60606.T.H9_MESODERM 286 bp overlap
TCF12 5 datasets
Motif DE_12h DE_12h-TCF12_MA1648.2 7 bp overlap
ChIP H1 ENCFF203EBH 251 bp overlap
ChIP Ishikawa ENCFF467DDW 460 bp overlap
ChIP Ishikawa ENCSR000BUV.TCF12.Ishikawa 234 bp overlap
ChIP WA01 ENCSR000BIT.TCF12.WA01 300 bp overlap
TCF3 2 datasets
Motif DE_12h DE_12h-TCF3_MA0522.4 7 bp overlap
ChIP NPC GSE154479.TCF3.NPC 311 bp overlap
TEAD1 1 dataset
ChIP WTC11 ENCFF502QUV 374 bp overlap
TFAP2B 2 datasets
Motif DE_12h DE_12h-TFAP2B_MA0811.2 11 bp overlap
Motif DE_24h DE_24h-TFAP2B_MA0811.2 11 bp overlap
TFAP2C 1 dataset
ChIP UCLA1-hESCs_TFAP2C--_1d_TFAP2C_Induc GSE101074.TFAP2C.UCLA1-hESCs_TFAP2C--_1d_TFAP2C_Induc 280 bp overlap
TP53 1 dataset
ChIP hESC_DAMAGED GSE39912.TP53.hESC_DAMAGED 286 bp overlap
TP63 11 datasets
ChIP HaCaT_LacZ_TGFB GSE60814.TP63.HaCaT_LacZ_TGFB 97 bp overlap
ChIP HaCaT_caRAS_TGFB GSE60814.TP63.HaCaT_caRAS_TGFB 258 bp overlap
ChIP MCF-10A_Nutlin3A GSE111009.TP63.MCF-10A_Nutlin3A 228 bp overlap
ChIP SUIT-2 GSE115461.TP63.SUIT-2 334 bp overlap
ChIP keratinocyte GSE56674.TP63.keratinocyte 83 bp overlap
ChIP keratinocyte_CISP GSE56674.TP63.keratinocyte_CISP 165 bp overlap
ChIP keratinocyte_D0 GSE59824.TP63.keratinocyte_D0 184 bp overlap
ChIP keratinocyte_D2 GSE59824.TP63.keratinocyte_D2 189 bp overlap
ChIP keratinocyte_D4 GSE59824.TP63.keratinocyte_D4 205 bp overlap
ChIP keratinocyte_D7 GSE59824.TP63.keratinocyte_D7 212 bp overlap
ChIP keratinocyte_epidermal GSE67382.TP63.keratinocyte_epidermal 117 bp overlap
USF2 1 dataset
ChIP IMR-90 ENCSR513UQG.USF2.IMR-90 114 bp overlap
Wt1 1 dataset
Motif DE_24h DE_24h-Wt1_MA1627.2 10 bp overlap
ZEB1 2 datasets
Motif DE_12h DE_12h-ZEB1_MA0103.4 6 bp overlap
Motif DE_24h DE_24h-ZEB1_MA0103.4 6 bp overlap
ZIC2 1 dataset
ChIP HEK293 ENCFF033NQQ 367 bp overlap
ZNF316 1 dataset
ChIP K562 ENCFF838QCD 81 bp overlap
ZNF410 1 dataset
Motif DE_24h DE_24h-ZNF410_MA0752.2 16 bp overlap
ZNF423 1 dataset
ChIP WTC11 ENCFF574PBR 317 bp overlap
ZNF449 1 dataset
Motif DE_24h DE_24h-ZNF449_MA1656.2 10 bp overlap
ZNF454 1 dataset
Motif DE_12h DE_12h-ZNF454_MA1712.2 17 bp overlap
ZNF667 1 dataset
Motif DE_24h DE_24h-ZNF667_MA1984.2 11 bp overlap
ZNF680 2 datasets
Motif DE_12h DE_12h-ZNF680_MA1729.2 11 bp overlap
Motif DE_24h DE_24h-ZNF680_MA1729.2 11 bp overlap
ZNF75A 1 dataset
Motif DE_12h DE_12h-ZNF75A_MA2097.1 12 bp overlap
Zic2 1 dataset
Motif DE_24h DE_24h-Zic2_MA1629.2 9 bp overlap
Zic3 1 dataset
Motif DE_24h DE_24h-Zic3_MA0697.3 7 bp overlap