chr12 : 54,047,114 54,047,863
749 bp 89 TFs 1 linked gene
This 749 bp open chromatin element is linked to HOXC4 and is bound by 89 transcription factors.
Linked Genes
1 gene
Gene Expression Dist. to TSS Distance Link type
HOXC4 6.0 kb Proximal Proximity
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr12:54,042,114 – 54,052,863
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
89 transcription factors
Source
Cell type
AATF 1 dataset
ChIP NALM-6 GSE93626.AATF.NALM-6 265 bp overlap
AGO1 3 datasets
ChIP Hep-G2 ENCSR555ZMV.AGO1.Hep-G2 183 bp overlap
ChIP Hep-G2 GSE120104.AGO1.Hep-G2 183 bp overlap
ChIP K-562 ENCSR641BSL.AGO1.K-562 181 bp overlap
AR 1 dataset
ChIP myofibroblast GSE90772.AR.myofibroblast 552 bp overlap
ASH2L 4 datasets
ChIP H1 ENCFF399KAM 635 bp overlap
ChIP H1 ENCFF399KAM 700 bp overlap
ChIP WA01 ENCSR850KIP.ASH2L.WA01 287 bp overlap
ChIP WA01 ENCSR850KIP.ASH2L.WA01 116 bp overlap
ATF2 1 dataset
ChIP WA01 ENCSR000BQU.ATF2.WA01 117 bp overlap
BCOR 1 dataset
ChIP WA01_KDM2B_KO GSE104690.BCOR.WA01_KDM2B_KO 749 bp overlap
BRD1 2 datasets
ChIP HUES-64 GSE104059.BRD1.HUES-64 248 bp overlap
ChIP HUES-64 GSE104059.BRD1.HUES-64 145 bp overlap
BRD4 10 datasets
ChIP 22Rv1_DHT-DMSO GSE118247.BRD4.22Rv1_DHT-DMSO 427 bp overlap
ChIP CLL_patient1_4h_CpG GSE109411.BRD4.CLL_patient1_4h_CpG 322 bp overlap
ChIP CLL_patient2_4h_CpG_BET GSE109411.BRD4.CLL_patient2_4h_CpG_BET 580 bp overlap
ChIP CLL_patient3_4h_CpG GSE109411.BRD4.CLL_patient3_4h_CpG 226 bp overlap
ChIP CLL_patient4 GSE109411.BRD4.CLL_patient4 303 bp overlap
ChIP CLL_patient4_4h_CpG_BET GSE109411.BRD4.CLL_patient4_4h_CpG_BET 140 bp overlap
ChIP COLO-741 GSE73319.BRD4.COLO-741 237 bp overlap
ChIP IMR-90_QUIES GSE74238.BRD4.IMR-90_QUIES 299 bp overlap
ChIP retina_AB1-FW15 GSE86981.BRD4.retina_AB1-FW15 697 bp overlap
ChIP retina_AB1-RB GSE86981.BRD4.retina_AB1-RB 297 bp overlap
CBFB 1 dataset
ChIP ME-1 GSE46044.CBFB.ME-1 509 bp overlap
CBX7 1 dataset
ChIP lymphocyte_DMSO GSE110139.CBX7.lymphocyte_DMSO 130 bp overlap
CBX8 1 dataset
ChIP H1 ENCFF095JHA 180 bp overlap
CHD1 2 datasets
ChIP WA01 ENCSR000AQK.CHD1.WA01 287 bp overlap
ChIP WA01 ENCSR000AQK.CHD1.WA01 240 bp overlap
CTBP2 4 datasets
ChIP H1 ENCFF329MAX 51 bp overlap
ChIP H1 ENCFF329MAX 465 bp overlap
ChIP H1 ENCFF329MAX 583 bp overlap
ChIP H1 ENCFF329MAX 154 bp overlap
CTCF 2 datasets
ChIP HAP1_WT GSE126634.CTCF.HAP1_WT 239 bp overlap
ChIP T-47D_Y537S GSE148277.CTCF.T-47D_Y537S 385 bp overlap
Cebpa 3 datasets
ChIP BLaER1 ENCFF031ISE 350 bp overlap
ChIP BLaER1 ENCFF364PUR 251 bp overlap
ChIP BLaER1 ENCFF364PUR 360 bp overlap
EED 2 datasets
ChIP ProEs GSE59087.EED.ProEs 207 bp overlap
ChIP ProEs GSE59087.EED.ProEs 350 bp overlap
ERG 4 datasets
ChIP CD34_NR29 GSE23730.ERG.CD34_NR29 318 bp overlap
ChIP K-562_DOX GSE23730.ERG.K-562_DOX 170 bp overlap
ChIP MCF-7 GSE23730.ERG.MCF-7 406 bp overlap
ChIP ME-1 GSE46044.ERG.ME-1 207 bp overlap
ESR1 12 datasets
ChIP Ishikawa GSE99905.ESR1.Ishikawa 494 bp overlap
ChIP Ishikawa_E2-1h GSE129803.ESR1.Ishikawa_E2-1h 749 bp overlap
ChIP Ishikawa_siETV4-72h GSE129803.ESR1.Ishikawa_siETV4-72h 146 bp overlap
ChIP Ishikawa_siNTC-72h GSE129803.ESR1.Ishikawa_siNTC-72h 209 bp overlap
ChIP Ishikawa_siNTC-96h GSE129803.ESR1.Ishikawa_siNTC-96h 110 bp overlap
ChIP MCF-7_PaPE-1 GSE93510.ESR1.MCF-7_PaPE-1 308 bp overlap
ChIP MCF-7_PaPE-1_OA GSE93510.ESR1.MCF-7_PaPE-1_OA 456 bp overlap
ChIP MCF-7_SHCTR_E2_TNF GSE59530.ESR1.MCF-7_SHCTR_E2_TNF 238 bp overlap
ChIP MCF-7_Y537S_DMSO GSE148277.ESR1.MCF-7_Y537S_DMSO 231 bp overlap
ChIP MCF-7_estradiol-aldosterone_45min GSE99626.ESR1.MCF-7_estradiol-aldosterone_45min 283 bp overlap
ChIP T-47D_DMSO GSE148277.ESR1.T-47D_DMSO 192 bp overlap
ChIP T-47D_Y537S GSE148277.ESR1.T-47D_Y537S 129 bp overlap
ETS1 1 dataset
ChIP cardiomyocyte_D0 GSE129986.ETS1.cardiomyocyte_D0 150 bp overlap
EZH2 30 datasets
ChIP B cell ENCFF803EMO 411 bp overlap
ChIP B cell ENCFF803EMO 425 bp overlap
ChIP GM12878 ENCFF635TDF 137 bp overlap
ChIP GM23338 ENCFF613YON 749 bp overlap
ChIP GM23338 ENCFF886DXX 749 bp overlap
ChIP H1 ENCFF232NZA 749 bp overlap
ChIP HUVEC-C GSE109625.EZH2.HUVEC-C 749 bp overlap
ChIP HUVEC-C_VEGF_12h GSE109625.EZH2.HUVEC-C_VEGF_12h 720 bp overlap
ChIP HUVEC-C_VEGF_1h GSE109625.EZH2.HUVEC-C_VEGF_1h 88 bp overlap
ChIP HUVEC-C_VEGF_1h GSE109625.EZH2.HUVEC-C_VEGF_1h 537 bp overlap
ChIP HUVEC-C_VEGF_4h GSE109625.EZH2.HUVEC-C_VEGF_4h 749 bp overlap
ChIP MM.1S ENCFF138BXR 439 bp overlap
ChIP RCH-ACV GSE135024.EZH2.RCH-ACV 190 bp overlap
ChIP SU-DHL-6 GSE134136.EZH2.SU-DHL-6 75 bp overlap
ChIP astrocyte ENCFF365JTP 705 bp overlap
ChIP endothelial cell of umbilical vein ENCFF539AKL 749 bp overlap
ChIP fibroblast of lung ENCFF479BAW 102 bp overlap
ChIP fibroblast of lung ENCFF479BAW 412 bp overlap
ChIP hESC GSE113817.EZH2.hESC 749 bp overlap
ChIP hESC_KO GSE113817.EZH2.hESC_KO 72 bp overlap
ChIP hESC_KO GSE113817.EZH2.hESC_KO 368 bp overlap
ChIP hESC_KO GSE113817.EZH2.hESC_KO 61 bp overlap
ChIP hepatocyte ENCFF118DKH 357 bp overlap
ChIP hepatocyte ENCFF552DZB 749 bp overlap
ChIP hiPSC_WTa_RNase-neg GSE128135.EZH2.hiPSC_WTa_RNase-neg 749 bp overlap
ChIP neural cell ENCFF610EPB 122 bp overlap
ChIP neural cell ENCFF610EPB 129 bp overlap
ChIP neural progenitor cell ENCFF018MKA 749 bp overlap
ChIP neural progenitor cell ENCFF472NFV 749 bp overlap
ChIP peripheral-blood-mononuclear-cell_B-cell GSE115772.EZH2.peripheral-blood-mononuclear-cell_B-cell 374 bp overlap
FOXA1 1 dataset
ChIP breast-cancer_heregulin GSE101407.FOXA1.breast-cancer_heregulin 521 bp overlap
FOXA2 1 dataset
ChIP BJ1-hTERT_FoxHnf1aCoExp GSE90454.FOXA2.BJ1-hTERT_FoxHnf1aCoExp 308 bp overlap
GABPA 3 datasets
ChIP H1 ENCFF739QFD 341 bp overlap
ChIP WA01 ENCSR000BIW.GABPA.WA01 159 bp overlap
ChIP WA01 ENCSR000BIW.GABPA.WA01 233 bp overlap
GATA2 1 dataset
ChIP ME-1_CBFB-MYH11-KD GSE117138.GATA2.ME-1_CBFB-MYH11-KD 242 bp overlap
GATA6 2 datasets
ChIP DE_D1 S41-DE-d1-GATA6-exp2 365 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 365 bp overlap
HEXIM1 1 dataset
ChIP HCT-116 GSE72622.HEXIM1.HCT-116 393 bp overlap
HIF1A 1 dataset
ChIP BEAS-2B_arsenic GSE145834.HIF1A.BEAS-2B_arsenic 240 bp overlap
IKZF3 2 datasets
ChIP HEK293 ENCSR304AMN.IKZF3.HEK293 163 bp overlap
ChIP HEK293 ENCSR304AMN.IKZF3.HEK293 110 bp overlap
JARID2 1 dataset
ChIP AMIPS8 GSE48516.JARID2.AMIPS8 749 bp overlap
JUN 3 datasets
ChIP ESC S24-ESC-d0-JUN-exp1 281 bp overlap
ChIP HUES-8 GSE109524.JUN.HUES-8 396 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.JUN.HUES-8_DE_ctrl 346 bp overlap
KDM1A 2 datasets
ChIP pancreatic-progenitor_PP1 GSE104840.KDM1A.pancreatic-progenitor_PP1 324 bp overlap
ChIP pancreatic-progenitor_PP2-KDM1A-inh GSE104840.KDM1A.pancreatic-progenitor_PP2-KDM1A-inh 273 bp overlap
KDM4A 1 dataset
ChIP hiPSC_IID12 GSE106870.KDM4A.hiPSC_IID12 175 bp overlap
KDM5B 1 dataset
ChIP SUM185_SHCTCF GSE46055.KDM5B.SUM185_SHCTCF 66 bp overlap
KDM6B 1 dataset
ChIP CUTLL1 GSE56694.KDM6B.CUTLL1 231 bp overlap
KLF5 2 datasets
ChIP ESO-26 GSE132680.KLF5.ESO-26 202 bp overlap
ChIP KATO-III GSE51705.KLF5.KATO-III 255 bp overlap
MBD2 1 dataset
ChIP HeLa GSE41006.MBD2.HeLa 159 bp overlap
MYC 2 datasets
ChIP CD34 GSE85488.MYC.CD34 123 bp overlap
ChIP P493-6 GSE77061.MYC.P493-6 150 bp overlap
MYOD1 1 dataset
ChIP IMR-90_GM GSE128527.MYOD1.IMR-90_GM 256 bp overlap
NANOG 2 datasets
ChIP WA01 ERP004238.NANOG.WA01 354 bp overlap
ChIP WA01_3IL ERP004238.NANOG.WA01_3IL 532 bp overlap
NCAPH2 1 dataset
ChIP HEK293 GSE97540.NCAPH2.HEK293 552 bp overlap
NCOR2 1 dataset
ChIP LS180 GSE39277.NCOR2.LS180 101 bp overlap
NOTCH3 1 dataset
ChIP TALL-1_DMSO GSE104261.NOTCH3.TALL-1_DMSO 179 bp overlap
NR2F2 2 datasets
ChIP MCF-7_E2 GSE132432.NR2F2.MCF-7_E2 316 bp overlap
ChIP MCF-7_Veh GSE132432.NR2F2.MCF-7_Veh 749 bp overlap
NUTM1 2 datasets
ChIP NUT_MZ1 GSE133122.NUTM1.NUT_MZ1 749 bp overlap
ChIP embryonic-kidney_induced GSE133122.NUTM1.embryonic-kidney_induced 257 bp overlap
OGG1 8 datasets
ChIP HEK293 GSE89017.OGG1.HEK293 182 bp overlap
ChIP HEK293 GSE89017.OGG1.HEK293 345 bp overlap
ChIP HEK293_15min GSE89017.OGG1.HEK293_15min 111 bp overlap
ChIP HEK293_15min GSE89017.OGG1.HEK293_15min 431 bp overlap
ChIP HEK293_30_min GSE89017.OGG1.HEK293_30_min 67 bp overlap
ChIP HEK293_30_min GSE89017.OGG1.HEK293_30_min 383 bp overlap
ChIP HEK293_60_min GSE89017.OGG1.HEK293_60_min 159 bp overlap
ChIP HEK293_60_min GSE89017.OGG1.HEK293_60_min 387 bp overlap
OLIG2 1 dataset
ChIP brain-prefrontal-cortex_2018006 GSE129039.OLIG2.brain-prefrontal-cortex_2018006 351 bp overlap
ONECUT1 1 dataset
ChIP pancreatic-progenitor_PP1 GSE104840.ONECUT1.pancreatic-progenitor_PP1 401 bp overlap
PCGF2 3 datasets
ChIP NT2-D1 GSE101538.PCGF2.NT2-D1 514 bp overlap
ChIP fibroblast_MET GSE55605.PCGF2.fibroblast_MET 369 bp overlap
ChIP fibroblast_MET GSE55605.PCGF2.fibroblast_MET 117 bp overlap
PDX1 1 dataset
ChIP pancreatic-progenitor GSE117136.PDX1.pancreatic-progenitor 400 bp overlap
POLR2A 1 dataset
ChIP sigmoid colon ENCFF748YVT 401 bp overlap
POU2F1 2 datasets
ChIP T-47D GSE148277.POU2F1.T-47D 305 bp overlap
ChIP T-47D GSE148277.POU2F1.T-47D 197 bp overlap
POU5F1 7 datasets
ChIP BG03 GSE21614.POU5F1.BG03 128 bp overlap
ChIP BG03 GSE21614.POU5F1.BG03 278 bp overlap
ChIP BJ1-hTERT GSE92491.POU5F1.BJ1-hTERT 167 bp overlap
ChIP DE_D1 DED1-OCT4_Batch_II 749 bp overlap
ChIP NCCIT_SNF5 GSE36134.POU5F1.NCCIT_SNF5 513 bp overlap
ChIP WA01_3IL ERP004238.POU5F1.WA01_3IL 749 bp overlap
ChIP hESC_NAIVE GSE69646.POU5F1.hESC_NAIVE 352 bp overlap
POU5F1_M 1 dataset
ChIP DE_D1 DED1-OCT4-M_Batch_II 749 bp overlap
PRDM14 1 dataset
ChIP NCCIT GSE71675.PRDM14.NCCIT 558 bp overlap
RARA 2 datasets
ChIP TSU-1621MT GSE60477.RARA.TSU-1621MT 223 bp overlap
ChIP hiPSC_D4 GSE109172.RARA.hiPSC_D4 261 bp overlap
RELA 1 dataset
ChIP HEK293_TNF-1h GSE75562.RELA.HEK293_TNF-1h 298 bp overlap
RNF2 4 datasets
ChIP H1 ENCFF239FFS 749 bp overlap
ChIP H1 ENCFF239FFS 749 bp overlap
ChIP HUES-64 GSE104059.RNF2.HUES-64 221 bp overlap
ChIP NCCIT GSE71675.RNF2.NCCIT 308 bp overlap
RORC 2 datasets
ChIP HCC70 GSE126380.RORC.HCC70 749 bp overlap
ChIP HCC70_XY018 GSE126380.RORC.HCC70_XY018 749 bp overlap
RUNX1 2 datasets
ChIP AML_age-50-70 GSE111821.RUNX1.AML_age-50-70 446 bp overlap
ChIP SKNO-1 GSE23730.RUNX1.SKNO-1 200 bp overlap
RUNX1T1 1 dataset
ChIP Kasumi-1_E2 GSE102697.RUNX1T1.Kasumi-1_E2 153 bp overlap
RXR 1 dataset
ChIP TSU-1621MT GSE60477.RXR.TSU-1621MT 391 bp overlap
SALL3 1 dataset
ChIP hiPSC GSE104863.SALL3.hiPSC 256 bp overlap
SIRT6 1 dataset
ChIP SK-MEL-239_L-C-B GSE102813.SIRT6.SK-MEL-239_L-C-B 159 bp overlap
SMAD2 1 dataset
ChIP hESC_YAP-_activinA_15h GSE99202.SMAD2.hESC_YAP-_activinA_15h 299 bp overlap
SMAD2-3 3 datasets
ChIP HUES-8 GSE109524.SMAD2-3.HUES-8 584 bp overlap
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 481 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.SMAD2-3.HUES-8_DE_JNKi 749 bp overlap
SMAD2_3 6 datasets
ChIP DE_D1 S07-DE-d1-SMAD2_3-exp1 415 bp overlap
ChIP DE_D1 S12-DE-d1-SMAD2_3-exp1 57 bp overlap
ChIP DE_D1 S12-DE-d1-SMAD2_3-exp1 496 bp overlap
ChIP DE_D1 S39-DE-d1-SMAD2_3-exp2 651 bp overlap
ChIP DE_D2 S01-DE-d2-SMAD2_3-exp1 261 bp overlap
ChIP ESC S33-ESC-d0-SMAD2_3-exp2 389 bp overlap
SMARCA4 3 datasets
ChIP 22Rv1 GSE115615.SMARCA4.22Rv1 292 bp overlap
ChIP 22Rv1_PTEN-KO GSE115615.SMARCA4.22Rv1_PTEN-KO 141 bp overlap
ChIP 22Rv1_PTEN-KO GSE115615.SMARCA4.22Rv1_PTEN-KO 345 bp overlap
SREBP2 2 datasets
ChIP HCC70 GSE126380.SREBP2.HCC70 729 bp overlap
ChIP HCC70_XY018 GSE126380.SREBP2.HCC70_XY018 723 bp overlap
STAT3 1 dataset
ChIP WA01 ERP004237.STAT3.WA01 478 bp overlap
SUZ12 11 datasets
ChIP GM12878 ENCSR744XTG.SUZ12.GM12878 749 bp overlap
ChIP GM12878 ENCSR091BOQ.SUZ12.GM12878 383 bp overlap
ChIP GM12878 ENCSR091BOQ.SUZ12.GM12878 94 bp overlap
ChIP H1 ENCFF507HGF 56 bp overlap
ChIP H1 ENCFF507HGF 232 bp overlap
ChIP H1 ENCFF881NFR 749 bp overlap
ChIP K-562 ENCSR000AUC.SUZ12.K-562 749 bp overlap
ChIP NT2-D1 GSE101538.SUZ12.NT2-D1 749 bp overlap
ChIP NT2/D1 ENCFF574SXS 749 bp overlap
ChIP WA01 ENCSR000EUQ.SUZ12.WA01 749 bp overlap
ChIP hiPSC_WTb_RNase-neg GSE128135.SUZ12.hiPSC_WTb_RNase-neg 749 bp overlap
TCF12 1 dataset
ChIP WA01 ENCSR000BIT.TCF12.WA01 138 bp overlap
TFAP2C 1 dataset
ChIP UCLA1-hESCs_TFAP2C--_1d_TFAP2C_Induc GSE101074.TFAP2C.UCLA1-hESCs_TFAP2C--_1d_TFAP2C_Induc 447 bp overlap
TFIIIC 2 datasets
ChIP HEK293 GSE119418.TFIIIC.HEK293 76 bp overlap
ChIP HEK293 GSE119418.TFIIIC.HEK293 487 bp overlap
TP63 1 dataset
ChIP keratinocyte_diff GSE33571.TP63.keratinocyte_diff 284 bp overlap
TRIM25 1 dataset
ChIP BT-549 GSE79588.TRIM25.BT-549 470 bp overlap
WDR5 1 dataset
ChIP SMMC-7721 GSE126982.WDR5.SMMC-7721 715 bp overlap
YY1 1 dataset
ChIP WA01 ENCSR000BKD.YY1.WA01 310 bp overlap
ZBTB48 2 datasets
ChIP U2OS GSE96776.ZBTB48.U2OS 152 bp overlap
ChIP U2OS_ZBTB48-KO GSE96776.ZBTB48.U2OS_ZBTB48-KO 749 bp overlap
ZBTB6 2 datasets
ChIP HEK293 ENCSR619OUC.ZBTB6.HEK293 103 bp overlap
ChIP HEK293 GSE76494.ZBTB6.HEK293 159 bp overlap
ZBTB7A 2 datasets
ChIP VCaP_CSS GSE123089.ZBTB7A.VCaP_CSS 247 bp overlap
ChIP VCaP_FBS GSE123089.ZBTB7A.VCaP_FBS 58 bp overlap
ZFX 2 datasets
ChIP HEK293T_22Rv1 GSE145160.ZFX.HEK293T_22Rv1 488 bp overlap
ChIP HEK293T_FLAG GSE145160.ZFX.HEK293T_FLAG 547 bp overlap
ZNF445 1 dataset
ChIP HEK293T GSE78099.ZNF445.HEK293T 663 bp overlap
ZNF468 1 dataset
ChIP HEK293T GSE78099.ZNF468.HEK293T 283 bp overlap
ZNF711 1 dataset
ChIP HEK293T GSE145160.ZNF711.HEK293T 749 bp overlap