chr11 : 128,174,626 – 128,174,981
355 bp 76 TFs 0 linked genes
This 355 bp open chromatin element has no linked target genes and is bound by 76 transcription factors.
Linked Genes
No linked genes
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr11:128,169,626 – 128,179,981
This element Other elements ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
76 transcription factors
Source
Cell type
BCOR 2 datasets
ChIP WA01_RNF2-R GSE104690.BCOR.WA01_RNF2-R 256 bp overlap
ChIP WA01_RNF2-R_Doxycyclin GSE104690.BCOR.WA01_RNF2-R_Doxycyclin 163 bp overlap
BRD2 5 datasets
ChIP MDA-MB-231_BAZ2BsiRNA GSE116879.BRD2.MDA-MB-231_BAZ2BsiRNA 251 bp overlap
ChIP MDA-MB-231_MGSK2801 GSE116879.BRD2.MDA-MB-231_MGSK2801 346 bp overlap
ChIP MDA-MB-231_MGSK2801_rDNA GSE116879.BRD2.MDA-MB-231_MGSK2801_rDNA 346 bp overlap
ChIP MDA-MB-231_NTsiRNA GSE116879.BRD2.MDA-MB-231_NTsiRNA 61 bp overlap
ChIP SK-MEL-147 GSE94488.BRD2.SK-MEL-147 186 bp overlap
BRD4 21 datasets
ChIP HCC1395 GSE63581.BRD4.HCC1395 210 bp overlap
ChIP Hs-352-Sk_PAX3-FOXO1-vector GSE83725.BRD4.Hs-352-Sk_PAX3-FOXO1-vector 250 bp overlap
ChIP IMR-90_QUIES GSE74238.BRD4.IMR-90_QUIES 304 bp overlap
ChIP MDA-MB-231_DMSO GSE116879.BRD4.MDA-MB-231_DMSO 355 bp overlap
ChIP MDA-MB-231_DMSO_rDNA GSE116879.BRD4.MDA-MB-231_DMSO_rDNA 355 bp overlap
ChIP MDA-MB-231_JQ1 GSE116879.BRD4.MDA-MB-231_JQ1 163 bp overlap
ChIP MDA-MB-231_JQ1-neg_L GSE136151.BRD4.MDA-MB-231_JQ1-neg_L 226 bp overlap
ChIP MDA-MB-231_JQ1-neg_S_3f GSE136151.BRD4.MDA-MB-231_JQ1-neg_S_3f 355 bp overlap
ChIP MDA-MB-231_JQ1_MGSK2801 GSE116879.BRD4.MDA-MB-231_JQ1_MGSK2801 300 bp overlap
ChIP MDA-MB-231_JQ1_MGSK2801_rDNA GSE116879.BRD4.MDA-MB-231_JQ1_MGSK2801_rDNA 300 bp overlap
ChIP MDA-MB-231_JQ1_rDNA GSE116879.BRD4.MDA-MB-231_JQ1_rDNA 163 bp overlap
ChIP MDA-MB-231_MGSK2801 GSE116879.BRD4.MDA-MB-231_MGSK2801 355 bp overlap
ChIP MDA-MB-231_MGSK2801_rDNA GSE116879.BRD4.MDA-MB-231_MGSK2801_rDNA 355 bp overlap
ChIP MPNST_PDJQ GSE62499.BRD4.MPNST_PDJQ 97 bp overlap
ChIP RH4 GSE83726.BRD4.RH4 296 bp overlap
ChIP RH4_shCHD4 GSE140115.BRD4.RH4_shCHD4 194 bp overlap
ChIP SUM1315_DMSO GSE63581.BRD4.SUM1315_DMSO 355 bp overlap
ChIP SUM159PT_30nMbortezomib_8h GSE87418.BRD4.SUM159PT_30nMbortezomib_8h 165 bp overlap
ChIP SUM159PT_DMSO_24h GSE87418.BRD4.SUM159PT_DMSO_24h 355 bp overlap
ChIP SUM159PT_KO GSE131097.BRD4.SUM159PT_KO 264 bp overlap
ChIP SUM159PT_scramble GSE131097.BRD4.SUM159PT_scramble 355 bp overlap
BRD9 2 datasets
ChIP MDA-MB-231_DMSO GSE116879.BRD9.MDA-MB-231_DMSO 355 bp overlap
ChIP MDA-MB-231_MGSK2801 GSE116879.BRD9.MDA-MB-231_MGSK2801 172 bp overlap
CASZ1 1 dataset
ChIP rhabdomyosarcoma_Trametinib GSE126143.CASZ1.rhabdomyosarcoma_Trametinib 215 bp overlap
CDKN1B 1 dataset
ChIP MDA-BoM-1833_shp27 GSE112444.CDKN1B.MDA-BoM-1833_shp27 221 bp overlap
CEBPB 6 datasets
ChIP A-549 ENCSR000BUB.CEBPB.A-549 221 bp overlap
ChIP H1 ENCFF871PTR 180 bp overlap
ChIP HFOB_DIFF GSE82295.CEBPB.HFOB_DIFF 211 bp overlap
ChIP IMR-90 ENCFF468UGY 162 bp overlap
ChIP WA01 ENCSR000EBV.CEBPB.WA01 149 bp overlap
ChIP hMSC GSE68864.CEBPB.hMSC 121 bp overlap
CHD4 1 dataset
ChIP SCMC GSE155861.CHD4.SCMC 325 bp overlap
CHD7 3 datasets
ChIP hiPSC_AF22_abD3F5 GSE108506.CHD7.hiPSC_AF22_abD3F5 257 bp overlap
ChIP hiPSC_derived_lt-NES GSE108506.CHD7.hiPSC_derived_lt-NES 327 bp overlap
ChIP hiPSC_derived_neural-crest-cell GSE108506.CHD7.hiPSC_derived_neural-crest-cell 82 bp overlap
CTCF 26 datasets
ChIP A-549 ENCSR000AUF.CTCF.A-549 133 bp overlap
ChIP A-673 ENCSR611JJS.CTCF.A-673 217 bp overlap
ChIP AG04449 ENCFF248MBD 157 bp overlap
ChIP D54 ENCSR000DKN.CTCF.D54 182 bp overlap
ChIP GM23338 ENCFF772DML 151 bp overlap
ChIP GSC23 GSE139416.CTCF.GSC23 202 bp overlap
ChIP H54 ENCFF255TVO 171 bp overlap
ChIP NB69 GSE101295.CTCF.NB69 109 bp overlap
ChIP OVCAR-8 GSE70764.CTCF.OVCAR-8 196 bp overlap
ChIP PC-3 ENCSR359LOD.CTCF.PC-3 163 bp overlap
ChIP RH4 GSE83726.CTCF.RH4 274 bp overlap
ChIP SH-SY5Y GSE101295.CTCF.SH-SY5Y 132 bp overlap
ChIP T-47D GSE111923.CTCF.T-47D 135 bp overlap
ChIP WERI-Rb-1 ENCFF349QKF 162 bp overlap
ChIP WERI-Rb-1 ENCSR000DXW.CTCF.WERI-Rb-1 143 bp overlap
ChIP brain microvascular endothelial cell ENCFF526VHE 193 bp overlap
ChIP chondrocyte ENCFF134ORZ 298 bp overlap
ChIP choroid-plexus_epithelial ENCSR000DTL.CTCF.choroid-plexus_epithelial 115 bp overlap
ChIP endothelial_brain-microvascular ENCSR000DTA.CTCF.endothelial_brain-microvascular 169 bp overlap
ChIP fibroblast_SKIN_LEG ENCSR000DPG.CTCF.fibroblast_SKIN_LEG 107 bp overlap
ChIP myoblast_skeletal_muscle ENCSR000ANE.CTCF.myoblast_skeletal_muscle 141 bp overlap
ChIP myotube ENCSR000ANS.CTCF.myotube 140 bp overlap
ChIP neuron_bipolar_doxy_4d ENCSR619IUE.CTCF.neuron_bipolar_doxy_4d 127 bp overlap
ChIP osteoblast ENCSR000APF.CTCF.osteoblast 156 bp overlap
ChIP osteocyte ENCFF929FPD 93 bp overlap
ChIP skeletal muscle myoblast ENCFF813BQI 199 bp overlap
CTCFL 1 dataset
ChIP OVCAR-8 GSE70764.CTCFL.OVCAR-8 209 bp overlap
DEAF1 1 dataset
ChIP keratinocyte GSE129965.DEAF1.keratinocyte 158 bp overlap
E2F1 1 dataset
ChIP U-87MG_GBM GSE99171.E2F1.U-87MG_GBM 96 bp overlap
EP300 2 datasets
ChIP SK-N-SH ENCFF451CNG 156 bp overlap
ChIP SK-N-SH ENCSR000BUA.EP300.SK-N-SH 112 bp overlap
ETS1 1 dataset
ChIP hESC ENCSR534VHI.ETS1.hESC 195 bp overlap
EZH2 2 datasets
ChIP GM23248 ENCFF506FWX 165 bp overlap
ChIP RH30_DMSO GSE85169.EZH2.RH30_DMSO 293 bp overlap
FOSL1 1 dataset
ChIP BT-549 GSE112961.FOSL1.BT-549 119 bp overlap
FOSL2 3 datasets
ChIP A-549 ENCSR448TVS.FOSL2.A-549 355 bp overlap
ChIP NPC_K755R-pos GSE122631.FOSL2.NPC_K755R-pos 355 bp overlap
ChIP NPC_R1159Q GSE122631.FOSL2.NPC_R1159Q 302 bp overlap
FOXO1-PAX3 2 datasets
ChIP RH4_DMSO-6H GSE116344.FOXO1-PAX3.RH4_DMSO-6H 320 bp overlap
ChIP RH4_Entinostat-6H GSE116344.FOXO1-PAX3.RH4_Entinostat-6H 278 bp overlap
GATA2 3 datasets
ChIP dermal-fibroblast_alone GSE51025.GATA2.dermal-fibroblast_alone 64 bp overlap
ChIP primary-endometrial-stromal-cell_IVD_P1 GSE145673.GATA2.primary-endometrial-stromal-cell_IVD_P1 71 bp overlap
ChIP primary-endometrial-stromal-cell_Veh_P1 GSE145673.GATA2.primary-endometrial-stromal-cell_Veh_P1 81 bp overlap
GATA6 5 datasets
ChIP DE_D1 S09-DE-d1-GATA6-exp1 164 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 128 bp overlap
ChIP DE_D1 S41-DE-d1-GATA6-exp2 239 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 188 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 201 bp overlap
HDAC2 2 datasets
ChIP RH4_DMSO-6H GSE116344.HDAC2.RH4_DMSO-6H 299 bp overlap
ChIP RH4_Entinostat-6H GSE116344.HDAC2.RH4_Entinostat-6H 236 bp overlap
HDAC3 1 dataset
ChIP RH4_DMSO-6H GSE116344.HDAC3.RH4_DMSO-6H 280 bp overlap
ISL1 1 dataset
ChIP SK-N-SH ENCFF285GEQ 198 bp overlap
JUN 6 datasets
ChIP DE_D1 S08-DE-d1-JUN-exp1 355 bp overlap
ChIP DE_D1 S40-DE-d1-JUN-exp2 355 bp overlap
ChIP DE_D2 S02-DE-d2-JUN-exp1 355 bp overlap
ChIP ESC S34-ESC-d0-JUN-exp2 147 bp overlap
ChIP HUES-8 GSE109524.JUN.HUES-8 336 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.JUN.HUES-8_DE_ctrl 355 bp overlap
JUND 1 dataset
ChIP SK-N-SH ENCFF551NEQ 101 bp overlap
KDM1A 1 dataset
ChIP NCI-H526_DMSO GSE66297.KDM1A.NCI-H526_DMSO 84 bp overlap
MED1 11 datasets
ChIP U-87MG GSE36354.MED1.U-87MG 220 bp overlap
ChIP hMSC-TERT4_4h GSE104537.MED1.hMSC-TERT4_4h 270 bp overlap
ChIP hMSC-TERT4_D0 GSE104537.MED1.hMSC-TERT4_D0 244 bp overlap
ChIP hMSC-TERT4_D14 GSE104537.MED1.hMSC-TERT4_D14 143 bp overlap
ChIP hMSC-TERT4_D3 GSE104537.MED1.hMSC-TERT4_D3 245 bp overlap
ChIP hMSC-TERT4_adipocyte-4H GSE113253.MED1.hMSC-TERT4_adipocyte-4H 191 bp overlap
ChIP hMSC-TERT4_osteoblast-4H GSE113253.MED1.hMSC-TERT4_osteoblast-4H 174 bp overlap
ChIP hMSC-TERT4_osteoblast-D1 GSE113253.MED1.hMSC-TERT4_osteoblast-D1 190 bp overlap
ChIP hMSC-TERT4_osteoblast-D3 GSE113253.MED1.hMSC-TERT4_osteoblast-D3 214 bp overlap
ChIP hMSC-TERT4_osteoblast-D7 GSE113253.MED1.hMSC-TERT4_osteoblast-D7 355 bp overlap
ChIP hMSC-TERT4_undifferentiated GSE113253.MED1.hMSC-TERT4_undifferentiated 262 bp overlap
MRTFB 1 dataset
ChIP A-673-clone-Asp114 GSE92738.MRTFB.A-673-clone-Asp114 355 bp overlap
MYC 1 dataset
ChIP GEN2-2 GSE70275.MYC.GEN2-2 113 bp overlap
MYCN 1 dataset
ChIP RH4 GSE83726.MYCN.RH4 278 bp overlap
MYF5 1 dataset
ChIP Rh18 GSE84628.MYF5.Rh18 253 bp overlap
MYOD1 7 datasets
ChIP RD GSE137168.MYOD1.RD 325 bp overlap
ChIP RD_shSNAI2 GSE137168.MYOD1.RD_shSNAI2 307 bp overlap
ChIP RH30_DMSO GSE85169.MYOD1.RH30_DMSO 313 bp overlap
ChIP RH4 GSE83726.MYOD1.RH4 255 bp overlap
ChIP SMS-CTR GSE137168.MYOD1.SMS-CTR 275 bp overlap
ChIP myoblast GSE50413.MYOD1.myoblast 311 bp overlap
ChIP rhabdomyosarcoma GSE50413.MYOD1.rhabdomyosarcoma 289 bp overlap
MYOG 3 datasets
ChIP RH4 GSE83726.MYOG.RH4 206 bp overlap
ChIP RH4_DMSO-6H GSE116344.MYOG.RH4_DMSO-6H 218 bp overlap
ChIP RH4_Entinostat-6H GSE116344.MYOG.RH4_Entinostat-6H 210 bp overlap
NANOG 5 datasets
ChIP GM23338 ENCFF065NZG 235 bp overlap
ChIP GM23338 ENCSR061DGF.NANOG.GM23338 167 bp overlap
ChIP HUES-8 GSE109524.NANOG.HUES-8 355 bp overlap
ChIP WA01 ERP004238.NANOG.WA01 216 bp overlap
ChIP WA09 GSE105028.NANOG.WA09 177 bp overlap
NEUROG2 6 datasets
ChIP MRC-5_NFD_02DPT GSE75910.NEUROG2.MRC-5_NFD_02DPT 156 bp overlap
ChIP MRC-5_NFD_05DPT GSE75910.NEUROG2.MRC-5_NFD_05DPT 235 bp overlap
ChIP MRC-5_NFD_1DPT GSE75910.NEUROG2.MRC-5_NFD_1DPT 168 bp overlap
ChIP MRC-5_N_02DPT GSE75910.NEUROG2.MRC-5_N_02DPT 196 bp overlap
ChIP MRC-5_N_05DPT GSE75910.NEUROG2.MRC-5_N_05DPT 165 bp overlap
ChIP MRC-5_N_1DPT GSE75910.NEUROG2.MRC-5_N_1DPT 179 bp overlap
NFIC 2 datasets
ChIP SK-N-SH ENCFF965AKM 222 bp overlap
ChIP SK-N-SH ENCSR000BSV.NFIC.SK-N-SH 115 bp overlap
NR2F2 2 datasets
ChIP liver ENCSR168SMX.NR2F2.liver 202 bp overlap
ChIP liver ENCSR338MMB.NR2F2.liver 224 bp overlap
NR3C1 2 datasets
ChIP BEAS-2B_TNF_IA1 GSE125623.NR3C1.BEAS-2B_TNF_IA1 270 bp overlap
ChIP BEAS-2B_shNR3C1_TNF_IA1 GSE125623.NR3C1.BEAS-2B_shNR3C1_TNF_IA1 91 bp overlap
PAX3-FOXO1 2 datasets
ChIP Hs-352-Sk_PAX3-FOXO1-vector GSE83725.PAX3-FOXO1.Hs-352-Sk_PAX3-FOXO1-vector 241 bp overlap
ChIP RH4 GSE140115.PAX3-FOXO1.RH4 187 bp overlap
PBX3 1 dataset
ChIP SK-N-SH ENCFF876BMC 155 bp overlap
POLR2A 2 datasets
ChIP SK-N-SH ENCFF683PFH 199 bp overlap
ChIP spleen ENCFF044PYR 195 bp overlap
POU2F3 1 dataset
ChIP NCI-H1048 GSE115123.POU2F3.NCI-H1048 206 bp overlap
POU4F2 1 dataset
ChIP HNPC_DIF GSE74814.POU4F2.HNPC_DIF 199 bp overlap
POU5F1 1 dataset
ChIP HUES-8 GSE109524.POU5F1.HUES-8 355 bp overlap
RAD21 7 datasets
ChIP A-549 ENCSR000BUC.RAD21.A-549 89 bp overlap
ChIP HEK293_siE1A GSE130135.RAD21.HEK293_siE1A 355 bp overlap
ChIP MCF-7 ENCSR703TNG.RAD21.MCF-7 115 bp overlap
ChIP RH4 GSE83726.RAD21.RH4 265 bp overlap
ChIP SLK_CTCF-KD GSE138105.RAD21.SLK_CTCF-KD 244 bp overlap
ChIP SLK_Ctrl GSE138105.RAD21.SLK_Ctrl 94 bp overlap
ChIP T-47D_triptolide GSE111923.RAD21.T-47D_triptolide 254 bp overlap
RBBP4 2 datasets
ChIP RH5 GSE155861.RBBP4.RH5 355 bp overlap
ChIP SCMC GSE155861.RBBP4.SCMC 296 bp overlap
RELA 2 datasets
ChIP FaDu_LPS GSE132018.RELA.FaDu_LPS 355 bp overlap
ChIP SGBS_TNF GSE64233.RELA.SGBS_TNF 193 bp overlap
RNF2 1 dataset
ChIP fibroblast GSE139053.RNF2.fibroblast 78 bp overlap
SMAD2-3 1 dataset
ChIP HGrC1_EV GSE138496.SMAD2-3.HGrC1_EV 113 bp overlap
SMAD3 1 dataset
ChIP BG03 GSE21614.SMAD3.BG03 210 bp overlap
SMARCA2 4 datasets
ChIP NPC_K755R-pos GSE122631.SMARCA2.NPC_K755R-pos 213 bp overlap
ChIP NPC_K755R-siCT GSE122631.SMARCA2.NPC_K755R-siCT 154 bp overlap
ChIP NPC_R1159Q GSE122631.SMARCA2.NPC_R1159Q 231 bp overlap
ChIP SK-N-MC_shEWSFLI1 GSE94275.SMARCA2.SK-N-MC_shEWSFLI1 115 bp overlap
SMARCA4 9 datasets
ChIP A-549_AG15722 GSE132290.SMARCA4.A-549_AG15722 87 bp overlap
ChIP A-549_AG15723 GSE132290.SMARCA4.A-549_AG15723 63 bp overlap
ChIP NPC_K755R-siAP GSE122631.SMARCA4.NPC_K755R-siAP 178 bp overlap
ChIP NPC_K755R-siCT GSE122631.SMARCA4.NPC_K755R-siCT 355 bp overlap
ChIP NPC_siCT GSE122631.SMARCA4.NPC_siCT 206 bp overlap
ChIP NPC_siSOX GSE122631.SMARCA4.NPC_siSOX 355 bp overlap
ChIP NSC GSE125033.SMARCA4.NSC 227 bp overlap
ChIP hiPSC GSE124903.SMARCA4.hiPSC 183 bp overlap
ChIP hiPSC_WT GSE124903.SMARCA4.hiPSC_WT 287 bp overlap
SMARCB1 2 datasets
ChIP hiPSC GSE124903.SMARCB1.hiPSC 176 bp overlap
ChIP proliferating-human-fibroblast_ASO_LINC GSE131711.SMARCB1.proliferating-human-fibroblast_ASO_LINC 276 bp overlap
SMARCC1 1 dataset
ChIP hiPSC_WT GSE124903.SMARCC1.hiPSC_WT 302 bp overlap
SMC1 1 dataset
ChIP DKO GSE131606.SMC1.DKO 232 bp overlap
SMC1A 1 dataset
ChIP A-549 GSE76893.SMC1A.A-549 119 bp overlap
SMC3 1 dataset
ChIP IMR-90 ENCSR000HPG.SMC3.IMR-90 57 bp overlap
SNAI2 1 dataset
ChIP RD GSE137168.SNAI2.RD 219 bp overlap
SOX2 1 dataset
ChIP HNSC GSE69479.SOX2.HNSC 341 bp overlap
SP1 1 dataset
ChIP WA01 ENCSR000BIR.SP1.WA01 164 bp overlap
T 1 dataset
ChIP H9_MESODERM GSE60606.T.H9_MESODERM 251 bp overlap
TAL1 1 dataset
ChIP HSPC-CD34pos GSE93372.TAL1.HSPC-CD34pos 103 bp overlap
TCF12 3 datasets
ChIP H1 ENCFF203EBH 251 bp overlap
ChIP SK-N-SH ENCFF147AHB 70 bp overlap
ChIP WA01 ENCSR000BIT.TCF12.WA01 219 bp overlap
TEAD1 3 datasets
ChIP HUCCT1 GSE68296.TEAD1.HUCCT1 320 bp overlap
ChIP WTC11 ENCFF502QUV 355 bp overlap
ChIP adipocyte GSE140782.TEAD1.adipocyte 129 bp overlap
TEAD4 6 datasets
ChIP H1 ENCFF778PAX 245 bp overlap
ChIP HUCCT1 GSE68296.TEAD4.HUCCT1 270 bp overlap
ChIP SK-MEL-147 GSE94488.TEAD4.SK-MEL-147 224 bp overlap
ChIP SK-N-SH ENCFF754TJT 107 bp overlap
ChIP SK-N-SH ENCSR000BUQ.TEAD4.SK-N-SH 95 bp overlap
ChIP hMSC-TERT4 GSE140782.TEAD4.hMSC-TERT4 258 bp overlap
TP53 1 dataset
ChIP MDA-MB-231 GSE95303.TP53.MDA-MB-231 166 bp overlap
TWIST1 4 datasets
ChIP SHEP-21N GSE80151.TWIST1.SHEP-21N 123 bp overlap
ChIP SHEP-21N_24h GSE80151.TWIST1.SHEP-21N_24h 142 bp overlap
ChIP SHEP-21N_DOX_0H GSE80151.TWIST1.SHEP-21N_DOX_0H 123 bp overlap
ChIP SHEP-21N_DOX_24H GSE80151.TWIST1.SHEP-21N_DOX_24H 229 bp overlap
VDR 1 dataset
ChIP LX2_CALCIPOTRIOL_TGFB1 GSE38103.VDR.LX2_CALCIPOTRIOL_TGFB1 239 bp overlap
YY1 1 dataset
ChIP RH4_DMSO-6H GSE116344.YY1.RH4_DMSO-6H 227 bp overlap
ZBTB33 1 dataset
ChIP SK-N-SH ENCFF667JYU 223 bp overlap
ZMYND8 1 dataset
ChIP MDA-MB-231 GSE108833.ZMYND8.MDA-MB-231 243 bp overlap
ZNF423 1 dataset
ChIP WTC11 ENCFF574PBR 224 bp overlap
ZNF462 2 datasets
ChIP GM23338 ENCFF896CCA 218 bp overlap
ChIP GM23338 ENCSR334UWP.ZNF462.GM23338 201 bp overlap