chr1 : 152,960,740 – 152,960,943
203 bp 62 TFs 0 linked genes
This 203 bp open chromatin element has no linked target genes and is bound by 62 transcription factors.
Linked Genes
No linked genes
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr1:152,955,740 – 152,965,943
This element Other elements ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
62 transcription factors
Source
Cell type
AR 3 datasets
ChIP LNCaP_D226N_shFOXA1_Ethanol GSE128883.AR.LNCaP_D226N_shFOXA1_Ethanol 159 bp overlap
ChIP LNCaP_GFP_shFOXA1_Ethanol GSE128883.AR.LNCaP_GFP_shFOXA1_Ethanol 178 bp overlap
ChIP MCF-7 GSE48930.AR.MCF-7 54 bp overlap
ARID2 1 dataset
ChIP K-562 ENCSR491EBY.ARID2.K-562 187 bp overlap
ARID3A 1 dataset
ChIP GM12878 ENCSR725LYT.ARID3A.GM12878 128 bp overlap
ATF1 1 dataset
ChIP K-562 ENCSR091GVJ.ATF1.K-562 149 bp overlap
BATF 1 dataset
ChIP GM12878 GSE97661.BATF.GM12878 95 bp overlap
BRD2 1 dataset
ChIP THP-1_iBET-BD2 GSE138084.BRD2.THP-1_iBET-BD2 116 bp overlap
CREB1 3 datasets
ChIP LNCaP GSE63034.CREB1.LNCaP 112 bp overlap
ChIP LNCaP-abl_SHCTR GSE63034.CREB1.LNCaP-abl_SHCTR 203 bp overlap
ChIP LNCaP-abl_SHFOXA1 GSE63034.CREB1.LNCaP-abl_SHFOXA1 203 bp overlap
CREM 2 datasets
ChIP K-562 ENCSR077DKV.CREM.K-562 148 bp overlap
ChIP K562 ENCFF180STA 203 bp overlap
CTCF 327 datasets
ChIP 22Rv1 ENCFF466OXN 203 bp overlap
ChIP 22Rv1 ENCFF466OXN 203 bp overlap
ChIP 22Rv1 ENCSR857PBV.CTCF.22Rv1 203 bp overlap
ChIP 22Rv1_hydroxy_10nM_4h ENCSR847XGE.CTCF.22Rv1_hydroxy_10nM_4h 203 bp overlap
ChIP 786-O_NORMOXIA GSE78113.CTCF.786-O_NORMOXIA 203 bp overlap
ChIP 81-3 ERP002246.CTCF.81-3 176 bp overlap
ChIP A-549 ENCSR000AUF.CTCF.A-549 203 bp overlap
ChIP A-549 ENCSR000DPF.CTCF.A-549 192 bp overlap
ChIP A-549 ENCSR000DNA.CTCF.A-549 171 bp overlap
ChIP A-549 ENCSR000DYD.CTCF.A-549 119 bp overlap
ChIP A-673 ENCSR611JJS.CTCF.A-673 203 bp overlap
ChIP A549 ENCFF034FVO 203 bp overlap
ChIP A549 ENCFF182TCQ 203 bp overlap
ChIP BC-3 GSE135740.CTCF.BC-3 203 bp overlap
ChIP BL41 GSE31485.CTCF.BL41 136 bp overlap
ChIP C4-2B ENCFF821XVN 203 bp overlap
ChIP C4-2B ENCFF821XVN 203 bp overlap
ChIP COLO-829 GSE81945.CTCF.COLO-829 203 bp overlap
ChIP Caco-2 ENCSR000DQN.CTCF.Caco-2 114 bp overlap
ChIP D54 ENCSR000DKN.CTCF.D54 164 bp overlap
ChIP DND-41 ENCFF913MRA 203 bp overlap
ChIP DND41 ENCSR000AQU.CTCF.DND41 202 bp overlap
ChIP GM04025_B-lymphocytes GSE111170.CTCF.GM04025_B-lymphocytes 203 bp overlap
ChIP GM04604 GSE148179.CTCF.GM04604 203 bp overlap
ChIP GM04648 GSE148179.CTCF.GM04648 203 bp overlap
ChIP GM06077 GSE148179.CTCF.GM06077 203 bp overlap
ChIP GM06990 ENCFF471OQT 76 bp overlap
ChIP GM06990 ENCSR000DQW.CTCF.GM06990 203 bp overlap
ChIP GM09236__B-lymphocytes GSE111170.CTCF.GM09236__B-lymphocytes 203 bp overlap
ChIP GM09237_B-lymphocytes GSE111170.CTCF.GM09237_B-lymphocytes 203 bp overlap
ChIP GM10266 ENCFF241YYF 177 bp overlap
ChIP GM10266 ENCFF892KUY 177 bp overlap
ChIP GM10266 ENCSR000DKR.CTCF.GM10266 144 bp overlap
ChIP GM12801 ENCSR000DQY.CTCF.GM12801 176 bp overlap
ChIP GM12864 ENCFF357DQE 203 bp overlap
ChIP GM12864 ENCSR000DRB.CTCF.GM12864 203 bp overlap
ChIP GM12865 ENCFF067GFI 203 bp overlap
ChIP GM12865 ENCSR000DRE.CTCF.GM12865 203 bp overlap
ChIP GM12866 ENCSR000DRF.CTCF.GM12866 203 bp overlap
ChIP GM12867 ENCSR000DRH.CTCF.GM12867 196 bp overlap
ChIP GM12868 ENCSR000DRI.CTCF.GM12868 203 bp overlap
ChIP GM12869 ENCSR000DRJ.CTCF.GM12869 203 bp overlap
ChIP GM12870 ENCSR000DRK.CTCF.GM12870 203 bp overlap
ChIP GM12871 ENCSR000DRL.CTCF.GM12871 203 bp overlap
ChIP GM12872 ENCFF697BYI 203 bp overlap
ChIP GM12872 ENCSR000DRN.CTCF.GM12872 202 bp overlap
ChIP GM12873 ENCFF711LOS 126 bp overlap
ChIP GM12873 ENCSR000DRP.CTCF.GM12873 203 bp overlap
ChIP GM12874 ENCFF942MTD 203 bp overlap
ChIP GM12874 ENCSR000DRR.CTCF.GM12874 189 bp overlap
ChIP GM12875 ENCFF081UCQ 203 bp overlap
ChIP GM12875 ENCSR000DRU.CTCF.GM12875 203 bp overlap
ChIP GM12878 ENCFF217EAX 203 bp overlap
ChIP GM12878 ENCFF485TGR 203 bp overlap
ChIP GM12878 ENCFF511URZ 106 bp overlap
ChIP GM12878 ENCFF635MMB 190 bp overlap
ChIP GM12878 GSE76922.CTCF.GM12878 203 bp overlap
ChIP GM12878 ENCSR000AKB.CTCF.GM12878 203 bp overlap
ChIP GM12878 ENCSR000DKV.CTCF.GM12878 203 bp overlap
ChIP GM12878 ENCSR000DRZ.CTCF.GM12878 182 bp overlap
ChIP GM12878 ENCSR000DZN.CTCF.GM12878 180 bp overlap
ChIP GM12878 ERP002246.CTCF.GM12878 136 bp overlap
ChIP GM13977 ENCFF528ESQ 171 bp overlap
ChIP GM13977 ENCSR000DLB.CTCF.GM13977 155 bp overlap
ChIP GM17942 GSE76922.CTCF.GM17942 203 bp overlap
ChIP GM23338 ENCFF531QOI 203 bp overlap
ChIP GM23338 ENCFF772DML 190 bp overlap
ChIP GP5D GSE51234.CTCF.GP5D 203 bp overlap
ChIP H1 ENCFF230QSV 165 bp overlap
ChIP H1 ENCFF414GZI 104 bp overlap
ChIP H1 ENCFF764RHO 177 bp overlap
ChIP H54 ENCFF255TVO 202 bp overlap
ChIP H9 ENCFF152GTF 203 bp overlap
ChIP HAP1 GSE152721.CTCF.HAP1 171 bp overlap
ChIP HAP1_clone21 GSE152721.CTCF.HAP1_clone21 184 bp overlap
ChIP HAP1_clone21_TSS-CRISPR GSE152721.CTCF.HAP1_clone21_TSS-CRISPR 177 bp overlap
ChIP HCT-116 GSE131606.CTCF.HCT-116 203 bp overlap
ChIP HCT-116 ENCSR240PRQ.CTCF.HCT-116 203 bp overlap
ChIP HCT-116 ENCSR000DTO.CTCF.HCT-116 198 bp overlap
ChIP HCT-116 GSE92879.CTCF.HCT-116 193 bp overlap
ChIP HCT-116 ENCSR000BSE.CTCF.HCT-116 187 bp overlap
ChIP HCT-116_RAD21-mAC GSE104888.CTCF.HCT-116_RAD21-mAC 203 bp overlap
ChIP HCT-116_RAD21-mAC_500uM_auxin GSE104888.CTCF.HCT-116_RAD21-mAC_500uM_auxin 172 bp overlap
ChIP HCT116 ENCFF003KHP 203 bp overlap
ChIP HCT116 ENCFF209YMI 203 bp overlap
ChIP HCT116 ENCFF373YMA 203 bp overlap
ChIP HEK293 ENCSR000DTW.CTCF.HEK293 119 bp overlap
ChIP HL-60 GSE131577.CTCF.HL-60 203 bp overlap
ChIP HMS001 GSE143026.CTCF.HMS001 147 bp overlap
ChIP HT-1080 GSE135580.CTCF.HT-1080 203 bp overlap
ChIP HT-1080_CBL0137-6h GSE122462.CTCF.HT-1080_CBL0137-6h 203 bp overlap
ChIP HUDEP-2_30min GSE104676.CTCF.HUDEP-2_30min 79 bp overlap
ChIP HUES-64 GSE97394.CTCF.HUES-64 203 bp overlap
ChIP HUES-64_DNMT-KO GSE97394.CTCF.HUES-64_DNMT-KO 203 bp overlap
ChIP HeLa GSE126990.CTCF.HeLa 159 bp overlap
ChIP HeLa-Kyoto GSE138405.CTCF.HeLa-Kyoto 159 bp overlap
ChIP HeLa-Kyoto_PDS5-depleted GSE102884.CTCF.HeLa-Kyoto_PDS5-depleted 156 bp overlap
ChIP HeLa-Kyoto_WAPL-depleted GSE102884.CTCF.HeLa-Kyoto_WAPL-depleted 203 bp overlap
ChIP HeLa-Kyoto_WAPL_PDS-depleted GSE102884.CTCF.HeLa-Kyoto_WAPL_PDS-depleted 203 bp overlap
ChIP HeLa-S3 ENCFF255ASZ 203 bp overlap
ChIP HeLa-S3 ENCFF565UFR 161 bp overlap
ChIP HeLa-S3 ENCFF626XQK 199 bp overlap
ChIP HeLa-S3 ENCSR000AOA.CTCF.HeLa-S3 203 bp overlap
ChIP HeLa-S3 ENCSR000DLO.CTCF.HeLa-S3 126 bp overlap
ChIP HeLa-S3 ENCSR000DUB.CTCF.HeLa-S3 163 bp overlap
ChIP HeLa-S3_biotin GSE108869.CTCF.HeLa-S3_biotin 167 bp overlap
ChIP HeLa-S3_shCtrl GSE137848.CTCF.HeLa-S3_shCtrl 203 bp overlap
ChIP HeLa-S3_unsynchro GSE108173.CTCF.HeLa-S3_unsynchro 203 bp overlap
ChIP HeLa_dC9Sun-D3AMut_MIR152 GSE107607.CTCF.HeLa_dC9Sun-D3AMut_MIR152 203 bp overlap
ChIP HeLa_dC9Sun-D3A_CDCC85C GSE107607.CTCF.HeLa_dC9Sun-D3A_CDCC85C 172 bp overlap
ChIP HeLa_dC9Sun-D3A_MIR152 GSE107607.CTCF.HeLa_dC9Sun-D3A_MIR152 198 bp overlap
ChIP HeLa_dC9Sun-D3A_SHB GSE107607.CTCF.HeLa_dC9Sun-D3A_SHB 203 bp overlap
ChIP Hep-G2 ENCSR000AMA.CTCF.Hep-G2 177 bp overlap
ChIP Hep-G2 ERP000209.CTCF.Hep-G2 163 bp overlap
ChIP Hep-G2 ENCSR000BIE.CTCF.Hep-G2 145 bp overlap
ChIP Hep-G2 ENCSR000DLS.CTCF.Hep-G2 100 bp overlap
ChIP HepG2 ENCFF194VBQ 203 bp overlap
ChIP HepG2 ENCFF348BUL 174 bp overlap
ChIP HepG2 ENCFF668CTD 144 bp overlap
ChIP HepG2 ENCFF757EKU 203 bp overlap
ChIP ID00014 GSE76922.CTCF.ID00014 203 bp overlap
ChIP ID00015 GSE76922.CTCF.ID00015 203 bp overlap
ChIP ID00016 GSE76922.CTCF.ID00016 203 bp overlap
ChIP Jurkat GSE68976.CTCF.Jurkat 203 bp overlap
ChIP Jurkat GSE130140.CTCF.Jurkat 145 bp overlap
ChIP Jurkat_DMSO GSE130140.CTCF.Jurkat_DMSO 157 bp overlap
ChIP Jurkat_GSI3d_w4hr GSE130140.CTCF.Jurkat_GSI3d_w4hr 172 bp overlap
ChIP K-562 ENCSR000AKO.CTCF.K-562 203 bp overlap
ChIP K-562 GSE92879.CTCF.K-562 203 bp overlap
ChIP K-562 GSE70482.CTCF.K-562 203 bp overlap
ChIP K-562 GSE70764.CTCF.K-562 203 bp overlap
ChIP K-562 ENCSR000BPJ.CTCF.K-562 203 bp overlap
ChIP K-562 GSE110681.CTCF.K-562 203 bp overlap
ChIP K-562 ENCSR000EGM.CTCF.K-562 200 bp overlap
ChIP K-562 ENCSR000DMA.CTCF.K-562 187 bp overlap
ChIP K-562 ENCSR000DWE.CTCF.K-562 187 bp overlap
ChIP K-562_11698 GSE131349.CTCF.K-562_11698 107 bp overlap
ChIP K-562_12040 GSE131349.CTCF.K-562_12040 114 bp overlap
ChIP K-562_13004 GSE131349.CTCF.K-562_13004 134 bp overlap
ChIP K-562_13189 GSE131349.CTCF.K-562_13189 102 bp overlap
ChIP K-562_14259 GSE131349.CTCF.K-562_14259 203 bp overlap
ChIP K-562_14376 GSE131349.CTCF.K-562_14376 122 bp overlap
ChIP K-562_15173 GSE131349.CTCF.K-562_15173 121 bp overlap
ChIP K-562_15776 GSE131349.CTCF.K-562_15776 155 bp overlap
ChIP K-562_15923 GSE131349.CTCF.K-562_15923 195 bp overlap
ChIP K-562_5635 GSE131349.CTCF.K-562_5635 164 bp overlap
ChIP K-562_7140 GSE131349.CTCF.K-562_7140 147 bp overlap
ChIP K-562_8004 GSE131349.CTCF.K-562_8004 186 bp overlap
ChIP K-562_8005 GSE131349.CTCF.K-562_8005 144 bp overlap
ChIP K-562_CRISPRa_N4293 GSE131349.CTCF.K-562_CRISPRa_N4293 147 bp overlap
ChIP K-562_CRISPRi_N4293 GSE131349.CTCF.K-562_CRISPRi_N4293 167 bp overlap
ChIP K-562_Dox GSE92879.CTCF.K-562_Dox 203 bp overlap
ChIP K-562_MonoC GSE137216.CTCF.K-562_MonoC 203 bp overlap
ChIP K-562_MonoN GSE137216.CTCF.K-562_MonoN 177 bp overlap
ChIP K-562_RF2-insertion GSE140868.CTCF.K-562_RF2-insertion 156 bp overlap
ChIP K-562_RF2-mutation GSE140868.CTCF.K-562_RF2-mutation 190 bp overlap
ChIP K-562_RabbitC GSE137216.CTCF.K-562_RabbitC 203 bp overlap
ChIP K-562_WT GSE140868.CTCF.K-562_WT 203 bp overlap
ChIP K-562_dCas9-KRAB GSE132212.CTCF.K-562_dCas9-KRAB 203 bp overlap
ChIP K-562_dCas9-LSD1 GSE132212.CTCF.K-562_dCas9-LSD1 203 bp overlap
ChIP K-562_enCRISPRi-KL GSE132212.CTCF.K-562_enCRISPRi-KL 203 bp overlap
ChIP K-562_enCRISPRi-LK GSE132212.CTCF.K-562_enCRISPRi-LK 203 bp overlap
ChIP K-562_sgGal4 GSE132212.CTCF.K-562_sgGal4 192 bp overlap
ChIP K562 ENCFF082GOI 115 bp overlap
ChIP K562 ENCFF111MGE 143 bp overlap
ChIP K562 ENCFF400DFR 142 bp overlap
ChIP K562 ENCFF430KTH 203 bp overlap
ChIP K562 ENCFF598YSU 203 bp overlap
ChIP KARPAS422 ENCSR113REG.CTCF.KARPAS422 203 bp overlap
ChIP KB_5Z GSE134435.CTCF.KB_5Z 126 bp overlap
ChIP KB_IL-1_5Z GSE134435.CTCF.KB_IL-1_5Z 131 bp overlap
ChIP Kelly_resistant GSE103030.CTCF.Kelly_resistant 203 bp overlap
ChIP Kelly_resistant_JQ1 GSE103030.CTCF.Kelly_resistant_JQ1 193 bp overlap
ChIP Kelly_sensitive GSE103030.CTCF.Kelly_sensitive 188 bp overlap
ChIP Kelly_shB4-res GSE115248.CTCF.Kelly_shB4-res 189 bp overlap
ChIP Kelly_shLUC-res GSE115248.CTCF.Kelly_shLUC-res 163 bp overlap
ChIP LNCAP ENCFF223HIG 203 bp overlap
ChIP LNCAP ENCFF700QXT 203 bp overlap
ChIP LNCaP ENCSR315NAC.CTCF.LNCaP 203 bp overlap
ChIP LNCaP ENCSR000DMF.CTCF.LNCaP 156 bp overlap
ChIP LNCaP ENCSR000DME.CTCF.LNCaP 116 bp overlap
ChIP LNCaP clone FGC ENCFF519YVI 181 bp overlap
ChIP LNCaP clone FGC ENCFF957KCI 181 bp overlap
ChIP LNCaP_Ctrl GSE125639.CTCF.LNCaP_Ctrl 200 bp overlap
ChIP LNCaP_hydroxy_10nM_4h ENCSR673WZL.CTCF.LNCaP_hydroxy_10nM_4h 203 bp overlap
ChIP Loucy ENCFF359TVQ 171 bp overlap
ChIP Loucy ENCSR464DKE.CTCF.Loucy 203 bp overlap
ChIP MCF-10A GSE98551.CTCF.MCF-10A 168 bp overlap
ChIP MCF-7 ENCFF139NQI 203 bp overlap
ChIP MCF-7 ENCFF162GNE 203 bp overlap
ChIP MCF-7 ENCFF198DQX 203 bp overlap
ChIP MCF-7 ENCFF210JUZ 203 bp overlap
ChIP MCF-7 ENCFF414SZG 96 bp overlap
ChIP MCF-7 ENCFF424NQR 105 bp overlap
ChIP MCF-7 ENCFF494VXA 203 bp overlap
ChIP MCF-7 ENCFF844STM 103 bp overlap
ChIP MCF-7 ENCFF954TUV 200 bp overlap
ChIP MCF-7 GSE137216.CTCF.MCF-7 203 bp overlap
ChIP MCF-7 GSE70764.CTCF.MCF-7 203 bp overlap
ChIP MCF-7 ENCSR560BUE.CTCF.MCF-7 203 bp overlap
ChIP MCF-7 ENCSR000DWH.CTCF.MCF-7 203 bp overlap
ChIP MCF-7 ENCSR000AHD.CTCF.MCF-7 198 bp overlap
ChIP MCF-7 ENCSR000DMR.CTCF.MCF-7 194 bp overlap
ChIP MCF-7 ENCSR000DMS.CTCF.MCF-7 162 bp overlap
ChIP MCF-7 ERP000380.CTCF.MCF-7 152 bp overlap
ChIP MCF-7L_TAMR GSE108787.CTCF.MCF-7L_TAMR 203 bp overlap
ChIP MCF-7L_t0 GSE108787.CTCF.MCF-7L_t0 203 bp overlap
ChIP MCF-7L_t1 GSE108787.CTCF.MCF-7L_t1 203 bp overlap
ChIP MCF-7L_t24 GSE108787.CTCF.MCF-7L_t24 203 bp overlap
ChIP MCF-7_E2 ERP000380.CTCF.MCF-7_E2 194 bp overlap
ChIP MCF-7_E2_SHCTCF ERP000209.CTCF.MCF-7_E2_SHCTCF 165 bp overlap
ChIP MCF-7_HYPOXIA GSE78113.CTCF.MCF-7_HYPOXIA 203 bp overlap
ChIP MCF-7_NORMOXIA GSE78113.CTCF.MCF-7_NORMOXIA 203 bp overlap
ChIP MCF-7_TAM ERP000380.CTCF.MCF-7_TAM 143 bp overlap
ChIP MCF-7_TAM ERP000783.CTCF.MCF-7_TAM 142 bp overlap
ChIP MDA-MB-157 GSE116868.CTCF.MDA-MB-157 203 bp overlap
ChIP MDA-MB-435_Asynchronous GSE102019.CTCF.MDA-MB-435_Asynchronous 180 bp overlap
ChIP MM.1S ENCFF869JMQ 203 bp overlap
ChIP MM1-S ENCSR402IDP.CTCF.MM1-S 203 bp overlap
ChIP MM1-S GSE43743.CTCF.MM1-S 197 bp overlap
ChIP NB4 ENCFF155DNY 203 bp overlap
ChIP NB4 ENCSR000DWN.CTCF.NB4 203 bp overlap
ChIP NB69 GSE101295.CTCF.NB69 182 bp overlap
ChIP NCI-H929 ENCFF305JAB 203 bp overlap
ChIP NCI-H929 ENCSR634OAQ.CTCF.NCI-H929 203 bp overlap
ChIP OCI-LY1 ENCFF455ESK 162 bp overlap
ChIP OCI-LY3 ENCFF939BYJ 203 bp overlap
ChIP OCI-Ly1 ENCSR072EUE.CTCF.OCI-Ly1 203 bp overlap
ChIP OCI-Ly3 ENCSR756ZKG.CTCF.OCI-Ly3 203 bp overlap
ChIP PC-3 ENCFF487TUI 203 bp overlap
ChIP PC-3 ENCSR359LOD.CTCF.PC-3 203 bp overlap
ChIP Panc1 ENCFF056JQX 203 bp overlap
ChIP RH4 GSE83726.CTCF.RH4 160 bp overlap
ChIP SEM GSE117864.CTCF.SEM 203 bp overlap
ChIP SH-SY5Y GSE101295.CTCF.SH-SY5Y 116 bp overlap
ChIP SK-MEL-147 GSE94488.CTCF.SK-MEL-147 203 bp overlap
ChIP SK-N-AS GSE101295.CTCF.SK-N-AS 192 bp overlap
ChIP SK-N-SH ENCSR000BLX.CTCF.SK-N-SH 92 bp overlap
ChIP SU-DHL-6 ENCFF116KKR 203 bp overlap
ChIP T-47D GSE111923.CTCF.T-47D 203 bp overlap
ChIP T-47D GSE120162.CTCF.T-47D 199 bp overlap
ChIP T-47D ENCSR000BNO.CTCF.T-47D 157 bp overlap
ChIP T-47D_NaCl-7-5min GSE111923.CTCF.T-47D_NaCl-7-5min 203 bp overlap
ChIP T-47D_NaCl-isotonic GSE111923.CTCF.T-47D_NaCl-isotonic 203 bp overlap
ChIP T-47D_NaCl-isotonic-triptolide GSE111923.CTCF.T-47D_NaCl-isotonic-triptolide 203 bp overlap
ChIP T-47D_Y537S GSE148277.CTCF.T-47D_Y537S 203 bp overlap
ChIP T-47D_serum GSE120162.CTCF.T-47D_serum 202 bp overlap
ChIP T-47D_triptolide GSE111923.CTCF.T-47D_triptolide 203 bp overlap
ChIP TALL-1_Pat1 GSE130140.CTCF.TALL-1_Pat1 161 bp overlap
ChIP THP-1_Pam3csk-020m-Flavo-000m GSE103477.CTCF.THP-1_Pam3csk-020m-Flavo-000m 163 bp overlap
ChIP THP-1_Pam3csk-030m-Flavo-000m GSE103477.CTCF.THP-1_Pam3csk-030m-Flavo-000m 155 bp overlap
ChIP THP-1_Pam3csk-360m-Flavo-240m GSE103477.CTCF.THP-1_Pam3csk-360m-Flavo-240m 198 bp overlap
ChIP VCaP ENCFF858YQT 203 bp overlap
ChIP VCaP ENCSR265ARE.CTCF.VCaP 203 bp overlap
ChIP WA01 ENCSR000BNH.CTCF.WA01 201 bp overlap
ChIP WA01 ENCSR000DLK.CTCF.WA01 168 bp overlap
ChIP WA01_NEUR GSE52457.CTCF.WA01_NEUR 192 bp overlap
ChIP WERI-Rb-1 ENCFF349QKF 203 bp overlap
ChIP WERI-Rb-1 ENCSR000DXW.CTCF.WERI-Rb-1 182 bp overlap
ChIP ZR751 ERP000783.CTCF.ZR751 162 bp overlap
ChIP body of pancreas ENCFF128ALM 203 bp overlap
ChIP body of pancreas ENCFF269EDN 122 bp overlap
ChIP body of pancreas ENCFF438KTE 182 bp overlap
ChIP body of pancreas ENCFF756FGB 203 bp overlap
ChIP body of pancreas ENCFF798MEO 167 bp overlap
ChIP body of pancreas ENCFF881RGF 203 bp overlap
ChIP bonchial_epithelial ENCSR000DXI.CTCF.bonchial_epithelial 186 bp overlap
ChIP bronchial epithelial cell ENCFF500SEA 203 bp overlap
ChIP cardiac-muscle ENCSR713SXF.CTCF.cardiac-muscle 197 bp overlap
ChIP delta-47 GSE70764.CTCF.delta-47 203 bp overlap
ChIP endodermal cell ENCFF471YCZ 203 bp overlap
ChIP epithelial cell of esophagus ENCFF946GGT 203 bp overlap
ChIP epithelial cell of prostate ENCFF086GTI 203 bp overlap
ChIP epithelial_esophagus ENCSR000DTR.CTCF.epithelial_esophagus 163 bp overlap
ChIP epithelial_mammary ENCSR000DUS.CTCF.epithelial_mammary 195 bp overlap
ChIP epithelial_mammary ENCSR000ALV.CTCF.epithelial_mammary 173 bp overlap
ChIP epithelial_retinal_pigment ENCSR000DVI.CTCF.epithelial_retinal_pigment 121 bp overlap
ChIP erythroblast_adult_erythroblasts GSE102184.CTCF.erythroblast_adult_erythroblasts 203 bp overlap
ChIP erythroid GSE67783.CTCF.erythroid 203 bp overlap
ChIP erythroid_Don001 GSE137982.CTCF.erythroid_Don001 190 bp overlap
ChIP fetal_erythroblasts GSE102184.CTCF.fetal_erythroblasts 203 bp overlap
ChIP foreskin keratinocyte ENCFF980OWR 203 bp overlap
ChIP hESC_NAIVE GSE69646.CTCF.hESC_NAIVE 203 bp overlap
ChIP hepatocyte ERP000395.CTCF.hepatocyte 201 bp overlap
ChIP hiPSC GSE115407.CTCF.hiPSC 189 bp overlap
ChIP hiPSC_CVI GSE106870.CTCF.hiPSC_CVI 198 bp overlap
ChIP hiPSC_HUES9 GSE106870.CTCF.hiPSC_HUES9 203 bp overlap
ChIP hiPSC_IB12 GSE106870.CTCF.hiPSC_IB12 203 bp overlap
ChIP hiPSC_IB7 GSE106870.CTCF.hiPSC_IB7 203 bp overlap
ChIP hiPSC_IIA11 GSE106870.CTCF.hiPSC_IIA11 203 bp overlap
ChIP hiPSC_IID12 GSE106870.CTCF.hiPSC_IID12 191 bp overlap
ChIP hiPSC_IIIB12 GSE106870.CTCF.hiPSC_IIIB12 150 bp overlap
ChIP hiPSC_IV8 GSE106870.CTCF.hiPSC_IV8 203 bp overlap
ChIP hiPSC_failed-genome-editing GSE106870.CTCF.hiPSC_failed-genome-editing 203 bp overlap
ChIP hiPSC_parental-line GSE106870.CTCF.hiPSC_parental-line 203 bp overlap
ChIP keratinocyte ENCFF046PBT 185 bp overlap
ChIP keratinocyte ENCFF291YDC 185 bp overlap
ChIP keratinocyte ENCFF667ULX 203 bp overlap
ChIP keratinocyte GSE154221.CTCF.keratinocyte 203 bp overlap
ChIP keratinocyte ENCSR000ALJ.CTCF.keratinocyte 176 bp overlap
ChIP keratinocyte ENCSR000DWX.CTCF.keratinocyte 203 bp overlap
ChIP keratinocyte ENCSR000DNC.CTCF.keratinocyte 147 bp overlap
ChIP leukemia GSE142161.CTCF.leukemia 203 bp overlap
ChIP leukemia_CEBPA-168h GSE142161.CTCF.leukemia_CEBPA-168h 203 bp overlap
ChIP leukemia_CEBPA-24h GSE142161.CTCF.leukemia_CEBPA-24h 203 bp overlap
ChIP leukemia_CEBPA-96h GSE142161.CTCF.leukemia_CEBPA-96h 203 bp overlap
ChIP leukemia_DMSO-24h GSE142161.CTCF.leukemia_DMSO-24h 203 bp overlap
ChIP lymphoblast_mut GSE155324.CTCF.lymphoblast_mut 203 bp overlap
ChIP mammary epithelial cell ENCFF164SPU 203 bp overlap
ChIP mammary epithelial cell ENCFF873ERE 203 bp overlap
ChIP neuron_bipolar_doxy_4d ENCSR619IUE.CTCF.neuron_bipolar_doxy_4d 136 bp overlap
ChIP pancreas ENCSR687APM.CTCF.pancreas 203 bp overlap
ChIP pancreas ENCSR585KBH.CTCF.pancreas 185 bp overlap
ChIP pancreas ENCSR000DND.CTCF.pancreas 124 bp overlap
ChIP pancreas_body ENCSR265PFQ.CTCF.pancreas_body 203 bp overlap
ChIP pancreas_body ENCSR307PFP.CTCF.pancreas_body 203 bp overlap
ChIP pancreas_body ENCSR408XTO.CTCF.pancreas_body 202 bp overlap
ChIP pancreas_body ENCSR484DDO.CTCF.pancreas_body 165 bp overlap
ChIP pancreas_body ENCSR572DUJ.CTCF.pancreas_body 156 bp overlap
ChIP placenta ENCFF029PHY 203 bp overlap
ChIP plasmablast GSE142493.CTCF.plasmablast 175 bp overlap
ChIP primary-epidermal-keratinocyte_diff_d0 GSE84657.CTCF.primary-epidermal-keratinocyte_diff_d0 171 bp overlap
ChIP primary-epidermal-keratinocyte_diff_d6 GSE84657.CTCF.primary-epidermal-keratinocyte_diff_d6 196 bp overlap
ChIP primary-glioblastoma_G583 GSE121601.CTCF.primary-glioblastoma_G583 190 bp overlap
ChIP progenitor cell of endocrine pancreas ENCFF492KHV 144 bp overlap
ChIP prostate_epithelial ENCSR196HOM.CTCF.prostate_epithelial 203 bp overlap
ChIP retinal pigment epithelial cell ENCFF810AAG 203 bp overlap
Cebpa 1 dataset
ChIP BLaER1 ENCFF335XTP 203 bp overlap
DEAF1 1 dataset
ChIP K562 ENCFF944USZ 203 bp overlap
E2F1 1 dataset
ChIP LNCaP_shRB GSE94958.E2F1.LNCaP_shRB 143 bp overlap
E2F4 1 dataset
ChIP K-562 ENCSR000EWL.E2F4.K-562 184 bp overlap
EP300 1 dataset
ChIP 697 GSE138031.EP300.697 197 bp overlap
ERG 2 datasets
ChIP SEM GSE117864.ERG.SEM 203 bp overlap
ChIP SKNO-1 GSE23730.ERG.SKNO-1 198 bp overlap
ESR1 13 datasets
ChIP MCF-7 GSE119057.ESR1.MCF-7 203 bp overlap
ChIP MCF-7_E2 GSE14664.ESR1.MCF-7_E2 120 bp overlap
ChIP MCF-7_ER_t10 GSE119057.ESR1.MCF-7_ER_t10 203 bp overlap
ChIP MCF-7_ER_t20 GSE119057.ESR1.MCF-7_ER_t20 203 bp overlap
ChIP MCF-7_ER_t30 GSE119057.ESR1.MCF-7_ER_t30 203 bp overlap
ChIP MCF-7_ER_t40 GSE119057.ESR1.MCF-7_ER_t40 203 bp overlap
ChIP MCF-7_ER_t50 GSE119057.ESR1.MCF-7_ER_t50 203 bp overlap
ChIP MCF-7_ER_t60 GSE119057.ESR1.MCF-7_ER_t60 203 bp overlap
ChIP MCF-7_ER_t70 GSE119057.ESR1.MCF-7_ER_t70 203 bp overlap
ChIP MCF-7_ER_t80 GSE119057.ESR1.MCF-7_ER_t80 203 bp overlap
ChIP MCF-7_ER_t90 GSE119057.ESR1.MCF-7_ER_t90 203 bp overlap
ChIP MCF-7_RUNX2_DOX GSE86538.ESR1.MCF-7_RUNX2_DOX 203 bp overlap
ChIP MCF-7_Veh GSE67295.ESR1.MCF-7_Veh 95 bp overlap
EZH2 2 datasets
ChIP Pfeiffer GSE45982.EZH2.Pfeiffer 137 bp overlap
ChIP T98G GSE112240.EZH2.T98G 203 bp overlap
FLI1 1 dataset
ChIP SEM GSE117864.FLI1.SEM 136 bp overlap
FOXO1 1 dataset
ChIP CD34 GSE80773.FOXO1.CD34 121 bp overlap
GFI1B 1 dataset
ChIP K-562 ENCSR509GDT.GFI1B.K-562 180 bp overlap
GRHL2 1 dataset
ChIP OVCAR-3 GSE71018.GRHL2.OVCAR-3 67 bp overlap
HDAC2 1 dataset
ChIP pre-B-cell GSE107886.HDAC2.pre-B-cell 203 bp overlap
HNRNPLL 1 dataset
ChIP K-562 ENCSR112RNT.HNRNPLL.K-562 200 bp overlap
IKZF3 1 dataset
ChIP pre-B-cell GSE107886.IKZF3.pre-B-cell 203 bp overlap
IRF2 1 dataset
ChIP K-562 ENCSR376WCJ.IRF2.K-562 108 bp overlap
JUND 1 dataset
ChIP K-562 ENCSR000EGN.JUND.K-562 133 bp overlap
KDM5B 1 dataset
ChIP T-47D GSE46055.KDM5B.T-47D 188 bp overlap
KMT2A 1 dataset
ChIP L826 GSE83671.KMT2A.L826 178 bp overlap
MAX 1 dataset
ChIP MCF-7 ENCSR000BUL.MAX.MCF-7 107 bp overlap
MED1 2 datasets
ChIP MCF-7_SHCTR GSE60270.MED1.MCF-7_SHCTR 157 bp overlap
ChIP MCF-7_SHRARS_E2 GSE60270.MED1.MCF-7_SHRARS_E2 157 bp overlap
MTA2 1 dataset
ChIP pre-B-cell GSE107886.MTA2.pre-B-cell 203 bp overlap
MYB 1 dataset
ChIP SEM GSE117864.MYB.SEM 151 bp overlap
MYC 1 dataset
ChIP K-562 ENCSR000EGJ.MYC.K-562 117 bp overlap
NFE2L2 1 dataset
ChIP K-562 ENCSR485GKE.NFE2L2.K-562 142 bp overlap
NR2F1 1 dataset
ChIP GM12878 ENCSR514VYD.NR2F1.GM12878 196 bp overlap
PBX1 1 dataset
ChIP RCH-ACV GSE85988.PBX1.RCH-ACV 172 bp overlap
POU2F1 1 dataset
ChIP T-47D_D538G GSE148277.POU2F1.T-47D_D538G 203 bp overlap
POU2F2 1 dataset
ChIP pre-B-cell GSE107886.POU2F2.pre-B-cell 203 bp overlap
PRDM14 1 dataset
ChIP NCCIT GSE71675.PRDM14.NCCIT 203 bp overlap
RAD21 62 datasets
ChIP A-549 ENCSR000BUC.RAD21.A-549 162 bp overlap
ChIP A549 ENCFF047SFC 203 bp overlap
ChIP GM12878 ENCFF046CBW 203 bp overlap
ChIP GM12878 ENCFF101UQZ 106 bp overlap
ChIP GM12878 ENCSR000EAC.RAD21.GM12878 198 bp overlap
ChIP GM12878 ENCSR000BMY.RAD21.GM12878 195 bp overlap
ChIP GP5D GSE51234.RAD21.GP5D 203 bp overlap
ChIP GP5D_SIRAD21 GSE51234.RAD21.GP5D_SIRAD21 194 bp overlap
ChIP H1 ENCFF698EWO 194 bp overlap
ChIP H1 ENCFF967OJF 168 bp overlap
ChIP HAP1 GSE152721.RAD21.HAP1 180 bp overlap
ChIP HCT-116 GSE131606.RAD21.HCT-116 203 bp overlap
ChIP HCT-116 GSE121355.RAD21.HCT-116 198 bp overlap
ChIP HCT-116 ENCSR000BSB.RAD21.HCT-116 188 bp overlap
ChIP HCT-116_RAD21-mAC GSE104888.RAD21.HCT-116_RAD21-mAC 203 bp overlap
ChIP HCT116 ENCFF568PEO 159 bp overlap
ChIP HeLa-S3 ENCFF775CHI 203 bp overlap
ChIP HeLa-S3_CTCF-s-OE GSE108869.RAD21.HeLa-S3_CTCF-s-OE 203 bp overlap
ChIP HeLa-Tet-On GSE112028.RAD21.HeLa-Tet-On 203 bp overlap
ChIP Hep-G2 ENCSR054FKH.RAD21.Hep-G2 158 bp overlap
ChIP Hep-G2 ENCSR000EEG.RAD21.Hep-G2 153 bp overlap
ChIP Hep-G2 ERP000209.RAD21.Hep-G2 143 bp overlap
ChIP HepG2 ENCFF360ZSW 199 bp overlap
ChIP HepG2 ENCFF906QIS 203 bp overlap
ChIP HepG2 ENCFF963UBJ 203 bp overlap
ChIP Ishikawa ENCFF570JVV 202 bp overlap
ChIP Ishikawa ENCSR000BTU.RAD21.Ishikawa 169 bp overlap
ChIP K-562 ENCSR000FAD.RAD21.K-562 203 bp overlap
ChIP K-562 ENCSR000BKV.RAD21.K-562 198 bp overlap
ChIP K-562_RF2-insertion GSE140868.RAD21.K-562_RF2-insertion 142 bp overlap
ChIP K562 ENCFF066JWO 168 bp overlap
ChIP K562 ENCFF169SQI 181 bp overlap
ChIP K562 ENCFF192VNH 192 bp overlap
ChIP K562 ENCFF634XYR 202 bp overlap
ChIP MCF-7 ENCFF724VCQ 195 bp overlap
ChIP MCF-7 GSE137216.RAD21.MCF-7 203 bp overlap
ChIP MCF-7 GSE72082.RAD21.MCF-7 200 bp overlap
ChIP MCF-7 ERP000209.RAD21.MCF-7 197 bp overlap
ChIP MCF-7 ENCSR000BTQ.RAD21.MCF-7 190 bp overlap
ChIP MCF-7 ENCSR703TNG.RAD21.MCF-7 187 bp overlap
ChIP MCF-7_E2 ERP000209.RAD21.MCF-7_E2 173 bp overlap
ChIP MCF-7_E2_SHCTCF ERP000209.RAD21.MCF-7_E2_SHCTCF 117 bp overlap
ChIP SLK_Ctrl GSE138105.RAD21.SLK_Ctrl 203 bp overlap
ChIP T-47D GSE111923.RAD21.T-47D 201 bp overlap
ChIP T-47D_NaCl-isotonic-triptolide GSE111923.RAD21.T-47D_NaCl-isotonic-triptolide 203 bp overlap
ChIP T-47D_triptolide GSE111923.RAD21.T-47D_triptolide 203 bp overlap
ChIP THP-1_Pam3csk-045m-Flavo-000m GSE103477.RAD21.THP-1_Pam3csk-045m-Flavo-000m 161 bp overlap
ChIP WA09 GSE105028.RAD21.WA09 203 bp overlap
ChIP WA09_heat-shock GSE105028.RAD21.WA09_heat-shock 191 bp overlap
ChIP hiPSC_CVI GSE106870.RAD21.hiPSC_CVI 203 bp overlap
ChIP hiPSC_HUES9 GSE106870.RAD21.hiPSC_HUES9 145 bp overlap
ChIP hiPSC_IB12 GSE106870.RAD21.hiPSC_IB12 202 bp overlap
ChIP hiPSC_IB7 GSE106870.RAD21.hiPSC_IB7 203 bp overlap
ChIP hiPSC_IIA11 GSE106870.RAD21.hiPSC_IIA11 203 bp overlap
ChIP hiPSC_IIA12 GSE106870.RAD21.hiPSC_IIA12 203 bp overlap
ChIP hiPSC_IID12 GSE106870.RAD21.hiPSC_IID12 185 bp overlap
ChIP hiPSC_IIIB12 GSE106870.RAD21.hiPSC_IIIB12 203 bp overlap
ChIP hiPSC_IV8 GSE106870.RAD21.hiPSC_IV8 203 bp overlap
ChIP hiPSC_failed-genome-editing GSE106870.RAD21.hiPSC_failed-genome-editing 203 bp overlap
ChIP hiPSC_parental-line GSE106870.RAD21.hiPSC_parental-line 203 bp overlap
ChIP lymphoblast GSE155324.RAD21.lymphoblast 203 bp overlap
ChIP lymphoblast_mut GSE155324.RAD21.lymphoblast_mut 203 bp overlap
RELB 1 dataset
ChIP GM12878 ENCSR387QUV.RELB.GM12878 115 bp overlap
REST 2 datasets
ChIP K-562 ENCSR137ZMQ.REST.K-562 203 bp overlap
ChIP K562 ENCFF688UKW 203 bp overlap
RNF2 1 dataset
ChIP K-562 ENCSR138FUZ.RNF2.K-562 130 bp overlap
RUNX1 3 datasets
ChIP 697 GSE138031.RUNX1.697 203 bp overlap
ChIP BCP-ALL_patient1 GSE109377.RUNX1.BCP-ALL_patient1 203 bp overlap
ChIP NALM-6 GSE126300.RUNX1.NALM-6 203 bp overlap
RUNX2 1 dataset
ChIP LNCaP-C4-2B GSE33889.RUNX2.LNCaP-C4-2B 203 bp overlap
SMAD1 2 datasets
ChIP BG03 GSE36578.SMAD1.BG03 102 bp overlap
ChIP CD34_PROG_BMP GSE29194.SMAD1.CD34_PROG_BMP 135 bp overlap
SMAD3 2 datasets
ChIP BG03 GSE36578.SMAD3.BG03 124 bp overlap
ChIP HCC1954 GSE104760.SMAD3.HCC1954 146 bp overlap
SMARCA5 1 dataset
ChIP GM12878 ENCSR706YUH.SMARCA5.GM12878 112 bp overlap
SMC1 3 datasets
ChIP DKO GSE131606.SMC1.DKO 203 bp overlap
ChIP HCT-116 GSE131606.SMC1.HCT-116 203 bp overlap
ChIP HCT-116_RAD21-mAC GSE104888.SMC1.HCT-116_RAD21-mAC 203 bp overlap
SMC1A 8 datasets
ChIP A-549 GSE76893.SMC1A.A-549 190 bp overlap
ChIP Hep-G2 GSE76893.SMC1A.Hep-G2 146 bp overlap
ChIP MCF-7 GSE115602.SMC1A.MCF-7 203 bp overlap
ChIP MCF-7 GSE76893.SMC1A.MCF-7 138 bp overlap
ChIP MCF-7_EtOH GSE115602.SMC1A.MCF-7_EtOH 203 bp overlap
ChIP primary-epidermal-keratinocyte GSE85526.SMC1A.primary-epidermal-keratinocyte 203 bp overlap
ChIP primary-epidermal-keratinocyte_diff_d0 GSE84657.SMC1A.primary-epidermal-keratinocyte_diff_d0 117 bp overlap
ChIP primary-epidermal-keratinocyte_diff_d6 GSE84657.SMC1A.primary-epidermal-keratinocyte_diff_d6 203 bp overlap
SMC3 12 datasets
ChIP GM12878 ENCFF085RLZ 203 bp overlap
ChIP GM12878 ENCSR000DZP.SMC3.GM12878 179 bp overlap
ChIP GP5D GSE51234.SMC3.GP5D 203 bp overlap
ChIP HeLa GSE126990.SMC3.HeLa 203 bp overlap
ChIP HeLa-Kyoto GSE102884.SMC3.HeLa-Kyoto 203 bp overlap
ChIP HeLa-Kyoto GSE138405.SMC3.HeLa-Kyoto 203 bp overlap
ChIP HeLa-Kyoto_PDS5-depleted GSE102884.SMC3.HeLa-Kyoto_PDS5-depleted 111 bp overlap
ChIP HeLa-Kyoto_WAPL_PDS-depleted GSE102884.SMC3.HeLa-Kyoto_WAPL_PDS-depleted 203 bp overlap
ChIP HeLa-S3 ENCFF992MML 203 bp overlap
ChIP HeLa-S3 ENCSR000ECS.SMC3.HeLa-S3 124 bp overlap
ChIP K-562 ENCSR000EGW.SMC3.K-562 191 bp overlap
ChIP K562 ENCFF582XIX 203 bp overlap
STAG1 10 datasets
ChIP HL-60 GSE131577.STAG1.HL-60 124 bp overlap
ChIP HeLa GSE126990.STAG1.HeLa 203 bp overlap
ChIP HeLa-Kyoto GSE138405.STAG1.HeLa-Kyoto 203 bp overlap
ChIP Hep-G2 ENCSR167MTG.STAG1.Hep-G2 171 bp overlap
ChIP Hep-G2 ERP000209.STAG1.Hep-G2 148 bp overlap
ChIP HepG2 ENCFF843EBZ 203 bp overlap
ChIP MCF-7 ERP000209.STAG1.MCF-7 202 bp overlap
ChIP MCF-7_E2 ERP000209.STAG1.MCF-7_E2 166 bp overlap
ChIP MCF-7_E2_SHCTCF ERP000209.STAG1.MCF-7_E2_SHCTCF 153 bp overlap
ChIP erythroid GSE67783.STAG1.erythroid 203 bp overlap
STAG2 1 dataset
ChIP HL-60 GSE131577.STAG2.HL-60 152 bp overlap
STAT3 1 dataset
ChIP T-47D_JC5056 GSE126004.STAT3.T-47D_JC5056 203 bp overlap
TCF12 1 dataset
ChIP GM12878 ENCSR000BGZ.TCF12.GM12878 155 bp overlap
TCF3 1 dataset
ChIP RCH-ACV GSE85988.TCF3.RCH-ACV 203 bp overlap
TFAP2A 1 dataset
ChIP MCF-7_E2 GSE60270.TFAP2A.MCF-7_E2 202 bp overlap
TFAP2C 1 dataset
ChIP HCC1954 GSE126897.TFAP2C.HCC1954 163 bp overlap
TRIM22 1 dataset
ChIP GM12878 ENCSR835XKS.TRIM22.GM12878 86 bp overlap
ZBTB2 1 dataset
ChIP GM12878 GSE97661.ZBTB2.GM12878 185 bp overlap
ZNF143 5 datasets
ChIP GM12878 ENCSR936XTK.ZNF143.GM12878 203 bp overlap
ChIP K-562 ENCSR000EGP.ZNF143.K-562 124 bp overlap
ChIP MCF-7 GSE76454.ZNF143.MCF-7 187 bp overlap
ChIP MCF-7_E2 GSE76454.ZNF143.MCF-7_E2 203 bp overlap
ChIP WA01 ENCSR000EBW.ZNF143.WA01 165 bp overlap
ZNF319 1 dataset
ChIP K562 ENCFF561ZSB 130 bp overlap
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