chr2 : 241,555,780 241,556,049
269 bp 87 TFs 2 linked genes
This 269 bp open chromatin element is linked to BOK and BOK-AS1 and is bound by 87 transcription factors.
Linked Genes
2 genes
Gene Expression Dist. to TSS Distance Link type
BOK 2.7 kb Proximal Proximity
BOK-AS1 2.9 kb Proximal Proximity
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr2:241,550,780 – 241,561,049
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
87 transcription factors
Source
Cell type
AR 3 datasets
ChIP LNCaP_FOXA1 ERP003503.AR.LNCaP_FOXA1 178 bp overlap
ChIP PC-3_R1881 GSE54110.AR.PC-3_R1881 269 bp overlap
ChIP VCaP_DHT24H_SHFOXA1 GSE58428.AR.VCaP_DHT24H_SHFOXA1 207 bp overlap
ARID4B 1 dataset
ChIP HepG2 ENCFF519OXJ 269 bp overlap
ATF3 1 dataset
ChIP HCT-116_CAMP GSE74355.ATF3.HCT-116_CAMP 144 bp overlap
BCOR 1 dataset
ChIP WA01_RNF2-R_Doxycyclin GSE104690.BCOR.WA01_RNF2-R_Doxycyclin 188 bp overlap
BMPR1A 1 dataset
ChIP HUVEC-C GSE60156.BMPR1A.HUVEC-C 269 bp overlap
BRD4 1 dataset
ChIP Jurkat GSE83777.BRD4.Jurkat 269 bp overlap
CDK7 2 datasets
ChIP Jurkat GSE83777.CDK7.Jurkat 210 bp overlap
ChIP Jurkat_DMSO GSE60027.CDK7.Jurkat_DMSO 176 bp overlap
CHD4 1 dataset
ChIP keratinocyte_CTR GSE139685.CHD4.keratinocyte_CTR 269 bp overlap
CHD7 1 dataset
ChIP hiPSC_derived_lt-NES GSE108506.CHD7.hiPSC_derived_lt-NES 217 bp overlap
CREBBP 1 dataset
ChIP keratinocyte GSE140991.CREBBP.keratinocyte 241 bp overlap
CTCF 1 dataset
ChIP SUM159 GSE46055.CTCF.SUM159 103 bp overlap
Cebpa 2 datasets
ChIP BLaER1 ENCFF335XTP 100 bp overlap
ChIP BLaER1 ENCFF364PUR 269 bp overlap
DRAP1 1 dataset
ChIP Hep-G2 ENCSR765MKZ.DRAP1.Hep-G2 148 bp overlap
E4F1 1 dataset
ChIP K-562 ENCSR731LHZ.E4F1.K-562 247 bp overlap
EGR1 1 dataset
ChIP Hep-G2 ENCSR026GSW.EGR1.Hep-G2 150 bp overlap
EHF 2 datasets
ChIP RWPE-1 GSE114241.EHF.RWPE-1 269 bp overlap
ChIP primary-bronchial-epithelial GSE85401.EHF.primary-bronchial-epithelial 261 bp overlap
ELF1 11 datasets
ChIP HCT-116 ENCSR000BVH.ELF1.HCT-116 253 bp overlap
ChIP Hep-G2 ENCSR321VGW.ELF1.Hep-G2 216 bp overlap
ChIP Hep-G2 ENCSR000BMZ.ELF1.Hep-G2 214 bp overlap
ChIP HepG2 ENCFF367ZWV 269 bp overlap
ChIP HepG2 ENCFF838BCU 239 bp overlap
ChIP K-562 ENCSR000BMD.ELF1.K-562 173 bp overlap
ChIP MCF-7 ENCFF305BNP 269 bp overlap
ChIP MCF-7 ENCFF687CWI 99 bp overlap
ChIP MCF-7 ENCSR000BSS.ELF1.MCF-7 269 bp overlap
ChIP MCF-7 ENCSR502NRF.ELF1.MCF-7 227 bp overlap
ChIP SK-N-MC_SHGFP_96H GSE61944.ELF1.SK-N-MC_SHGFP_96H 226 bp overlap
ELF3 4 datasets
ChIP HepG2 ENCFF633ULY 128 bp overlap
ChIP PDAC GSE64557.ELF3.PDAC 269 bp overlap
ChIP PDAC_KOKLF5 GSE64557.ELF3.PDAC_KOKLF5 269 bp overlap
ChIP PDAC_SHCTR GSE64557.ELF3.PDAC_SHCTR 269 bp overlap
EOMES 1 dataset
ChIP hESC GSE26097.EOMES.hESC 157 bp overlap
ERG 13 datasets
ChIP HUVEC-C GSE124891.ERG.HUVEC-C 260 bp overlap
ChIP HUVEC-C GSE128382.ERG.HUVEC-C 185 bp overlap
ChIP Jurkat GSE49091.ERG.Jurkat 262 bp overlap
ChIP RWPE-1 GSE114241.ERG.RWPE-1 269 bp overlap
ChIP RWPE-1 GSE37752.ERG.RWPE-1 180 bp overlap
ChIP TSU-1621MT_ATRA GSE60477.ERG.TSU-1621MT_ATRA 269 bp overlap
ChIP aortic-endothelial-cell_D14 GSE139377.ERG.aortic-endothelial-cell_D14 194 bp overlap
ChIP aortic-endothelial-cell_D19 GSE139377.ERG.aortic-endothelial-cell_D19 195 bp overlap
ChIP aortic-endothelial-cell_D28 GSE139377.ERG.aortic-endothelial-cell_D28 237 bp overlap
ChIP aortic-endothelial-cell_D39 GSE139377.ERG.aortic-endothelial-cell_D39 181 bp overlap
ChIP aortic-endothelial-cell_D40 GSE139377.ERG.aortic-endothelial-cell_D40 165 bp overlap
ChIP aortic-endothelial-cell_D47 GSE139377.ERG.aortic-endothelial-cell_D47 198 bp overlap
ChIP aortic-endothelial-cell_D5 GSE139377.ERG.aortic-endothelial-cell_D5 201 bp overlap
ESR1 2 datasets
ChIP MCF-7_IKK7 GSE67295.ESR1.MCF-7_IKK7 159 bp overlap
ChIP breast_tumor_Male_14 GSE104399.ESR1.breast_tumor_Male_14 192 bp overlap
ETS1 13 datasets
ChIP CTV-1_DSG GSE128835.ETS1.CTV-1_DSG 269 bp overlap
ChIP CTV-1_DSG_SPI1-mut GSE128835.ETS1.CTV-1_DSG_SPI1-mut 262 bp overlap
ChIP GM23338 ENCFF701IZH 269 bp overlap
ChIP HUVEC-C GSE41166.ETS1.HUVEC-C 228 bp overlap
ChIP HUVEC-C_12h GSE93030.ETS1.HUVEC-C_12h 269 bp overlap
ChIP HUVEC-C_4h GSE93030.ETS1.HUVEC-C_4h 269 bp overlap
ChIP HUVEC-C_VEGF_12H GSE41166.ETS1.HUVEC-C_VEGF_12H 269 bp overlap
ChIP HUVEC-C_VEGF_12h GSE109625.ETS1.HUVEC-C_VEGF_12h 269 bp overlap
ChIP HUVEC-C_VEGF_4H GSE41166.ETS1.HUVEC-C_VEGF_4H 269 bp overlap
ChIP HUVEC-C_VEGF_4h GSE109625.ETS1.HUVEC-C_VEGF_4h 269 bp overlap
ChIP SCC-25 GSE109884.ETS1.SCC-25 201 bp overlap
ChIP THP-6_shCtrl GSE138516.ETS1.THP-6_shCtrl 160 bp overlap
ChIP hESC ENCSR534VHI.ETS1.hESC 239 bp overlap
ETV1 2 datasets
ChIP GIST-T1 GSE106624.ETV1.GIST-T1 182 bp overlap
ChIP LNCaP GSE47120.ETV1.LNCaP 212 bp overlap
ETV2 1 dataset
ChIP induced-endothelial-cell_Veh GSE123906.ETV2.induced-endothelial-cell_Veh 240 bp overlap
ETV5 1 dataset
ChIP HepG2 ENCFF456LSA 263 bp overlap
ETV6 1 dataset
ChIP WTC11 ENCFF812SCD 269 bp overlap
EZH2 1 dataset
ChIP SF8628 GSE94834.EZH2.SF8628 239 bp overlap
FLI1 5 datasets
ChIP A-673_1_KRAB_GFP GSE106914.FLI1.A-673_1_KRAB_GFP 257 bp overlap
ChIP A-673_2_KRAB_eSOX2_1 GSE106914.FLI1.A-673_2_KRAB_eSOX2_1 191 bp overlap
ChIP CTV-1_DSG GSE128835.FLI1.CTV-1_DSG 269 bp overlap
ChIP CTV-1_DSG_SPI1-mut GSE128835.FLI1.CTV-1_DSG_SPI1-mut 242 bp overlap
ChIP HUVEC-C GSE109695.FLI1.HUVEC-C 257 bp overlap
FOS 1 dataset
ChIP endothelial_umbilical-vein ENCSR000EVU.FOS.endothelial_umbilical-vein 129 bp overlap
FOSL2 1 dataset
ChIP Hep-G2 ENCSR000BHP.FOSL2.Hep-G2 112 bp overlap
FOXA1 1 dataset
ChIP breast_tumor_Male_14 GSE104399.FOXA1.breast_tumor_Male_14 213 bp overlap
GABPA 7 datasets
ChIP Hep-G2 GSE72082.GABPA.Hep-G2 269 bp overlap
ChIP Hep-G2 ENCSR269TNX.GABPA.Hep-G2 268 bp overlap
ChIP Hep-G2 ENCSR000BJK.GABPA.Hep-G2 145 bp overlap
ChIP HepG2 ENCFF180FFY 238 bp overlap
ChIP MCF-7 ENCFF735CHO 197 bp overlap
ChIP MCF-7 ENCSR000BUK.GABPA.MCF-7 230 bp overlap
ChIP VCaP_R1881 GSE49091.GABPA.VCaP_R1881 188 bp overlap
GABPB1 2 datasets
ChIP K-562 ENCSR138YYY.GABPB1.K-562 269 bp overlap
ChIP K562 ENCFF015GDS 269 bp overlap
GATAD2B 1 dataset
ChIP HepG2 ENCFF829IBY 52 bp overlap
HBP1 1 dataset
ChIP Hep-G2 ENCSR616WEG.HBP1.Hep-G2 124 bp overlap
HIF1A 1 dataset
ChIP PC-3_siOC2 GSE106305.HIF1A.PC-3_siOC2 243 bp overlap
HSF1 1 dataset
ChIP MO91_CHX_10UM GSE45852.HSF1.MO91_CHX_10UM 143 bp overlap
IRF1 1 dataset
ChIP PDAC GSE64557.IRF1.PDAC 212 bp overlap
JUN 11 datasets
ChIP DE_D1 S08-DE-d1-JUN-exp1 269 bp overlap
ChIP DE_D1 S40-DE-d1-JUN-exp2 269 bp overlap
ChIP DE_D2 S02-DE-d2-JUN-exp1 269 bp overlap
ChIP DE_D2 S54-DE-d2-JUN-exp2 269 bp overlap
ChIP ESC S24-ESC-d0-JUN-exp1 269 bp overlap
ChIP ESC S34-ESC-d0-JUN-exp2 269 bp overlap
ChIP HUES-8 GSE109524.JUN.HUES-8 269 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.JUN.HUES-8_DE_ctrl 269 bp overlap
ChIP MDA-BoM-1833 GSE112444.JUN.MDA-BoM-1833 212 bp overlap
ChIP MDA-MB-231 GSE112444.JUN.MDA-MB-231 189 bp overlap
ChIP endothelial_umbilical-vein ENCSR000EFA.JUN.endothelial_umbilical-vein 201 bp overlap
KAT8 1 dataset
ChIP HepG2 ENCFF890JFC 269 bp overlap
KLF6 1 dataset
ChIP PDAC GSE64557.KLF6.PDAC 112 bp overlap
LMO1 1 dataset
ChIP Jurkat GSE94391.LMO1.Jurkat 214 bp overlap
MAF 1 dataset
ChIP CD4_Th1 GSE72266.MAF.CD4_Th1 161 bp overlap
MAX 1 dataset
ChIP Hep-G2 ENCSR168DYA.MAX.Hep-G2 161 bp overlap
MED1 1 dataset
ChIP MM1-S_JQ1_50NM GSE49224.MED1.MM1-S_JQ1_50NM 61 bp overlap
MED26 2 datasets
ChIP HCT-116 GSE121355.MED26.HCT-116 269 bp overlap
ChIP HCT-116_aux GSE121355.MED26.HCT-116_aux 178 bp overlap
MTERF4 2 datasets
ChIP HepG2 ENCFF831NAM 225 bp overlap
ChIP HepG2 ENCFF831NAM 269 bp overlap
MYB 1 dataset
ChIP Jurkat GSE59657.MYB.Jurkat 211 bp overlap
NANOG 3 datasets
ChIP HUES-8 GSE109524.NANOG.HUES-8 269 bp overlap
ChIP WA01 ERP004238.NANOG.WA01 183 bp overlap
ChIP WA09 GSE105028.NANOG.WA09 269 bp overlap
PGR 3 datasets
ChIP T-47D-A_E2_R5020 GSE80358.PGR.T-47D-A_E2_R5020 204 bp overlap
ChIP T-47D_R5020 GSE68355.PGR.T-47D_R5020 175 bp overlap
ChIP T-47D_R5020-MTVL GSE126859.PGR.T-47D_R5020-MTVL 166 bp overlap
PHIP 3 datasets
ChIP HCT-116_MLL1-KO__ab833 GSE101646.PHIP.HCT-116_MLL1-KO__ab833 269 bp overlap
ChIP HCT-116_ab833 GSE101646.PHIP.HCT-116_ab833 79 bp overlap
ChIP HCT-116_ab833 GSE101646.PHIP.HCT-116_ab833 128 bp overlap
POLR2A 2 datasets
ChIP HCT116 ENCFF508RDJ 269 bp overlap
ChIP endothelial cell of umbilical vein ENCFF303XUJ 269 bp overlap
POU2F1 1 dataset
ChIP HepG2 ENCFF422JZU 239 bp overlap
POU5F1 2 datasets
ChIP DE_D1 DED1-OCT4_Batch_II 269 bp overlap
ChIP HUES-8 GSE109524.POU5F1.HUES-8 224 bp overlap
POU5F1_M 1 dataset
ChIP DE_D1 DED1-OCT4-M_Batch_II 167 bp overlap
RBPJ 2 datasets
ChIP SCC_0h GSE156486.RBPJ.SCC_0h 250 bp overlap
ChIP SCC_4h GSE156486.RBPJ.SCC_4h 191 bp overlap
RELA 12 datasets
ChIP Detroit-562_LPS GSE91018.RELA.Detroit-562_LPS 224 bp overlap
ChIP Detroit-562_Pam2CSK4 GSE91018.RELA.Detroit-562_Pam2CSK4 269 bp overlap
ChIP Detroit-562_Poly-I GSE91018.RELA.Detroit-562_Poly-I 269 bp overlap
ChIP Detroit-562_TNFa GSE91018.RELA.Detroit-562_TNFa 269 bp overlap
ChIP Detroit-562_tri-DAP GSE91018.RELA.Detroit-562_tri-DAP 269 bp overlap
ChIP HUVEC-C_TNF-60min GSE121890.RELA.HUVEC-C_TNF-60min 219 bp overlap
ChIP aortic-endothelial-cell_IL1B_D14 GSE139377.RELA.aortic-endothelial-cell_IL1B_D14 206 bp overlap
ChIP aortic-endothelial-cell_IL1B_D16 GSE139377.RELA.aortic-endothelial-cell_IL1B_D16 161 bp overlap
ChIP aortic-endothelial-cell_IL1B_D21 GSE139377.RELA.aortic-endothelial-cell_IL1B_D21 234 bp overlap
ChIP aortic-endothelial-cell_IL1B_D40 GSE139377.RELA.aortic-endothelial-cell_IL1B_D40 237 bp overlap
ChIP aortic-endothelial-cell_IL1B_D7 GSE139377.RELA.aortic-endothelial-cell_IL1B_D7 146 bp overlap
ChIP aortic-endothelial-cell_IL1B_D9 GSE139377.RELA.aortic-endothelial-cell_IL1B_D9 150 bp overlap
SAP130 1 dataset
ChIP HepG2 ENCFF892EHZ 269 bp overlap
SIN3A 1 dataset
ChIP PFSK-1 ENCFF218MAY 240 bp overlap
SMAD2-3 2 datasets
ChIP HUES-8 GSE109524.SMAD2-3.HUES-8 269 bp overlap
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 269 bp overlap
SMAD3 4 datasets
ChIP BG03 GSE21614.SMAD3.BG03 220 bp overlap
ChIP BG03_DIFF_48H GSE36578.SMAD3.BG03_DIFF_48H 127 bp overlap
ChIP PC-3_hypoxia GSE106305.SMAD3.PC-3_hypoxia 195 bp overlap
ChIP WTC11 ENCFF815YYQ 254 bp overlap
SMARCA4 6 datasets
ChIP 22Rv1 GSE115615.SMARCA4.22Rv1 207 bp overlap
ChIP CTV-1_DSG GSE128835.SMARCA4.CTV-1_DSG 269 bp overlap
ChIP CTV-1_DSG_SPI1-mut GSE128835.SMARCA4.CTV-1_DSG_SPI1-mut 269 bp overlap
ChIP CTV-1_FLAG GSE128835.SMARCA4.CTV-1_FLAG 269 bp overlap
ChIP CTV-1_delQ GSE128835.SMARCA4.CTV-1_delQ 190 bp overlap
ChIP endometrial-epithelial-cells_KO GSE106660.SMARCA4.endometrial-epithelial-cells_KO 267 bp overlap
SMARCB1 1 dataset
ChIP hiPSC_WT GSE124903.SMARCB1.hiPSC_WT 170 bp overlap
SMARCC1 2 datasets
ChIP ESC S25-ESC-d0-BAF155-exp1 245 bp overlap
ChIP SK-N-MC GSE94275.SMARCC1.SK-N-MC 269 bp overlap
SMC1 1 dataset
ChIP HCAEC GSE101921.SMC1.HCAEC 269 bp overlap
SNAI2 1 dataset
ChIP keratinocyte_LacZ GSE55421.SNAI2.keratinocyte_LacZ 238 bp overlap
SP1 1 dataset
ChIP Hep-G2 ENCSR460YAM.SP1.Hep-G2 173 bp overlap
SP5 1 dataset
ChIP Hep-G2 ENCSR019NPF.SP5.Hep-G2 187 bp overlap
SPI1 3 datasets
ChIP CTV-1_FLAG_15perc GSE128835.SPI1.CTV-1_FLAG_15perc 269 bp overlap
ChIP CTV-1_delA GSE128835.SPI1.CTV-1_delA 203 bp overlap
ChIP CTV-1_delQ GSE128835.SPI1.CTV-1_delQ 269 bp overlap
STAG2 1 dataset
ChIP HCAEC GSE101921.STAG2.HCAEC 171 bp overlap
SUPT5H 2 datasets
ChIP HCT-116_DMSO_pThr806 GSE138548.SUPT5H.HCT-116_DMSO_pThr806 225 bp overlap
ChIP HCT-116_pThr806 GSE138548.SUPT5H.HCT-116_pThr806 269 bp overlap
TBP 1 dataset
ChIP hESC_8h GSE122298.TBP.hESC_8h 122 bp overlap
TCF12 1 dataset
ChIP Kasumi-1 GSE23730.TCF12.Kasumi-1 227 bp overlap
TFAP2A 2 datasets
ChIP MCF-7 GSE60270.TFAP2A.MCF-7 222 bp overlap
ChIP MCF-7_E2 GSE60270.TFAP2A.MCF-7_E2 209 bp overlap
TFAP2C 5 datasets
ChIP HCC1954 GSE126897.TFAP2C.HCC1954 238 bp overlap
ChIP MCF-7_E2 GSE23852.TFAP2C.MCF-7_E2 203 bp overlap
ChIP MCF-7_ETOH GSE23852.TFAP2C.MCF-7_ETOH 222 bp overlap
ChIP UCLA1-hESCs_TFAP2C--_1d_TFAP2C_Induc GSE101074.TFAP2C.UCLA1-hESCs_TFAP2C--_1d_TFAP2C_Induc 269 bp overlap
ChIP UCLA1-hESCs_TFAP2C--_2d_TFAP2C_Induc GSE101074.TFAP2C.UCLA1-hESCs_TFAP2C--_2d_TFAP2C_Induc 269 bp overlap
TP63 3 datasets
ChIP HCC95 GSE46837.TP63.HCC95 133 bp overlap
ChIP SUIT-2 GSE115461.TP63.SUIT-2 269 bp overlap
ChIP breast-organoid GSE113909.TP63.breast-organoid 89 bp overlap
TRIM28 1 dataset
ChIP hESC GSE115387.TRIM28.hESC 138 bp overlap
YY1 1 dataset
ChIP HCT-116 ENCSR000BNX.YY1.HCT-116 108 bp overlap
ZBTB38 1 dataset
ChIP HepG2 ENCFF875UQX 269 bp overlap
ZNF44 1 dataset
ChIP Hep-G2 ENCSR914HPP.ZNF44.Hep-G2 169 bp overlap
ZNF724 1 dataset
ChIP HepG2 ENCFF318TJD 61 bp overlap