chr14 : 54,490,562 54,490,737
175 bp 72 TFs 1 linked gene
This 175 bp open chromatin element is linked to GMFB and is bound by 72 transcription factors.
Linked Genes
1 gene
Gene Expression Dist. to TSS Distance Link type
GMFB 1.5 kb Proximal Proximity
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr14:54,485,562 – 54,495,737
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
72 transcription factors
Source
Cell type
AR 17 datasets
ChIP LNCaP GSE110655.AR.LNCaP 173 bp overlap
ChIP LNCaP-95_NT GSE125552.AR.LNCaP-95_NT 175 bp overlap
ChIP LNCaP-clone-FGC_DHT-DMSO GSE118247.AR.LNCaP-clone-FGC_DHT-DMSO 175 bp overlap
ChIP LNCaP_ETOH GSE69043.AR.LNCaP_ETOH 149 bp overlap
ChIP LNCaP_R1881 GSE69043.AR.LNCaP_R1881 175 bp overlap
ChIP LNCaP_SHGATA2_ETOH GSE69043.AR.LNCaP_SHGATA2_ETOH 137 bp overlap
ChIP LNCaP_SHGATA2_R1881 GSE69043.AR.LNCaP_SHGATA2_R1881 175 bp overlap
ChIP LNCaP_androgen-N_hypoxia-N GSE114732.AR.LNCaP_androgen-N_hypoxia-N 103 bp overlap
ChIP MDA-MB-453 ERP001226.AR.MDA-MB-453 152 bp overlap
ChIP MDA-MB-453_FOXA1 ERP003503.AR.MDA-MB-453_FOXA1 133 bp overlap
ChIP MDA-MB-453_R1881_SICTR GSE70161.AR.MDA-MB-453_R1881_SICTR 119 bp overlap
ChIP MDA-MB-453_R1881_SIPIAS1 GSE70161.AR.MDA-MB-453_R1881_SIPIAS1 143 bp overlap
ChIP prostate-cancer GSE136128.AR.prostate-cancer 175 bp overlap
ChIP prostate-cancer_C4-2-CON GSE136128.AR.prostate-cancer_C4-2-CON 149 bp overlap
ChIP prostate-cancer_PDX_167 GSE130408.AR.prostate-cancer_PDX_167 100 bp overlap
ChIP prostate-cancer_PDX_70 GSE130408.AR.prostate-cancer_PDX_70 83 bp overlap
ChIP prostate-cancer_PDX_78 GSE130408.AR.prostate-cancer_PDX_78 77 bp overlap
ARID1A 4 datasets
ChIP MCF-7 GSE123284.ARID1A.MCF-7 160 bp overlap
ChIP MCF-7_4-hydroxytamoxifen GSE123284.ARID1A.MCF-7_4-hydroxytamoxifen 175 bp overlap
ChIP MCF-7_DCDT GSE123284.ARID1A.MCF-7_DCDT 175 bp overlap
ChIP MCF-7_Fulvestrant GSE123284.ARID1A.MCF-7_Fulvestrant 175 bp overlap
ARID1B 2 datasets
ChIP K-562 ENCSR822CCM.ARID1B.K-562 139 bp overlap
ChIP K562 ENCFF938UXQ 139 bp overlap
ARID2 1 dataset
ChIP BIN-67_ctrl GSE117734.ARID2.BIN-67_ctrl 175 bp overlap
ARNT 1 dataset
ChIP K-562 ENCSR613NUC.ARNT.K-562 175 bp overlap
ATF1 1 dataset
ChIP K-562 ENCSR091GVJ.ATF1.K-562 175 bp overlap
BRD4 1 dataset
ChIP T-47D_DMSO GSE63581.BRD4.T-47D_DMSO 175 bp overlap
CTBP1 1 dataset
ChIP K-562 ENCSR201NQZ.CTBP1.K-562 175 bp overlap
DPF2 5 datasets
ChIP BIN-67_ctrl GSE117734.DPF2.BIN-67_ctrl 175 bp overlap
ChIP BIN-67_lentivirus-SMARCA4-K785R GSE117734.DPF2.BIN-67_lentivirus-SMARCA4-K785R 175 bp overlap
ChIP BIN-67_lentivirus-SMARCA4-T910M GSE117734.DPF2.BIN-67_lentivirus-SMARCA4-T910M 175 bp overlap
ChIP K-562 ENCSR219BXP.DPF2.K-562 171 bp overlap
ChIP MCF-7 ENCFF712EXQ 175 bp overlap
E4F1 1 dataset
ChIP K-562 ENCSR731LHZ.E4F1.K-562 175 bp overlap
ELF1 6 datasets
ChIP MCF-7 ENCFF305BNP 175 bp overlap
ChIP MCF-7 ENCFF687CWI 175 bp overlap
ChIP MCF-7 ENCSR502NRF.ELF1.MCF-7 163 bp overlap
ChIP MCF-7 ENCSR000BSS.ELF1.MCF-7 170 bp overlap
ChIP SK-N-MC_SHFLI_96H GSE61944.ELF1.SK-N-MC_SHFLI_96H 151 bp overlap
ChIP SK-N-MC_SHGFP_96H GSE61944.ELF1.SK-N-MC_SHGFP_96H 168 bp overlap
ELF3 2 datasets
ChIP PDAC GSE64557.ELF3.PDAC 175 bp overlap
ChIP PDAC_SHCTR GSE64557.ELF3.PDAC_SHCTR 175 bp overlap
EOMES 1 dataset
ChIP hESC GSE26097.EOMES.hESC 132 bp overlap
ERG 3 datasets
ChIP TSU-1621MT_ATRA GSE60477.ERG.TSU-1621MT_ATRA 151 bp overlap
ChIP VCaP GSE28950.ERG.VCaP 150 bp overlap
ChIP VCaP_shARID1A GSE110655.ERG.VCaP_shARID1A 160 bp overlap
ESR1 5 datasets
ChIP MCF-7_E2+4OHT_SRC-3 GSE119702.ESR1.MCF-7_E2+4OHT_SRC-3 175 bp overlap
ChIP MCF-7_SRC3_OHT GSE119702.ESR1.MCF-7_SRC3_OHT 175 bp overlap
ChIP T-47D GSE148277.ESR1.T-47D 175 bp overlap
ChIP T-47D ENCSR000BKN.ESR1.T-47D 146 bp overlap
ChIP ZR751 ERP000783.ESR1.ZR751 137 bp overlap
ESRRA 1 dataset
ChIP BT-474 GSE81651.ESRRA.BT-474 175 bp overlap
ETS1 1 dataset
ChIP CTV-1_DSG GSE128835.ETS1.CTV-1_DSG 175 bp overlap
ETV1 2 datasets
ChIP LNCaP GSE47120.ETV1.LNCaP 133 bp overlap
ChIP MDA-Pca-2b GSE106624.ETV1.MDA-Pca-2b 109 bp overlap
FLI1 6 datasets
ChIP A-673_1_KRAB_GFP GSE106914.FLI1.A-673_1_KRAB_GFP 175 bp overlap
ChIP A-673_2_KRAB_eSOX2_1 GSE106914.FLI1.A-673_2_KRAB_eSOX2_1 175 bp overlap
ChIP A-673_D11 GSE129155.FLI1.A-673_D11 175 bp overlap
ChIP A-673_D14 GSE129155.FLI1.A-673_D14 175 bp overlap
ChIP A-673_D17 GSE129155.FLI1.A-673_D17 175 bp overlap
ChIP CTV-1_DSG GSE128835.FLI1.CTV-1_DSG 175 bp overlap
FOXA1 3 datasets
ChIP LNCaP GSE64656.FOXA1.LNCaP 62 bp overlap
ChIP LNCaP_DHT24H GSE58428.FOXA1.LNCaP_DHT24H 175 bp overlap
ChIP VCaP_ETOH24H GSE58428.FOXA1.VCaP_ETOH24H 175 bp overlap
FOXA2 1 dataset
ChIP DE DE-FOXA2-2 73 bp overlap
GABPA 2 datasets
ChIP MCF-7 ENCFF735CHO 175 bp overlap
ChIP MCF-7 ENCSR000BUK.GABPA.MCF-7 130 bp overlap
GATA1 5 datasets
ChIP HUDEP-2_30min GSE104676.GATA1.HUDEP-2_30min 89 bp overlap
ChIP K-562 GSE107726.GATA1.K-562 175 bp overlap
ChIP K-562_dCas9-LSD1 GSE132212.GATA1.K-562_dCas9-LSD1 116 bp overlap
ChIP K-562_enCRISPRi-LK GSE132212.GATA1.K-562_enCRISPRi-LK 132 bp overlap
ChIP erythroid_Don001 GSE137982.GATA1.erythroid_Don001 96 bp overlap
GATA2 17 datasets
ChIP ESF GSE108408.GATA2.ESF 175 bp overlap
ChIP K-562 ENCSR000DKA.GATA2.K-562 164 bp overlap
ChIP K-562 ENCSR000EWG.GATA2.K-562 146 bp overlap
ChIP LNCaP GSE38391.GATA2.LNCaP 175 bp overlap
ChIP LNCaP GSE52725.GATA2.LNCaP 175 bp overlap
ChIP LNCaP_CSFCS GSE69043.GATA2.LNCaP_CSFCS 175 bp overlap
ChIP LNCaP_FBS GSE69043.GATA2.LNCaP_FBS 175 bp overlap
ChIP SH-SY5Y ENCFF485YIB 175 bp overlap
ChIP SH-SY5Y ENCSR000EYB.GATA2.SH-SY5Y 171 bp overlap
ChIP SKH1 GSE87283.GATA2.SKH1 175 bp overlap
ChIP SKH1_RUNX1-EVI1_KD GSE87283.GATA2.SKH1_RUNX1-EVI1_KD 175 bp overlap
ChIP TF1 GSE73207.GATA2.TF1 175 bp overlap
ChIP TSU-1621MT GSE60477.GATA2.TSU-1621MT 153 bp overlap
ChIP VCaP GSE125236.GATA2.VCaP 175 bp overlap
ChIP VCaP_JQ1 GSE125236.GATA2.VCaP_JQ1 175 bp overlap
ChIP endothelial cell of umbilical vein ENCFF148NLK 175 bp overlap
ChIP endothelial_umbilical-vein ENCSR000EVW.GATA2.endothelial_umbilical-vein 175 bp overlap
GATA3 25 datasets
ChIP A1A3_Brg1KD_Dex GSE112491.GATA3.A1A3_Brg1KD_Dex 142 bp overlap
ChIP A1A3_Dex GSE112491.GATA3.A1A3_Dex 172 bp overlap
ChIP Jurkat GSE29180.GATA3.Jurkat 117 bp overlap
ChIP Jurkat GSE120063.GATA3.Jurkat 175 bp overlap
ChIP Jurkat GSE76181.GATA3.Jurkat 175 bp overlap
ChIP MCF-7 ENCFF352QVM 175 bp overlap
ChIP MCF-7 ENCFF437NQS 175 bp overlap
ChIP MCF-7 ENCFF695MAU 175 bp overlap
ChIP MCF-7 GSE128445.GATA3.MCF-7 175 bp overlap
ChIP MCF-7 GSE122847.GATA3.MCF-7 175 bp overlap
ChIP MCF-7 GSE51274.GATA3.MCF-7 175 bp overlap
ChIP MCF-7 GSE117492.GATA3.MCF-7 162 bp overlap
ChIP MCF-7 GSE133072.GATA3.MCF-7 175 bp overlap
ChIP MCF-7_DMSO GSE29073.GATA3.MCF-7_DMSO 153 bp overlap
ChIP MCF-7_E2 GSE29073.GATA3.MCF-7_E2 123 bp overlap
ChIP MCF-7_E2 GSE40129.GATA3.MCF-7_E2 111 bp overlap
ChIP MCF-7_E2 GSE81510.GATA3.MCF-7_E2 149 bp overlap
ChIP MCF-7_E2 GSE60270.GATA3.MCF-7_E2 175 bp overlap
ChIP MCF-7_E2_Dex GSE81510.GATA3.MCF-7_E2_Dex 119 bp overlap
ChIP MCF-7_sgScr GSE133072.GATA3.MCF-7_sgScr 175 bp overlap
ChIP SK-N-SH ENCFF040SSB 108 bp overlap
ChIP T-47D GSE51274.GATA3.T-47D 175 bp overlap
ChIP T-47D ENCSR000BMX.GATA3.T-47D 175 bp overlap
ChIP T-47D_sc GSE122847.GATA3.T-47D_sc 151 bp overlap
ChIP thymocyte GSE71751.GATA3.thymocyte 138 bp overlap
GATA3_Cter 2 datasets
ChIP T-47D_CR3flp GSE99479.GATA3_Cter.T-47D_CR3flp 166 bp overlap
ChIP T-47D_flp-ctrl GSE99479.GATA3_Cter.T-47D_flp-ctrl 147 bp overlap
GATA3_Nter 2 datasets
ChIP T-47D_CR3flp GSE99479.GATA3_Nter.T-47D_CR3flp 175 bp overlap
ChIP T-47D_flp-ctrl GSE99479.GATA3_Nter.T-47D_flp-ctrl 175 bp overlap
GATA4 3 datasets
ChIP DE DE-GATA4-1 175 bp overlap
ChIP DE DE-GATA4-2 175 bp overlap
ChIP foregut GSE117136.GATA4.foregut 108 bp overlap
GATA6 10 datasets
ChIP DE DE-GATA6-1 175 bp overlap
ChIP DE DE-GATA6-2 175 bp overlap
ChIP DE_D1 S09-DE-d1-GATA6-exp1 175 bp overlap
ChIP DE_D1 S41-DE-d1-GATA6-exp2 153 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 175 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 165 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 175 bp overlap
ChIP OACP4-C GSE132680.GATA6.OACP4-C 175 bp overlap
ChIP YCC-3 GSE51705.GATA6.YCC-3 125 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 175 bp overlap
GLIS1 1 dataset
ChIP HEK293 ENCSR482BBZ.GLIS1.HEK293 175 bp overlap
GRHL1 2 datasets
ChIP MCF-7 GSE140185.GRHL1.MCF-7 161 bp overlap
ChIP MCF-7_ARID1A-KO GSE140185.GRHL1.MCF-7_ARID1A-KO 168 bp overlap
GRHL2 6 datasets
ChIP LNCaP GSE80256.GRHL2.LNCaP 175 bp overlap
ChIP MCF-7 GSE109820.GRHL2.MCF-7 175 bp overlap
ChIP MCF-7-WS8_VEH GSE113092.GRHL2.MCF-7-WS8_VEH 175 bp overlap
ChIP MCF-7_E2_45min GSE109820.GRHL2.MCF-7_E2_45min 164 bp overlap
ChIP OVCA429 GSE71018.GRHL2.OVCA429 151 bp overlap
ChIP T-47D GSE99680.GRHL2.T-47D 162 bp overlap
HDAC1 2 datasets
ChIP K-562 ENCSR387UWP.HDAC1.K-562 164 bp overlap
ChIP MCF-7_parental_4-hydroxytamoxifen GSE123284.HDAC1.MCF-7_parental_4-hydroxytamoxifen 175 bp overlap
HDAC2 2 datasets
ChIP K-562 ENCSR893WSB.HDAC2.K-562 165 bp overlap
ChIP MCF-7 ENCSR000BTP.HDAC2.MCF-7 123 bp overlap
HOXB13 9 datasets
ChIP LNCaP GSE56288.HOXB13.LNCaP 175 bp overlap
ChIP LNCaP GSE96652.HOXB13.LNCaP 89 bp overlap
ChIP prostate-cancer_PDX_141 GSE130408.HOXB13.prostate-cancer_PDX_141 92 bp overlap
ChIP prostate-cancer_PDX_167 GSE130408.HOXB13.prostate-cancer_PDX_167 175 bp overlap
ChIP prostate-cancer_PDX_70 GSE130408.HOXB13.prostate-cancer_PDX_70 103 bp overlap
ChIP prostate-cancer_PDX_92 GSE130408.HOXB13.prostate-cancer_PDX_92 69 bp overlap
ChIP prostate_P23_T GSE130408.HOXB13.prostate_P23_T 165 bp overlap
ChIP prostate_P29_T GSE130408.HOXB13.prostate_P29_T 175 bp overlap
ChIP prostate_P5_T GSE130408.HOXB13.prostate_P5_T 175 bp overlap
IKZF1 3 datasets
ChIP K-562 ENCSR395HWC.IKZF1.K-562 175 bp overlap
ChIP K562 ENCFF348IBL 150 bp overlap
ChIP K562 ENCFF771OHZ 123 bp overlap
KDM5B 3 datasets
ChIP HCC2157 GSE46055.KDM5B.HCC2157 133 bp overlap
ChIP SUM185 GSE46055.KDM5B.SUM185 66 bp overlap
ChIP SUM185_SHCTCF GSE46055.KDM5B.SUM185_SHCTCF 96 bp overlap
KMT2C 2 datasets
ChIP BIN-67_ctrl GSE117734.KMT2C.BIN-67_ctrl 175 bp overlap
ChIP BIN-67_lentivirus-SMARCA4 GSE117734.KMT2C.BIN-67_lentivirus-SMARCA4 175 bp overlap
KMT2D 3 datasets
ChIP BIN-67_ctrl GSE117734.KMT2D.BIN-67_ctrl 175 bp overlap
ChIP BIN-67_lentivirus-SMARCA4 GSE117734.KMT2D.BIN-67_lentivirus-SMARCA4 136 bp overlap
ChIP BIN-67_lentivirus-SMARCA4-T910M GSE117734.KMT2D.BIN-67_lentivirus-SMARCA4-T910M 116 bp overlap
LEF1 1 dataset
ChIP K-562 ENCSR343ELW.LEF1.K-562 75 bp overlap
MTA3 1 dataset
ChIP K-562 ENCSR914NEI.MTA3.K-562 65 bp overlap
MYBL2 1 dataset
ChIP A-673 GSE119971.MYBL2.A-673 135 bp overlap
NANOG 1 dataset
ChIP LNCaP_pNanog8_Dox GSE74799.NANOG.LNCaP_pNanog8_Dox 84 bp overlap
NR2F1 1 dataset
ChIP K-562 ENCSR970NKQ.NR2F1.K-562 175 bp overlap
NR2F2 2 datasets
ChIP K-562 ENCSR000BRS.NR2F2.K-562 92 bp overlap
ChIP MCF-7 ENCSR000BUY.NR2F2.MCF-7 158 bp overlap
NR3C1 2 datasets
ChIP MDA-MB-453 GSE152203.NR3C1.MDA-MB-453 162 bp overlap
ChIP T47D-A1-2_Dex GSE112491.NR3C1.T47D-A1-2_Dex 158 bp overlap
PHIP 1 dataset
ChIP HCT-116_ab833 GSE101646.PHIP.HCT-116_ab833 72 bp overlap
POU5F1 1 dataset
ChIP hESC_NAIVE GSE69646.POU5F1.hESC_NAIVE 144 bp overlap
RCOR1 1 dataset
ChIP keratinocyte_diff GSE57702.RCOR1.keratinocyte_diff 175 bp overlap
RELA 2 datasets
ChIP LNCaP_SIFOXA1_TNFA GSE83860.RELA.LNCaP_SIFOXA1_TNFA 109 bp overlap
ChIP aortic-endothelial-cell_IL1B_D21 GSE139377.RELA.aortic-endothelial-cell_IL1B_D21 130 bp overlap
SMAD3 1 dataset
ChIP HCC1954 GSE104760.SMAD3.HCC1954 175 bp overlap
SMARCA2 1 dataset
ChIP SK-N-MC_shEWSFLI1 GSE94275.SMARCA2.SK-N-MC_shEWSFLI1 175 bp overlap
SMARCA4 7 datasets
ChIP BIN-67_lentivirus-SMARCA4 GSE117734.SMARCA4.BIN-67_lentivirus-SMARCA4 165 bp overlap
ChIP CTV-1_DSG GSE128835.SMARCA4.CTV-1_DSG 175 bp overlap
ChIP CTV-1_FLAG GSE128835.SMARCA4.CTV-1_FLAG 175 bp overlap
ChIP CTV-1_delQ GSE128835.SMARCA4.CTV-1_delQ 142 bp overlap
ChIP K-562 ENCSR587OQL.SMARCA4.K-562 136 bp overlap
ChIP MCF-7 GSE140185.SMARCA4.MCF-7 175 bp overlap
ChIP MCF-7_parental GSE123284.SMARCA4.MCF-7_parental 167 bp overlap
SMARCB1 1 dataset
ChIP MCF-7_Fulvestrant GSE123284.SMARCB1.MCF-7_Fulvestrant 95 bp overlap
SMARCC1 6 datasets
ChIP BIN-67_ctrl GSE117734.SMARCC1.BIN-67_ctrl 175 bp overlap
ChIP BIN-67_lentivirus-SMARCA4 GSE117734.SMARCC1.BIN-67_lentivirus-SMARCA4 62 bp overlap
ChIP BIN-67_lentivirus-SMARCA4-K785R GSE117734.SMARCC1.BIN-67_lentivirus-SMARCA4-K785R 157 bp overlap
ChIP BIN-67_lentivirus-SMARCA4-T910M GSE117734.SMARCC1.BIN-67_lentivirus-SMARCA4-T910M 175 bp overlap
ChIP MCF-7 GSE124225.SMARCC1.MCF-7 155 bp overlap
ChIP SK-N-MC GSE94275.SMARCC1.SK-N-MC 175 bp overlap
SMARCE1 3 datasets
ChIP K-562 ENCSR157TCS.SMARCE1.K-562 175 bp overlap
ChIP MCF-7 ENCFF890MHF 175 bp overlap
ChIP MCF-7 ENCSR431TLD.SMARCE1.MCF-7 142 bp overlap
SPI1 4 datasets
ChIP CTV-1_FLAG_15perc GSE128835.SPI1.CTV-1_FLAG_15perc 153 bp overlap
ChIP CTV-1_delQ GSE128835.SPI1.CTV-1_delQ 152 bp overlap
ChIP DC_96h_donorM GSE128834.SPI1.DC_96h_donorM 130 bp overlap
ChIP THP-1 GSE128834.SPI1.THP-1 117 bp overlap
SS18 3 datasets
ChIP BIN-67_lentivirus-SMARCA4 GSE117734.SS18.BIN-67_lentivirus-SMARCA4 164 bp overlap
ChIP BIN-67_lentivirus-SMARCA4-K785R GSE117734.SS18.BIN-67_lentivirus-SMARCA4-K785R 110 bp overlap
ChIP BIN-67_lentivirus-SMARCA4-T910M GSE117734.SS18.BIN-67_lentivirus-SMARCA4-T910M 175 bp overlap
STAT3 3 datasets
ChIP HCC70_EtOH GSE85579.STAT3.HCC70_EtOH 125 bp overlap
ChIP MDA-MB-453 GSE152203.STAT3.MDA-MB-453 175 bp overlap
ChIP T-47D_JC5056 GSE126004.STAT3.T-47D_JC5056 164 bp overlap
TFAP2A 2 datasets
ChIP MCF-7_E2 GSE60270.TFAP2A.MCF-7_E2 175 bp overlap
ChIP WA09 GSE105081.TFAP2A.WA09 145 bp overlap
TFAP2B 1 dataset
ChIP SK-N-SH ENCFF869XXQ 175 bp overlap
TFAP2C 1 dataset
ChIP HCC1954 GSE126897.TFAP2C.HCC1954 157 bp overlap
TLE3 2 datasets
ChIP LNCaP GSE94682.TLE3.LNCaP 112 bp overlap
ChIP LNCaP_r1881 GSE94682.TLE3.LNCaP_r1881 167 bp overlap
TRPS1 1 dataset
ChIP T-47D GSE107013.TRPS1.T-47D 108 bp overlap
ZBTB2 1 dataset
ChIP K-562 ENCSR230PTV.ZBTB2.K-562 175 bp overlap
ZBTB7A 1 dataset
ChIP K-562 GSE103445.ZBTB7A.K-562 143 bp overlap
ZNF750 1 dataset
ChIP keratinocyte_diff GSE57702.ZNF750.keratinocyte_diff 175 bp overlap