chr12 : 88,942,685 88,944,156
1,471 bp 122 TFs 2 linked genes
This 1.5 kb open chromatin element is linked to TMTC3 and CEP290 and is bound by 122 transcription factors.
Linked Genes
2 genes
Gene Expression Dist. to TSS Distance Link type
TMTC3 801.2 kb Distal Multiome+HiCAR
CEP290 801.5 kb Distal Multiome+HiCAR
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr12:88,937,685 – 88,949,156
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
122 transcription factors
Source
Cell type
AR 2 datasets
ChIP LNCaP_1F5_SIFOXA1 GSE30623.AR.LNCaP_1F5_SIFOXA1 126 bp overlap
ChIP MCF-7 GSE48930.AR.MCF-7 167 bp overlap
ARGFX 2 datasets
Motif DE_48h DE_48h-ARGFX_MA1463.2 8 bp overlap
Motif DE_60h DE_60h-ARGFX_MA1463.2 8 bp overlap
Arx 2 datasets
Motif DE_48h DE_48h-Arx_MA0874.2 10 bp overlap
Motif DE_60h DE_60h-Arx_MA0874.2 10 bp overlap
BARHL1 1 dataset
Motif DE_60h DE_60h-BARHL1_MA0877.4 6 bp overlap
BARHL2 1 dataset
Motif DE_60h DE_60h-BARHL2_MA0635.2 6 bp overlap
BARX1 2 datasets
Motif DE_48h DE_48h-BARX1_MA0875.2 6 bp overlap
Motif DE_60h DE_60h-BARX1_MA0875.2 6 bp overlap
BRD2 1 dataset
ChIP MDA-MB-231_NTsiRNA GSE116879.BRD2.MDA-MB-231_NTsiRNA 84 bp overlap
BRD4 3 datasets
ChIP HCT-116 GSE57628.BRD4.HCT-116 180 bp overlap
ChIP PC-3_GDC-R GSE137207.BRD4.PC-3_GDC-R 52 bp overlap
ChIP PC-3_GDC-R_SAHA GSE137207.BRD4.PC-3_GDC-R_SAHA 63 bp overlap
BSX 2 datasets
Motif DE_48h DE_48h-BSX_MA0876.2 6 bp overlap
Motif DE_60h DE_60h-BSX_MA0876.2 6 bp overlap
CBX3 2 datasets
ChIP HCT116 ENCFF947BOL 431 bp overlap
ChIP HCT116 ENCFF947BOL 431 bp overlap
CDK8 1 dataset
ChIP HCT-116_NOMO GSE38258.CDK8.HCT-116_NOMO 231 bp overlap
CREBBP 1 dataset
ChIP PC-3_GDC-resist GSE147455.CREBBP.PC-3_GDC-resist 124 bp overlap
DLX1 2 datasets
Motif DE_48h DE_48h-DLX1_MA0879.3 6 bp overlap
Motif DE_60h DE_60h-DLX1_MA0879.3 6 bp overlap
DLX6 2 datasets
Motif DE_48h DE_48h-DLX6_MA0882.2 6 bp overlap
Motif DE_60h DE_60h-DLX6_MA0882.2 6 bp overlap
Dlx2 2 datasets
Motif DE_48h DE_48h-Dlx2_MA0885.3 8 bp overlap
Motif DE_60h DE_60h-Dlx2_MA0885.3 8 bp overlap
Dlx3 2 datasets
Motif DE_48h DE_48h-Dlx3_MA0880.2 6 bp overlap
Motif DE_60h DE_60h-Dlx3_MA0880.2 6 bp overlap
Dlx4 2 datasets
Motif DE_48h DE_48h-Dlx4_MA0881.2 6 bp overlap
Motif DE_60h DE_60h-Dlx4_MA0881.2 6 bp overlap
Dlx5 2 datasets
Motif DE_48h DE_48h-Dlx5_MA1476.3 8 bp overlap
Motif DE_60h DE_60h-Dlx5_MA1476.3 8 bp overlap
EOMES 2 datasets
ChIP hESC GSE26097.EOMES.hESC 490 bp overlap
ChIP hESC GSE26097.EOMES.hESC 228 bp overlap
EP300 2 datasets
ChIP PC-3 GSE147455.EP300.PC-3 156 bp overlap
ChIP PC-3 GSE147455.EP300.PC-3 110 bp overlap
ESR1 1 dataset
ChIP MCF-7_E2 GSE14664.ESR1.MCF-7_E2 169 bp overlap
Elf5 1 dataset
Motif DE_60h DE_60h-Elf5_MA0136.4 8 bp overlap
FIGLA 1 dataset
Motif DE_60h DE_60h-FIGLA_MA0820.2 6 bp overlap
FOS 1 dataset
ChIP CFPAC-1 GSE119930.FOS.CFPAC-1 171 bp overlap
FOSL1 1 dataset
ChIP K562 ENCFF455MKD 53 bp overlap
FOXA1 1 dataset
ChIP NCI-H3122 GSE39998.FOXA1.NCI-H3122 162 bp overlap
FOXA2 3 datasets
ChIP DE DE-FOXA2-1 1160 bp overlap
ChIP DE DE-FOXA2-2 1254 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.FOXA2.pancreatic-progenitor_PP1 286 bp overlap
FOXB1 1 dataset
Motif DE_60h DE_60h-FOXB1_MA0845.1 11 bp overlap
FOXC1 1 dataset
Motif DE_60h DE_60h-FOXC1_MA0032.2 11 bp overlap
FOXC2 1 dataset
Motif DE_60h DE_60h-FOXC2_MA0846.2 11 bp overlap
FOXL2 2 datasets
ChIP HGrC1_C134W-TGF GSE138496.FOXL2.HGrC1_C134W-TGF 131 bp overlap
ChIP HGrC1_C134W-TGF_parental GSE138496.FOXL2.HGrC1_C134W-TGF_parental 303 bp overlap
FOXP2 1 dataset
Motif DE_60h DE_60h-FOXP2_MA0593.2 9 bp overlap
GATA2 6 datasets
Motif DE_48h DE_48h-GATA2_MA0036.4 7 bp overlap
Motif DE_60h DE_60h-GATA2_MA0036.4 7 bp overlap
Motif DE_72h DE_72h-GATA2_MA0036.4 7 bp overlap
ChIP primary-endometrial-stromal-cell_IVD_P2 GSE145673.GATA2.primary-endometrial-stromal-cell_IVD_P2 259 bp overlap
ChIP primary-endometrial-stromal-cell_Veh_P1 GSE145673.GATA2.primary-endometrial-stromal-cell_Veh_P1 490 bp overlap
ChIP primary-endometrial-stromal-cell_Veh_P2 GSE145673.GATA2.primary-endometrial-stromal-cell_Veh_P2 378 bp overlap
GATA3 1 dataset
ChIP Jurkat GSE76181.GATA3.Jurkat 278 bp overlap
GATA4 8 datasets
ChIP DE DE-GATA4-1 1324 bp overlap
ChIP DE DE-GATA4-2 1458 bp overlap
Motif DE_48h DE_48h-GATA4_MA0482.3 8 bp overlap
Motif DE_60h DE_60h-GATA4_MA0482.3 8 bp overlap
Motif DE_72h DE_72h-GATA4_MA0482.3 8 bp overlap
ChIP foregut GSE117136.GATA4.foregut 560 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA4.pancreatic-progenitor 468 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA4.pancreatic-progenitor_PP1 490 bp overlap
GATA5 4 datasets
Motif DE_48h DE_48h-GATA5_MA0766.3 8 bp overlap
Motif DE_60h DE_60h-GATA5_MA0766.3 8 bp overlap
Motif DE_60h DE_60h-GATA5_MA0766.3 8 bp overlap
Motif DE_72h DE_72h-GATA5_MA0766.3 8 bp overlap
GATA6 16 datasets
ChIP DE DE-GATA6-1 1350 bp overlap
ChIP DE DE-GATA6-2 1318 bp overlap
Motif DE_48h DE_48h-GATA6_MA1104.3 8 bp overlap
Motif DE_60h DE_60h-GATA6_MA1104.3 8 bp overlap
Motif DE_72h DE_72h-GATA6_MA1104.3 8 bp overlap
ChIP DE_D1 S09-DE-d1-GATA6-exp1 414 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 1145 bp overlap
ChIP DE_D1 S41-DE-d1-GATA6-exp2 480 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 1354 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 1269 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 1047 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 1190 bp overlap
ChIP foregut GSE117136.GATA6.foregut 311 bp overlap
ChIP foregut GSE117136.GATA6.foregut 472 bp overlap
ChIP foregut_KO GSE117136.GATA6.foregut_KO 299 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA6.pancreatic-progenitor 384 bp overlap
GBX2 2 datasets
Motif DE_48h DE_48h-GBX2_MA0890.2 6 bp overlap
Motif DE_60h DE_60h-GBX2_MA0890.2 6 bp overlap
GRHL2 1 dataset
ChIP LNCaP GSE80256.GRHL2.LNCaP 252 bp overlap
Gata3 3 datasets
Motif DE_48h DE_48h-Gata3_MA0037.5 8 bp overlap
Motif DE_60h DE_60h-Gata3_MA0037.5 8 bp overlap
Motif DE_72h DE_72h-Gata3_MA0037.5 8 bp overlap
HDAC2 2 datasets
ChIP PC-3 GSE147455.HDAC2.PC-3 126 bp overlap
ChIP PC-3_GDC-resist GSE147455.HDAC2.PC-3_GDC-resist 96 bp overlap
HESX1 2 datasets
Motif DE_48h DE_48h-HESX1_MA0894.2 6 bp overlap
Motif DE_60h DE_60h-HESX1_MA0894.2 6 bp overlap
HNF4A 1 dataset
Motif DE_60h DE_60h-HNF4A_MA1494.2 14 bp overlap
HOXA7 2 datasets
Motif DE_48h DE_48h-HOXA7_MA1498.3 6 bp overlap
Motif DE_60h DE_60h-HOXA7_MA1498.3 6 bp overlap
Hoxa13 1 dataset
Motif DE_60h DE_60h-Hoxa13_MA0650.4 8 bp overlap
IKZF2 1 dataset
Motif DE_60h DE_60h-IKZF2_MA2326.1 6 bp overlap
JUN 1 dataset
ChIP DE_D2 S02-DE-d2-JUN-exp1 276 bp overlap
JUNB 1 dataset
ChIP CFPAC-1 GSE119930.JUNB.CFPAC-1 331 bp overlap
LBX2 2 datasets
Motif DE_48h DE_48h-LBX2_MA0699.2 6 bp overlap
Motif DE_60h DE_60h-LBX2_MA0699.2 6 bp overlap
LHX2 3 datasets
Motif DE_48h DE_48h-LHX2_MA0700.3 6 bp overlap
Motif DE_60h DE_60h-LHX2_MA0700.3 6 bp overlap
ChIP retina_pigment GSE60024.LHX2.retina_pigment 312 bp overlap
MAX 1 dataset
ChIP HCT116 ENCFF810LEN 497 bp overlap
MED1 5 datasets
ChIP HCT-116 GSE121798.MED1.HCT-116 971 bp overlap
ChIP HCT-116_shLuc GSE121798.MED1.HCT-116_shLuc 1203 bp overlap
ChIP HCT-116_shLuc_thaps GSE121798.MED1.HCT-116_shLuc_thaps 284 bp overlap
ChIP HCT-116_shLuc_thaps GSE121798.MED1.HCT-116_shLuc_thaps 332 bp overlap
ChIP HCT-116_thaps GSE121798.MED1.HCT-116_thaps 914 bp overlap
MED26 1 dataset
ChIP HCT-116_aux GSE121355.MED26.HCT-116_aux 61 bp overlap
MEF2C 3 datasets
Motif DE_48h DE_48h-MEF2C_MA0497.2 11 bp overlap
Motif DE_60h DE_60h-MEF2C_MA0497.2 11 bp overlap
Motif DE_72h DE_72h-MEF2C_MA0497.2 11 bp overlap
MEIS1 1 dataset
Motif DE_60h DE_60h-MEIS1_MA0498.3 5 bp overlap
MITF 4 datasets
ChIP 501-mel GSE137522.MITF.501-mel 455 bp overlap
ChIP 501-mel GSE61965.MITF.501-mel 261 bp overlap
ChIP 501-mel_20ng_K243R GSE137522.MITF.501-mel_20ng_K243R 382 bp overlap
ChIP 501-mel_K243Q GSE137522.MITF.501-mel_K243Q 411 bp overlap
MSX1 2 datasets
Motif DE_48h DE_48h-MSX1_MA0666.3 6 bp overlap
Motif DE_60h DE_60h-MSX1_MA0666.3 6 bp overlap
MSX2 2 datasets
Motif DE_48h DE_48h-MSX2_MA0708.3 6 bp overlap
Motif DE_60h DE_60h-MSX2_MA0708.3 6 bp overlap
MTF1 1 dataset
Motif DE_60h DE_60h-MTF1_MA0863.1 14 bp overlap
MYC 1 dataset
ChIP CD34 GSE85488.MYC.CD34 116 bp overlap
Mecom 3 datasets
Motif DE_48h DE_48h-Mecom_MA0029.2 11 bp overlap
Motif DE_60h DE_60h-Mecom_MA0029.2 11 bp overlap
Motif DE_72h DE_72h-Mecom_MA0029.2 11 bp overlap
Msx3 2 datasets
Motif DE_48h DE_48h-Msx3_MA0709.2 6 bp overlap
Motif DE_60h DE_60h-Msx3_MA0709.2 6 bp overlap
NKX2-1 1 dataset
ChIP NCI-H3122 GSE39998.NKX2-1.NCI-H3122 172 bp overlap
NR1D1 1 dataset
Motif DE_60h DE_60h-NR1D1_MA1531.2 14 bp overlap
Nfat5 1 dataset
Motif DE_60h DE_60h-Nfat5_MA0606.3 8 bp overlap
Nfe2l2 1 dataset
Motif DE_60h DE_60h-Nfe2l2_MA0150.3 11 bp overlap
Nobox 2 datasets
Motif DE_48h DE_48h-Nobox_MA0125.2 6 bp overlap
Motif DE_60h DE_60h-Nobox_MA0125.2 6 bp overlap
Nr2F6 1 dataset
Motif DE_60h DE_60h-Nr2F6_MA0728.1 15 bp overlap
Nr2f6 1 dataset
Motif DE_60h DE_60h-Nr2f6_MA0677.2 13 bp overlap
PAF1 2 datasets
ChIP HCT-116_ab20662 GSE97527.PAF1.HCT-116_ab20662 314 bp overlap
ChIP HCT-116_ab20662 GSE97527.PAF1.HCT-116_ab20662 324 bp overlap
PHIP 4 datasets
ChIP HCT-116_MLL1-KO__ab833 GSE101646.PHIP.HCT-116_MLL1-KO__ab833 59 bp overlap
ChIP HCT-116_MLL1-KO__ab833 GSE101646.PHIP.HCT-116_MLL1-KO__ab833 384 bp overlap
ChIP HCT-116_MLL1-KO__ab833 GSE101646.PHIP.HCT-116_MLL1-KO__ab833 239 bp overlap
ChIP HCT-116_ab833 GSE101646.PHIP.HCT-116_ab833 807 bp overlap
PLAG1 1 dataset
Motif DE_60h DE_60h-PLAG1_MA0163.1 14 bp overlap
PRDM9 3 datasets
Motif DE_48h DE_48h-PRDM9_MA1723.2 20 bp overlap
Motif DE_60h DE_60h-PRDM9_MA1723.2 20 bp overlap
Motif DE_60h DE_60h-PRDM9_MA1723.2 20 bp overlap
Prdm5 2 datasets
Motif DE_48h DE_48h-Prdm5_MA1999.2 11 bp overlap
Motif DE_60h DE_60h-Prdm5_MA1999.2 11 bp overlap
RAD51 1 dataset
ChIP GM12878 ENCSR482TWQ.RAD51.GM12878 305 bp overlap
RAX 2 datasets
Motif DE_48h DE_48h-RAX_MA0718.2 6 bp overlap
Motif DE_60h DE_60h-RAX_MA0718.2 6 bp overlap
RBPJ 1 dataset
Motif DE_60h DE_60h-RBPJ_MA1116.2 6 bp overlap
RXRB 1 dataset
Motif DE_60h DE_60h-RXRB_MA0855.1 14 bp overlap
Rarb 1 dataset
Motif DE_60h DE_60h-Rarb_MA0857.1 16 bp overlap
Rarg 1 dataset
Motif DE_60h DE_60h-Rarg_MA0859.2 15 bp overlap
SIX2 3 datasets
Motif DE_48h DE_48h-SIX2_MA1119.2 11 bp overlap
Motif DE_60h DE_60h-SIX2_MA1119.2 11 bp overlap
Motif DE_72h DE_72h-SIX2_MA1119.2 11 bp overlap
SMAD2 1 dataset
ChIP endoderm GSE29422.SMAD2.endoderm 169 bp overlap
SMAD2-3 3 datasets
ChIP HGrC1_C134W-TGF GSE138496.SMAD2-3.HGrC1_C134W-TGF 275 bp overlap
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 1134 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.SMAD2-3.HUES-8_DE_JNKi 1011 bp overlap
SMAD2_3 3 datasets
ChIP DE_D1 S12-DE-d1-SMAD2_3-exp1 611 bp overlap
ChIP DE_D2 S01-DE-d2-SMAD2_3-exp1 1215 bp overlap
ChIP DE_D2 S53-DE-d2-SMAD2_3-exp2 1077 bp overlap
SMAD3 1 dataset
ChIP NCI-H441_SIITTF1 GSE51509.SMAD3.NCI-H441_SIITTF1 307 bp overlap
SMAD4 1 dataset
ChIP endoderm GSE29422.SMAD4.endoderm 222 bp overlap
SMARCC1 1 dataset
ChIP Aska-SS_shSSX GSE108025.SMARCC1.Aska-SS_shSSX 197 bp overlap
SNAI2 1 dataset
ChIP SMS-CTR_shSNAI2 GSE137168.SNAI2.SMS-CTR_shSNAI2 185 bp overlap
SOX17 1 dataset
ChIP DE_D2 DED2-SOX17_Batch_II 994 bp overlap
SOX17_M 1 dataset
ChIP DE_D2 DED2-SOX17-M_Batch_II 1030 bp overlap
Spz1 1 dataset
Motif DE_60h DE_60h-Spz1_MA0111.1 11 bp overlap
Stat4 1 dataset
Motif DE_60h DE_60h-Stat4_MA0518.2 10 bp overlap
Stat6 1 dataset
Motif DE_60h DE_60h-Stat6_MA0520.2 10 bp overlap
TBX5 1 dataset
Motif DE_60h DE_60h-TBX5_MA0807.1 8 bp overlap
TCF7L1 2 datasets
Motif DE_48h DE_48h-TCF7L1_MA1421.1 12 bp overlap
Motif DE_60h DE_60h-TCF7L1_MA1421.1 12 bp overlap
TCF7L2 3 datasets
ChIP HCT-116 ENCSR000EUV.TCF7L2.HCT-116 382 bp overlap
ChIP HCT116 ENCFF038POZ 371 bp overlap
ChIP LNCaP GSE51621.TCF7L2.LNCaP 294 bp overlap
TFAP2A 1 dataset
Motif DE_60h DE_60h-TFAP2A_MA0003.5 9 bp overlap
TFAP2C 1 dataset
Motif DE_60h DE_60h-TFAP2C_MA0814.3 9 bp overlap
TFE3 1 dataset
ChIP K562 ENCFF697ABG 317 bp overlap
TRPS1 3 datasets
Motif DE_48h DE_48h-TRPS1_MA1970.2 8 bp overlap
Motif DE_60h DE_60h-TRPS1_MA1970.2 8 bp overlap
Motif DE_72h DE_72h-TRPS1_MA1970.2 8 bp overlap
Tbx6 1 dataset
Motif DE_60h DE_60h-Tbx6_MA1567.3 9 bp overlap
Thap11 1 dataset
Motif DE_60h DE_60h-Thap11_MA1573.2 14 bp overlap
USF1 2 datasets
ChIP HCT-116 ENCSR000BVK.USF1.HCT-116 228 bp overlap
ChIP HCT116 ENCFF330PYP 365 bp overlap
USF2 2 datasets
ChIP GM12878 GSE97661.USF2.GM12878 211 bp overlap
ChIP K-562 ENCSR359NFW.USF2.K-562 204 bp overlap
ZEB1 1 dataset
Motif DE_60h DE_60h-ZEB1_MA0103.4 6 bp overlap
ZFP57 1 dataset
ChIP hESC GSE115387.ZFP57.hESC 91 bp overlap
ZNF133 2 datasets
ChIP HEK293 ENCFF844RST 242 bp overlap
ChIP HEK293 ENCSR283MWQ.ZNF133.HEK293 134 bp overlap
ZNF257 2 datasets
Motif DE_48h DE_48h-ZNF257_MA1710.2 10 bp overlap
Motif DE_60h DE_60h-ZNF257_MA1710.2 10 bp overlap
ZNF283 1 dataset
ChIP HEK293T GSE78099.ZNF283.HEK293T 215 bp overlap
ZNF320 1 dataset
Motif DE_60h DE_60h-ZNF320_MA1976.2 20 bp overlap
ZNF528 1 dataset
Motif DE_60h DE_60h-ZNF528_MA1597.1 17 bp overlap
ZNF652 1 dataset
Motif DE_60h DE_60h-ZNF652_MA1657.2 9 bp overlap
ZNF677 1 dataset
Motif DE_60h DE_60h-ZNF677_MA2101.1 12 bp overlap