chr1 : 39,276,096 39,276,654
558 bp 93 TFs 5 linked genes
This 558 bp open chromatin element is linked to 5 target genes and is bound by 93 transcription factors.
Linked Genes
5 genes
Gene Expression Dist. to TSS Distance Link type
MACF1 171.0 kb Distal Multiome
NDUFS5 250.1 kb Distal Multiome
PPIEL 283.0 kb Distal Multiome
AKIRIN1 285.1 kb Distal Multiome
PABPC4 300.4 kb Distal Multiome
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr1:39,271,096 – 39,281,654
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
93 transcription factors
Source
Cell type
ARID2 1 dataset
ChIP BIN-67_lentivirus-SMARCA4 GSE117734.ARID2.BIN-67_lentivirus-SMARCA4 340 bp overlap
ATF1 1 dataset
ChIP K-562 ENCSR091GVJ.ATF1.K-562 354 bp overlap
ATF3 3 datasets
ChIP K-562 ENCSR028UIU.ATF3.K-562 447 bp overlap
ChIP K-562 ENCSR632DCH.ATF3.K-562 272 bp overlap
ChIP K562 ENCFF921JQW 558 bp overlap
ATF7 2 datasets
ChIP K-562 ENCSR972ZBV.ATF7.K-562 328 bp overlap
ChIP K562 ENCFF308SKS 491 bp overlap
BHLHE22 3 datasets
Motif DE_36h DE_36h-BHLHE22_MA1635.2 6 bp overlap
Motif DE_48h DE_48h-BHLHE22_MA1635.2 6 bp overlap
Motif DE_72h DE_72h-BHLHE22_MA1635.2 6 bp overlap
CBFA2T3 2 datasets
ChIP K-562 ENCSR697YLJ.CBFA2T3.K-562 199 bp overlap
ChIP K562 ENCFF673OEZ 411 bp overlap
CDX2 2 datasets
ChIP Caco-2_DIFF GSE23436.CDX2.Caco-2_DIFF 321 bp overlap
ChIP Caco-2_PROLIF GSE23436.CDX2.Caco-2_PROLIF 298 bp overlap
CEBPD 1 dataset
ChIP K-562 ENCSR000BVY.CEBPD.K-562 138 bp overlap
CHD4 1 dataset
ChIP HaCaT GSE139685.CHD4.HaCaT 177 bp overlap
CRX 3 datasets
ChIP retina_Hu20 GSE137311.CRX.retina_Hu20 333 bp overlap
ChIP retina_Hu3 GSE137311.CRX.retina_Hu3 294 bp overlap
ChIP retina_Hu6 GSE137311.CRX.retina_Hu6 392 bp overlap
CTBP1 1 dataset
ChIP K-562 ENCSR201NQZ.CTBP1.K-562 242 bp overlap
EOMES 1 dataset
ChIP hESC GSE26097.EOMES.hESC 429 bp overlap
ETS1 1 dataset
ChIP cardiomyocyte_D9 GSE129986.ETS1.cardiomyocyte_D9 192 bp overlap
FOXA1 1 dataset
ChIP pancreatic-progenitor_PP1 GSE104840.FOXA1.pancreatic-progenitor_PP1 250 bp overlap
FOXA2 6 datasets
ChIP BJ1-hTERT_FOXA2_GATA4_Coexp GSE92491.FOXA2.BJ1-hTERT_FOXA2_GATA4_Coexp 409 bp overlap
ChIP BJ1-hTERT_GATA4 GSE90454.FOXA2.BJ1-hTERT_GATA4 259 bp overlap
ChIP Caco-2 GSE66218.FOXA2.Caco-2 283 bp overlap
ChIP DE DE-FOXA2-1 507 bp overlap
ChIP DE DE-FOXA2-2 471 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.FOXA2.pancreatic-progenitor_PP1 340 bp overlap
GATA1 7 datasets
ChIP CD34_ERYTH_BIO GSE29194.GATA1.CD34_ERYTH_BIO 215 bp overlap
ChIP CD34_ERYTH_BMP GSE29194.GATA1.CD34_ERYTH_BMP 258 bp overlap
ChIP K-562 GSE107726.GATA1.K-562 410 bp overlap
ChIP K-562 ENCSR000EFT.GATA1.K-562 181 bp overlap
ChIP K-562_dCas9-LSD1 GSE132212.GATA1.K-562_dCas9-LSD1 319 bp overlap
ChIP K-562_enCRISPRi-LK GSE132212.GATA1.K-562_enCRISPRi-LK 275 bp overlap
ChIP K562 ENCFF094CMK 251 bp overlap
GATA1::TAL1 3 datasets
Motif DE_36h DE_36h-GATA1TAL1_MA0140.3 17 bp overlap
Motif DE_48h DE_48h-GATA1TAL1_MA0140.3 17 bp overlap
Motif DE_72h DE_72h-GATA1TAL1_MA0140.3 17 bp overlap
GATA2 10 datasets
Motif DE_36h DE_36h-GATA2_MA0036.4 7 bp overlap
Motif DE_48h DE_48h-GATA2_MA0036.4 7 bp overlap
Motif DE_72h DE_72h-GATA2_MA0036.4 7 bp overlap
ChIP K-562 ENCSR000DKA.GATA2.K-562 344 bp overlap
ChIP K562 ENCFF830LLA 558 bp overlap
ChIP K562 ENCFF830LLA 372 bp overlap
ChIP LNCaP_CSFCS GSE69043.GATA2.LNCaP_CSFCS 181 bp overlap
ChIP TF1 GSE73207.GATA2.TF1 237 bp overlap
ChIP WA09 GSE105081.GATA2.WA09 302 bp overlap
ChIP dermal-fibroblast_alone GSE51025.GATA2.dermal-fibroblast_alone 158 bp overlap
GATA3 1 dataset
ChIP WA09 GSE105081.GATA3.WA09 200 bp overlap
GATA4 12 datasets
ChIP A-549 GSE85002.GATA4.A-549 306 bp overlap
ChIP BJ1-hTERT_FOXA2_GATA4_Coexp GSE92491.GATA4.BJ1-hTERT_FOXA2_GATA4_Coexp 259 bp overlap
ChIP DE DE-GATA4-1 558 bp overlap
ChIP DE DE-GATA4-2 558 bp overlap
Motif DE_36h DE_36h-GATA4_MA0482.3 8 bp overlap
Motif DE_48h DE_48h-GATA4_MA0482.3 8 bp overlap
Motif DE_72h DE_72h-GATA4_MA0482.3 8 bp overlap
ChIP Hep-G2 GSE135714.GATA4.Hep-G2 558 bp overlap
ChIP YCC-3 GSE51705.GATA4.YCC-3 317 bp overlap
ChIP foregut GSE117136.GATA4.foregut 558 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA4.pancreatic-progenitor 558 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA4.pancreatic-progenitor_PP1 466 bp overlap
GATA5 3 datasets
Motif DE_36h DE_36h-GATA5_MA0766.3 8 bp overlap
Motif DE_48h DE_48h-GATA5_MA0766.3 8 bp overlap
Motif DE_72h DE_72h-GATA5_MA0766.3 8 bp overlap
GATA6 22 datasets
ChIP AGS GSE51705.GATA6.AGS 441 bp overlap
ChIP AGS GSE51936.GATA6.AGS 157 bp overlap
ChIP Caco-2_DIFF GSE23436.GATA6.Caco-2_DIFF 229 bp overlap
ChIP Caco-2_PROLIF GSE23436.GATA6.Caco-2_PROLIF 436 bp overlap
ChIP DE DE-GATA6-1 558 bp overlap
ChIP DE DE-GATA6-2 558 bp overlap
Motif DE_36h DE_36h-GATA6_MA1104.3 8 bp overlap
Motif DE_48h DE_48h-GATA6_MA1104.3 8 bp overlap
Motif DE_72h DE_72h-GATA6_MA1104.3 8 bp overlap
ChIP DE_D1 S09-DE-d1-GATA6-exp1 558 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 558 bp overlap
ChIP DE_D1 S41-DE-d1-GATA6-exp2 558 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 558 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 558 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 558 bp overlap
ChIP YCC-3 GSE51705.GATA6.YCC-3 412 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 558 bp overlap
ChIP foregut GSE117136.GATA6.foregut 558 bp overlap
ChIP foregut_KO GSE117136.GATA6.foregut_KO 472 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA6.pancreatic-progenitor 548 bp overlap
ChIP pancreatic-progenitor_KO GSE117136.GATA6.pancreatic-progenitor_KO 554 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA6.pancreatic-progenitor_PP1 479 bp overlap
Gata3 3 datasets
Motif DE_36h DE_36h-Gata3_MA0037.5 8 bp overlap
Motif DE_48h DE_48h-Gata3_MA0037.5 8 bp overlap
Motif DE_72h DE_72h-Gata3_MA0037.5 8 bp overlap
HDAC2 1 dataset
ChIP K-562 ENCSR893WSB.HDAC2.K-562 367 bp overlap
HMBOX1 3 datasets
ChIP K-562 ENCSR757IIU.HMBOX1.K-562 300 bp overlap
ChIP K562 ENCFF055GAZ 501 bp overlap
ChIP K562 ENCFF317JJX 496 bp overlap
HNF4A 3 datasets
ChIP Caco-2_DIFF GSE23436.HNF4A.Caco-2_DIFF 309 bp overlap
ChIP Caco-2_PROLIF GSE23436.HNF4A.Caco-2_PROLIF 215 bp overlap
Motif DE_72h DE_72h-HNF4A_MA0114.5 9 bp overlap
HNF4G 1 dataset
Motif DE_72h DE_72h-HNF4G_MA0484.3 9 bp overlap
IKZF1 2 datasets
ChIP K-562 ENCSR395HWC.IKZF1.K-562 267 bp overlap
ChIP K562 ENCFF348IBL 391 bp overlap
IRF2 1 dataset
ChIP keratinocyte_DOX GSE135677.IRF2.keratinocyte_DOX 205 bp overlap
IRF6 2 datasets
Motif DE_36h DE_36h-IRF6_MA1509.1 9 bp overlap
Motif DE_72h DE_72h-IRF6_MA1509.1 9 bp overlap
JUN 6 datasets
ChIP DE_D1 S08-DE-d1-JUN-exp1 495 bp overlap
ChIP DE_D1 S13-DE-d1-JUN-exp1 517 bp overlap
ChIP DE_D1 S40-DE-d1-JUN-exp2 491 bp overlap
ChIP DE_D2 S02-DE-d2-JUN-exp1 558 bp overlap
ChIP DE_D2 S54-DE-d2-JUN-exp2 467 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.JUN.HUES-8_DE_ctrl 507 bp overlap
JUND 2 datasets
ChIP K-562 ENCSR000EGN.JUND.K-562 185 bp overlap
ChIP K562 ENCFF830LVJ 281 bp overlap
KDM1A 1 dataset
ChIP keratinocyte_diff GSE57702.KDM1A.keratinocyte_diff 191 bp overlap
KLF3 1 dataset
ChIP HEK293 GSE69739.KLF3.HEK293 170 bp overlap
KLF4 1 dataset
ChIP keratinocyte_diff GSE57702.KLF4.keratinocyte_diff 59 bp overlap
LDB1 1 dataset
ChIP K-562 GSE142227.LDB1.K-562 188 bp overlap
MAML1 1 dataset
ChIP SCC_4h GSE156486.MAML1.SCC_4h 321 bp overlap
Mecom 3 datasets
Motif DE_36h DE_36h-Mecom_MA0029.2 11 bp overlap
Motif DE_48h DE_48h-Mecom_MA0029.2 11 bp overlap
Motif DE_72h DE_72h-Mecom_MA0029.2 11 bp overlap
NKX2-5 1 dataset
ChIP hESC_ab3584 GSE89457.NKX2-5.hESC_ab3584 197 bp overlap
NR3C1 5 datasets
ChIP HCC1937 GSE152203.NR3C1.HCC1937 223 bp overlap
ChIP LNCaP_1F5_SIFOXA1 GSE30623.NR3C1.LNCaP_1F5_SIFOXA1 141 bp overlap
ChIP SUM159PT GSE152203.NR3C1.SUM159PT 241 bp overlap
ChIP U2OS_GLUCC ERP007081.NR3C1.U2OS_GLUCC 55 bp overlap
ChIP U2OS_SHHIC5 GSE65847.NR3C1.U2OS_SHHIC5 232 bp overlap
NRL 1 dataset
ChIP retina_Hu13 GSE137311.NRL.retina_Hu13 242 bp overlap
Neurod2 3 datasets
Motif DE_36h DE_36h-Neurod2_MA1993.2 6 bp overlap
Motif DE_48h DE_48h-Neurod2_MA1993.2 6 bp overlap
Motif DE_72h DE_72h-Neurod2_MA1993.2 6 bp overlap
OSR2 3 datasets
ChIP HEK293 ENCFF875BDB 210 bp overlap
ChIP HEK293 ENCSR324LTM.OSR2.HEK293 313 bp overlap
ChIP HEK293 GSE76494.OSR2.HEK293 205 bp overlap
Olig2 3 datasets
Motif DE_36h DE_36h-Olig2_MA1997.2 6 bp overlap
Motif DE_48h DE_48h-Olig2_MA1997.2 6 bp overlap
Motif DE_72h DE_72h-Olig2_MA1997.2 6 bp overlap
PATZ1 1 dataset
ChIP HEK293 ENCSR966ULI.PATZ1.HEK293 301 bp overlap
POLR2H 1 dataset
ChIP K562 ENCFF377NHG 281 bp overlap
PRDM1 2 datasets
ChIP HEK293 ENCFF302TBP 222 bp overlap
ChIP HEK293 GSE76494.PRDM1.HEK293 152 bp overlap
PRDM10 1 dataset
ChIP HEK293 ENCFF145WQQ 460 bp overlap
PRDM6 1 dataset
ChIP HEK293 ENCSR892QHR.PRDM6.HEK293 221 bp overlap
RCOR1 1 dataset
ChIP keratinocyte_diff GSE57702.RCOR1.keratinocyte_diff 498 bp overlap
RELA 1 dataset
ChIP FaDu_LPS GSE132018.RELA.FaDu_LPS 314 bp overlap
RORB 1 dataset
ChIP retina_Hu22 GSE137311.RORB.retina_Hu22 217 bp overlap
SETDB1 3 datasets
ChIP HEK293 ENCFF676PLV 491 bp overlap
ChIP HEK293 ENCSR348AGV.SETDB1.HEK293 401 bp overlap
ChIP U2OS ENCSR000EYD.SETDB1.U2OS 408 bp overlap
SMAD2-3 4 datasets
ChIP HGrC1_C134W-TGF GSE138496.SMAD2-3.HGrC1_C134W-TGF 120 bp overlap
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 458 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.SMAD2-3.HUES-8_DE_JNKi 524 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.SMAD2-3.HUES-8_DE_ctrl 411 bp overlap
SMAD2_3 4 datasets
ChIP DE_D1 S07-DE-d1-SMAD2_3-exp1 396 bp overlap
ChIP DE_D1 S12-DE-d1-SMAD2_3-exp1 460 bp overlap
ChIP DE_D2 S01-DE-d2-SMAD2_3-exp1 435 bp overlap
ChIP DE_D2 S53-DE-d2-SMAD2_3-exp2 537 bp overlap
SMAD4 1 dataset
ChIP Caco-2 GSE112946.SMAD4.Caco-2 163 bp overlap
SMARCA4 2 datasets
ChIP K-562 ENCSR587OQL.SMARCA4.K-562 276 bp overlap
ChIP hiPSC GSE124903.SMARCA4.hiPSC 307 bp overlap
SMARCC1 2 datasets
ChIP DE_D1 S15-DE-d1-BAF155-exp1 327 bp overlap
ChIP hiPSC GSE124903.SMARCC1.hiPSC 227 bp overlap
SNAI2 1 dataset
ChIP SMS-CTR_T48 GSE137168.SNAI2.SMS-CTR_T48 170 bp overlap
SOX17 1 dataset
ChIP DE_D2 DED2-SOX17_Batch_II 369 bp overlap
SOX17_M 1 dataset
ChIP DE_D2 DED2-SOX17-M_Batch_II 401 bp overlap
SOX6 2 datasets
ChIP K-562 ENCSR788RSW.SOX6.K-562 294 bp overlap
ChIP K562 ENCFF059YCJ 437 bp overlap
SP7 2 datasets
ChIP HEK293 ENCFF733RBE 244 bp overlap
ChIP HEK293 ENCSR468IJT.SP7.HEK293 558 bp overlap
SPI1 1 dataset
ChIP BDMC_donorG GSE128834.SPI1.BDMC_donorG 152 bp overlap
T 1 dataset
ChIP H9_ENDODERM GSE60606.T.H9_ENDODERM 304 bp overlap
TAL1 2 datasets
ChIP K-562 GSE107726.TAL1.K-562 170 bp overlap
ChIP K-562_enCRISPRi-KL GSE132212.TAL1.K-562_enCRISPRi-KL 141 bp overlap
TP63 15 datasets
ChIP BxPC-3 GSE115461.TP63.BxPC-3 396 bp overlap
ChIP HaCaT_LacZ_TGFB GSE60814.TP63.HaCaT_LacZ_TGFB 260 bp overlap
ChIP HaCaT_caRAS_TGFB GSE60814.TP63.HaCaT_caRAS_TGFB 212 bp overlap
ChIP HaCaT_dnRAS_TGFB GSE60814.TP63.HaCaT_dnRAS_TGFB 252 bp overlap
ChIP JHU-029 GSE88859.TP63.JHU-029 184 bp overlap
ChIP MCF-10A_Nutlin3A GSE111009.TP63.MCF-10A_Nutlin3A 558 bp overlap
ChIP SUIT-2 GSE115461.TP63.SUIT-2 447 bp overlap
ChIP keratinocyte GSE56674.TP63.keratinocyte 305 bp overlap
ChIP keratinocyte_CISP GSE56674.TP63.keratinocyte_CISP 263 bp overlap
ChIP keratinocyte_D0 GSE59824.TP63.keratinocyte_D0 475 bp overlap
ChIP keratinocyte_D2 GSE59824.TP63.keratinocyte_D2 405 bp overlap
ChIP keratinocyte_D4 GSE59824.TP63.keratinocyte_D4 372 bp overlap
ChIP keratinocyte_D7 GSE59824.TP63.keratinocyte_D7 388 bp overlap
ChIP keratinocyte_epidermal GSE67382.TP63.keratinocyte_epidermal 330 bp overlap
ChIP keratinocyte_epidermal_KDPAF GSE67382.TP63.keratinocyte_epidermal_KDPAF 206 bp overlap
TP73_TA 1 dataset
ChIP SaOS-2 GSE15780.TP73_TA.SaOS-2 344 bp overlap
TRIM28 6 datasets
ChIP HEK293 ENCFF265CEM 546 bp overlap
ChIP HEK293 ENCFF265CEM 458 bp overlap
ChIP HEK293 ENCFF582MWI 335 bp overlap
ChIP HEK293 ENCSR000EUZ.TRIM28.HEK293 454 bp overlap
ChIP HEK293 ENCSR618HNF.TRIM28.HEK293 309 bp overlap
ChIP U2OS ENCSR000EYC.TRIM28.U2OS 314 bp overlap
TRPS1 3 datasets
Motif DE_36h DE_36h-TRPS1_MA1970.2 8 bp overlap
Motif DE_48h DE_48h-TRPS1_MA1970.2 8 bp overlap
Motif DE_72h DE_72h-TRPS1_MA1970.2 8 bp overlap
Tcf12 3 datasets
Motif DE_36h DE_36h-Tcf12_MA0521.3 6 bp overlap
Motif DE_48h DE_48h-Tcf12_MA0521.3 6 bp overlap
Motif DE_72h DE_72h-Tcf12_MA0521.3 6 bp overlap
Twist2 3 datasets
Motif DE_36h DE_36h-Twist2_MA0633.3 6 bp overlap
Motif DE_48h DE_48h-Twist2_MA0633.3 6 bp overlap
Motif DE_72h DE_72h-Twist2_MA0633.3 6 bp overlap
YY1 1 dataset
ChIP Hep-G2_RBM25-KD GSE120104.YY1.Hep-G2_RBM25-KD 218 bp overlap
ZBTB17 1 dataset
ChIP HEK293 ENCFF865LIO 345 bp overlap
ZBTB21 2 datasets
ChIP HEK293 ENCFF509WYZ 406 bp overlap
ChIP HEK293 ENCSR321MSF.ZBTB21.HEK293 255 bp overlap
ZEB2 2 datasets
ChIP HEK293 ENCFF847JIE 476 bp overlap
ChIP HEK293 ENCSR417VWF.ZEB2.HEK293 425 bp overlap
ZNF184 2 datasets
ChIP HEK293 ENCFF221CII 357 bp overlap
ChIP HEK293 ENCSR020UPN.ZNF184.HEK293 403 bp overlap
ZNF2 1 dataset
ChIP HEK293 ENCSR011CKE.ZNF2.HEK293 435 bp overlap
ZNF35 1 dataset
ChIP HEK293 GSE76494.ZNF35.HEK293 144 bp overlap
ZNF354C 2 datasets
Motif DE_36h DE_36h-ZNF354C_MA0130.1 6 bp overlap
Motif DE_72h DE_72h-ZNF354C_MA0130.1 6 bp overlap
ZNF366 2 datasets
ChIP HEK293 ENCFF799ATK 148 bp overlap
ChIP HEK293 ENCSR106EBH.ZNF366.HEK293 558 bp overlap
ZNF528 1 dataset
ChIP HEK293 GSE76494.ZNF528.HEK293 188 bp overlap
ZNF629 2 datasets
ChIP HEK293 ENCFF096ELQ 202 bp overlap
ChIP HEK293 ENCSR351NON.ZNF629.HEK293 401 bp overlap
ZSCAN21 2 datasets
ChIP HEK293 ENCFF582WUP 234 bp overlap
ChIP HEK293 ENCSR253CKN.ZSCAN21.HEK293 424 bp overlap
ZXDB 1 dataset
ChIP HEK293 ENCSR559IOZ.ZXDB.HEK293 440 bp overlap
Zfp809 3 datasets
Motif DE_36h DE_36h-Zfp809_MA2125.1 9 bp overlap
Motif DE_48h DE_48h-Zfp809_MA2125.1 9 bp overlap
Motif DE_72h DE_72h-Zfp809_MA2125.1 9 bp overlap