chr1 : 171,805,179 171,806,145
966 bp 92 TFs 0 linked genes
This 966 bp open chromatin element has no linked target genes and is bound by 92 transcription factors.
Linked Genes
No linked genes
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr1:171,800,179 – 171,811,145
This element Other elements ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
92 transcription factors
Source
Cell type
ARNTL 1 dataset
ChIP U2OS_DMOG GSE85096.ARNTL.U2OS_DMOG 230 bp overlap
ATF2 6 datasets
Motif DE_36h DE_36h-ATF2_MA1632.2 10 bp overlap
Motif DE_48h DE_48h-ATF2_MA1632.2 10 bp overlap
Motif DE_60h DE_60h-ATF2_MA1632.2 10 bp overlap
Motif DE_72h DE_72h-ATF2_MA1632.2 10 bp overlap
ChIP GM12878 ENCFF521LQJ 359 bp overlap
ChIP GM12878 ENCFF521LQJ 160 bp overlap
ATF7 1 dataset
ChIP GM12878 ENCSR014YCR.ATF7.GM12878 295 bp overlap
BATF 2 datasets
ChIP GM12878 ENCFF954REE 69 bp overlap
ChIP OCI-Ly3 GSE56857.BATF.OCI-Ly3 152 bp overlap
BCL11A 1 dataset
ChIP CD34_Day9_30min GSE104676.BCL11A.CD34_Day9_30min 54 bp overlap
BRD4 4 datasets
ChIP HEK293_sgNT GSE129407.BRD4.HEK293_sgNT 216 bp overlap
ChIP MM1-S_JQ1_500NM GSE42355.BRD4.MM1-S_JQ1_500NM 296 bp overlap
ChIP MM1-S_JQ1_50NM GSE42355.BRD4.MM1-S_JQ1_50NM 379 bp overlap
ChIP MM1-S_JQ1_5NM GSE42355.BRD4.MM1-S_JQ1_5NM 206 bp overlap
BRD7 2 datasets
ChIP SUM159PT_DMSO GSE131097.BRD7.SUM159PT_DMSO 225 bp overlap
ChIP SUM159PT_JQ1 GSE131097.BRD7.SUM159PT_JQ1 302 bp overlap
Bcl11B 2 datasets
Motif DE_60h DE_60h-Bcl11B_MA1989.2 9 bp overlap
Motif DE_72h DE_72h-Bcl11B_MA1989.2 9 bp overlap
CDK9 3 datasets
ChIP MM1-S_DMSO GSE42161.CDK9.MM1-S_DMSO 182 bp overlap
ChIP MM1-S_JQ1_5000NM GSE49224.CDK9.MM1-S_JQ1_5000NM 228 bp overlap
ChIP MM1-S_JQ1_500NM GSE42355.CDK9.MM1-S_JQ1_500NM 213 bp overlap
CREB1 4 datasets
Motif DE_36h DE_36h-CREB1_MA0018.5 8 bp overlap
Motif DE_48h DE_48h-CREB1_MA0018.5 8 bp overlap
Motif DE_60h DE_60h-CREB1_MA0018.5 8 bp overlap
Motif DE_72h DE_72h-CREB1_MA0018.5 8 bp overlap
CREM 4 datasets
Motif DE_36h DE_36h-CREM_MA0609.3 10 bp overlap
Motif DE_48h DE_48h-CREM_MA0609.3 10 bp overlap
Motif DE_60h DE_60h-CREM_MA0609.3 10 bp overlap
Motif DE_72h DE_72h-CREM_MA0609.3 10 bp overlap
CRY1 2 datasets
ChIP U2OS GSE130602.CRY1.U2OS 259 bp overlap
ChIP U2OS_DMSO GSE130506.CRY1.U2OS_DMSO 258 bp overlap
CTCF 2 datasets
ChIP CD14-positive monocyte ENCFF087XLR 450 bp overlap
ChIP dorsolateral prefrontal cortex ENCFF277CZQ 440 bp overlap
Crx 4 datasets
Motif DE_36h DE_36h-Crx_MA0467.3 6 bp overlap
Motif DE_48h DE_48h-Crx_MA0467.3 6 bp overlap
Motif DE_60h DE_60h-Crx_MA0467.3 6 bp overlap
Motif DE_72h DE_72h-Crx_MA0467.3 6 bp overlap
DPF2 1 dataset
ChIP GM12878 ENCSR509FWH.DPF2.GM12878 207 bp overlap
DPRX 4 datasets
Motif DE_36h DE_36h-DPRX_MA1480.2 9 bp overlap
Motif DE_48h DE_48h-DPRX_MA1480.2 9 bp overlap
Motif DE_60h DE_60h-DPRX_MA1480.2 9 bp overlap
Motif DE_72h DE_72h-DPRX_MA1480.2 9 bp overlap
DUX4 4 datasets
Motif DE_36h DE_36h-DUX4_MA0468.1 11 bp overlap
Motif DE_48h DE_48h-DUX4_MA0468.1 11 bp overlap
Motif DE_60h DE_60h-DUX4_MA0468.1 11 bp overlap
Motif DE_72h DE_72h-DUX4_MA0468.1 11 bp overlap
Dmbx1 4 datasets
Motif DE_36h DE_36h-Dmbx1_MA0883.2 10 bp overlap
Motif DE_48h DE_48h-Dmbx1_MA0883.2 10 bp overlap
Motif DE_60h DE_60h-Dmbx1_MA0883.2 10 bp overlap
Motif DE_72h DE_72h-Dmbx1_MA0883.2 10 bp overlap
EBF1 1 dataset
ChIP GM12878 ENCFF813OXE 265 bp overlap
ESR2 4 datasets
Motif DE_36h DE_36h-ESR2_MA0258.2 15 bp overlap
Motif DE_48h DE_48h-ESR2_MA0258.2 15 bp overlap
Motif DE_60h DE_60h-ESR2_MA0258.2 15 bp overlap
Motif DE_72h DE_72h-ESR2_MA0258.2 15 bp overlap
ETS1 1 dataset
ChIP SCC-25 GSE109884.ETS1.SCC-25 227 bp overlap
FOXA2 2 datasets
ChIP DE DE-FOXA2-1 305 bp overlap
ChIP DE DE-FOXA2-2 394 bp overlap
FOXE1 2 datasets
Motif DE_60h DE_60h-FOXE1_MA1487.3 12 bp overlap
Motif DE_72h DE_72h-FOXE1_MA1487.3 12 bp overlap
GATA1 1 dataset
ChIP K-562_enCRISPRi-LK GSE132212.GATA1.K-562_enCRISPRi-LK 262 bp overlap
GATA2 4 datasets
ChIP K-562 ENCSR000DKA.GATA2.K-562 426 bp overlap
ChIP K-562 ENCSR000EWG.GATA2.K-562 116 bp overlap
ChIP K562 ENCFF830LLA 577 bp overlap
ChIP K562 ENCFF830LLA 577 bp overlap
GATA3 4 datasets
ChIP Kelly GSE65664.GATA3.Kelly 161 bp overlap
ChIP T-47D GSE51274.GATA3.T-47D 258 bp overlap
ChIP T-47D_sc GSE122847.GATA3.T-47D_sc 185 bp overlap
ChIP T47D-A1-2_EtOH GSE112491.GATA3.T47D-A1-2_EtOH 338 bp overlap
GATA3_Nter 2 datasets
ChIP T-47D_CR3flp GSE99479.GATA3_Nter.T-47D_CR3flp 447 bp overlap
ChIP T-47D_flp-ctrl GSE99479.GATA3_Nter.T-47D_flp-ctrl 266 bp overlap
GATA4 2 datasets
ChIP DE DE-GATA4-1 544 bp overlap
ChIP DE DE-GATA4-2 566 bp overlap
GATA6 7 datasets
ChIP DE DE-GATA6-1 609 bp overlap
ChIP DE DE-GATA6-2 563 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 289 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 368 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 463 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 542 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 400 bp overlap
GCM1 1 dataset
Motif DE_60h DE_60h-GCM1_MA0646.2 10 bp overlap
GSC 4 datasets
Motif DE_36h DE_36h-GSC_MA0648.2 6 bp overlap
Motif DE_48h DE_48h-GSC_MA0648.2 6 bp overlap
Motif DE_60h DE_60h-GSC_MA0648.2 6 bp overlap
Motif DE_72h DE_72h-GSC_MA0648.2 6 bp overlap
GSC2 4 datasets
Motif DE_36h DE_36h-GSC2_MA0891.2 6 bp overlap
Motif DE_48h DE_48h-GSC2_MA0891.2 6 bp overlap
Motif DE_60h DE_60h-GSC2_MA0891.2 6 bp overlap
Motif DE_72h DE_72h-GSC2_MA0891.2 6 bp overlap
HNF4A 2 datasets
Motif DE_60h DE_60h-HNF4A_MA1494.2 14 bp overlap
Motif DE_72h DE_72h-HNF4A_MA1494.2 14 bp overlap
IRF1 8 datasets
ChIP AsPC-1_IFNg GSE141606.IRF1.AsPC-1_IFNg 251 bp overlap
ChIP AsPC-1_ZBED2-cDNA GSE141606.IRF1.AsPC-1_ZBED2-cDNA 256 bp overlap
ChIP CD14_LPS GSE43036.IRF1.CD14_LPS 185 bp overlap
ChIP HAEC_TNFa_4h GSE89970.IRF1.HAEC_TNFa_4h 141 bp overlap
ChIP K-562 ENCSR854MCV.IRF1.K-562 469 bp overlap
ChIP K-562_EPZ GSE129380.IRF1.K-562_EPZ 367 bp overlap
ChIP K562 ENCFF277KTJ 438 bp overlap
ChIP K562 ENCFF277KTJ 238 bp overlap
IRF2 2 datasets
ChIP K-562 ENCSR376WCJ.IRF2.K-562 168 bp overlap
ChIP keratinocyte_DOX GSE135677.IRF2.keratinocyte_DOX 266 bp overlap
IRF4 8 datasets
ChIP B-cell GSE142493.IRF4.B-cell 192 bp overlap
ChIP B-cell GSE142493.IRF4.B-cell 294 bp overlap
ChIP BC-3 GSE132777.IRF4.BC-3 240 bp overlap
ChIP GM12878 ENCSR000BGY.IRF4.GM12878 172 bp overlap
ChIP KK-1 GSE94732.IRF4.KK-1 98 bp overlap
ChIP OCI-Ly3 GSE142493.IRF4.OCI-Ly3 174 bp overlap
ChIP OCI-Ly3_SHCTR GSE56857.IRF4.OCI-Ly3_SHCTR 217 bp overlap
ChIP T-cell GSE136853.IRF4.T-cell 358 bp overlap
JUN 5 datasets
Motif DE_36h DE_36h-JUN_MA0488.2 10 bp overlap
Motif DE_48h DE_48h-JUN_MA0488.2 10 bp overlap
Motif DE_60h DE_60h-JUN_MA0488.2 10 bp overlap
Motif DE_72h DE_72h-JUN_MA0488.2 10 bp overlap
ChIP Karpas-299 GSE151413.JUN.Karpas-299 309 bp overlap
JUNB 2 datasets
ChIP GM12878 ENCFF667EJQ 392 bp overlap
ChIP GM12878 ENCSR897MMC.JUNB.GM12878 293 bp overlap
JUND 4 datasets
Motif DE_36h DE_36h-JUND_MA0492.2 11 bp overlap
Motif DE_48h DE_48h-JUND_MA0492.2 11 bp overlap
Motif DE_60h DE_60h-JUND_MA0492.2 11 bp overlap
Motif DE_72h DE_72h-JUND_MA0492.2 11 bp overlap
KLF11 1 dataset
Motif DE_60h DE_60h-KLF11_MA1512.2 10 bp overlap
KLF13 1 dataset
Motif DE_60h DE_60h-KLF13_MA0657.2 17 bp overlap
KLF16 1 dataset
Motif DE_60h DE_60h-KLF16_MA0741.1 11 bp overlap
KLF6 1 dataset
Motif DE_60h DE_60h-KLF6_MA1517.2 9 bp overlap
MAZ 1 dataset
ChIP K562 ENCFF982GSZ 517 bp overlap
MED1 3 datasets
ChIP MM1-S_JQ1_500NM GSE42355.MED1.MM1-S_JQ1_500NM 176 bp overlap
ChIP MM1-S_JQ1_50NM GSE49224.MED1.MM1-S_JQ1_50NM 166 bp overlap
ChIP MM1-S_JQ1_5NM GSE49224.MED1.MM1-S_JQ1_5NM 169 bp overlap
MSANTD3 4 datasets
Motif DE_36h DE_36h-MSANTD3_MA1523.2 7 bp overlap
Motif DE_48h DE_48h-MSANTD3_MA1523.2 7 bp overlap
Motif DE_60h DE_60h-MSANTD3_MA1523.2 7 bp overlap
Motif DE_72h DE_72h-MSANTD3_MA1523.2 7 bp overlap
MTA2 1 dataset
ChIP GM12878 ENCSR293QAR.MTA2.GM12878 348 bp overlap
NFIA 4 datasets
Motif DE_36h DE_36h-NFIA_MA0670.2 6 bp overlap
Motif DE_48h DE_48h-NFIA_MA0670.2 6 bp overlap
Motif DE_60h DE_60h-NFIA_MA0670.2 6 bp overlap
Motif DE_72h DE_72h-NFIA_MA0670.2 6 bp overlap
NFIC 4 datasets
Motif DE_36h DE_36h-NFIC_MA0161.3 7 bp overlap
Motif DE_48h DE_48h-NFIC_MA0161.3 7 bp overlap
Motif DE_60h DE_60h-NFIC_MA0161.3 7 bp overlap
Motif DE_72h DE_72h-NFIC_MA0161.3 7 bp overlap
NFIX 4 datasets
Motif DE_36h DE_36h-NFIX_MA0671.2 6 bp overlap
Motif DE_48h DE_48h-NFIX_MA0671.2 6 bp overlap
Motif DE_60h DE_60h-NFIX_MA0671.2 6 bp overlap
Motif DE_72h DE_72h-NFIX_MA0671.2 6 bp overlap
NR2C1 4 datasets
Motif DE_36h DE_36h-NR2C1_MA1535.2 6 bp overlap
Motif DE_48h DE_48h-NR2C1_MA1535.2 6 bp overlap
Motif DE_60h DE_60h-NR2C1_MA1535.2 6 bp overlap
Motif DE_72h DE_72h-NR2C1_MA1535.2 6 bp overlap
NR2C2 4 datasets
Motif DE_36h DE_36h-NR2C2_MA1536.2 6 bp overlap
Motif DE_48h DE_48h-NR2C2_MA1536.2 6 bp overlap
Motif DE_60h DE_60h-NR2C2_MA1536.2 6 bp overlap
Motif DE_72h DE_72h-NR2C2_MA1536.2 6 bp overlap
Nfat5 1 dataset
Motif DE_60h DE_60h-Nfat5_MA0606.3 8 bp overlap
Nr1H2 4 datasets
Motif DE_36h DE_36h-Nr1H2_MA1996.2 6 bp overlap
Motif DE_48h DE_48h-Nr1H2_MA1996.2 6 bp overlap
Motif DE_60h DE_60h-Nr1H2_MA1996.2 6 bp overlap
Motif DE_72h DE_72h-Nr1H2_MA1996.2 6 bp overlap
Nr1H4 4 datasets
Motif DE_36h DE_36h-Nr1H4_MA1110.3 6 bp overlap
Motif DE_48h DE_48h-Nr1H4_MA1110.3 6 bp overlap
Motif DE_60h DE_60h-Nr1H4_MA1110.3 6 bp overlap
Motif DE_72h DE_72h-Nr1H4_MA1110.3 6 bp overlap
Nr1h3 4 datasets
Motif DE_36h DE_36h-Nr1h3_MA2337.1 6 bp overlap
Motif DE_48h DE_48h-Nr1h3_MA2337.1 6 bp overlap
Motif DE_60h DE_60h-Nr1h3_MA2337.1 6 bp overlap
Motif DE_72h DE_72h-Nr1h3_MA2337.1 6 bp overlap
OTX1 4 datasets
Motif DE_36h DE_36h-OTX1_MA0711.2 6 bp overlap
Motif DE_48h DE_48h-OTX1_MA0711.2 6 bp overlap
Motif DE_60h DE_60h-OTX1_MA0711.2 6 bp overlap
Motif DE_72h DE_72h-OTX1_MA0711.2 6 bp overlap
PHOX2A 4 datasets
Motif DE_36h DE_36h-PHOX2A_MA0713.1 11 bp overlap
Motif DE_48h DE_48h-PHOX2A_MA0713.1 11 bp overlap
Motif DE_60h DE_60h-PHOX2A_MA0713.1 11 bp overlap
Motif DE_72h DE_72h-PHOX2A_MA0713.1 11 bp overlap
PHOX2B 4 datasets
Motif DE_36h DE_36h-PHOX2B_MA0681.3 12 bp overlap
Motif DE_48h DE_48h-PHOX2B_MA0681.3 12 bp overlap
Motif DE_60h DE_60h-PHOX2B_MA0681.3 12 bp overlap
Motif DE_72h DE_72h-PHOX2B_MA0681.3 12 bp overlap
PITX1 4 datasets
Motif DE_36h DE_36h-PITX1_MA0682.3 6 bp overlap
Motif DE_48h DE_48h-PITX1_MA0682.3 6 bp overlap
Motif DE_60h DE_60h-PITX1_MA0682.3 6 bp overlap
Motif DE_72h DE_72h-PITX1_MA0682.3 6 bp overlap
PITX3 4 datasets
Motif DE_36h DE_36h-PITX3_MA0714.2 6 bp overlap
Motif DE_48h DE_48h-PITX3_MA0714.2 6 bp overlap
Motif DE_60h DE_60h-PITX3_MA0714.2 6 bp overlap
Motif DE_72h DE_72h-PITX3_MA0714.2 6 bp overlap
PRDM9 3 datasets
Motif DE_36h DE_36h-PRDM9_MA1723.2 20 bp overlap
Motif DE_60h DE_60h-PRDM9_MA1723.2 20 bp overlap
Motif DE_72h DE_72h-PRDM9_MA1723.2 20 bp overlap
PROP1 4 datasets
Motif DE_36h DE_36h-PROP1_MA0715.1 11 bp overlap
Motif DE_48h DE_48h-PROP1_MA0715.1 11 bp overlap
Motif DE_60h DE_60h-PROP1_MA0715.1 11 bp overlap
Motif DE_72h DE_72h-PROP1_MA0715.1 11 bp overlap
Pgr 2 datasets
Motif DE_60h DE_60h-Pgr_MA2323.1 17 bp overlap
Motif DE_72h DE_72h-Pgr_MA2323.1 17 bp overlap
Prdm5 1 dataset
Motif DE_60h DE_60h-Prdm5_MA1999.2 11 bp overlap
RBPJ 1 dataset
ChIP LCL GSE75503.RBPJ.LCL 148 bp overlap
RFX1 4 datasets
Motif DE_36h DE_36h-RFX1_MA0509.3 16 bp overlap
Motif DE_48h DE_48h-RFX1_MA0509.3 16 bp overlap
Motif DE_60h DE_60h-RFX1_MA0509.3 16 bp overlap
Motif DE_72h DE_72h-RFX1_MA0509.3 16 bp overlap
RFX2 4 datasets
Motif DE_36h DE_36h-RFX2_MA0600.3 14 bp overlap
Motif DE_48h DE_48h-RFX2_MA0600.3 14 bp overlap
Motif DE_60h DE_60h-RFX2_MA0600.3 14 bp overlap
Motif DE_72h DE_72h-RFX2_MA0600.3 14 bp overlap
RFX3 4 datasets
Motif DE_36h DE_36h-RFX3_MA0798.3 16 bp overlap
Motif DE_48h DE_48h-RFX3_MA0798.3 16 bp overlap
Motif DE_60h DE_60h-RFX3_MA0798.3 16 bp overlap
Motif DE_72h DE_72h-RFX3_MA0798.3 16 bp overlap
RFX5 4 datasets
Motif DE_36h DE_36h-RFX5_MA0510.3 14 bp overlap
Motif DE_48h DE_48h-RFX5_MA0510.3 14 bp overlap
Motif DE_60h DE_60h-RFX5_MA0510.3 14 bp overlap
Motif DE_72h DE_72h-RFX5_MA0510.3 14 bp overlap
RFX7 1 dataset
Motif DE_60h DE_60h-RFX7_MA1554.2 8 bp overlap
RHOXF1 4 datasets
Motif DE_36h DE_36h-RHOXF1_MA0719.2 6 bp overlap
Motif DE_48h DE_48h-RHOXF1_MA0719.2 6 bp overlap
Motif DE_60h DE_60h-RHOXF1_MA0719.2 6 bp overlap
Motif DE_72h DE_72h-RHOXF1_MA0719.2 6 bp overlap
RREB1 1 dataset
Motif DE_60h DE_60h-RREB1_MA0073.2 19 bp overlap
Rfx6 4 datasets
Motif DE_36h DE_36h-Rfx6_MA1724.2 9 bp overlap
Motif DE_48h DE_48h-Rfx6_MA1724.2 9 bp overlap
Motif DE_60h DE_60h-Rfx6_MA1724.2 9 bp overlap
Motif DE_72h DE_72h-Rfx6_MA1724.2 9 bp overlap
SMAD2 1 dataset
ChIP endoderm GSE29422.SMAD2.endoderm 162 bp overlap
SMAD2-3 2 datasets
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 633 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.SMAD2-3.HUES-8_DE_JNKi 417 bp overlap
SMAD2_3 3 datasets
ChIP DE_D1 S12-DE-d1-SMAD2_3-exp1 330 bp overlap
ChIP DE_D2 S01-DE-d2-SMAD2_3-exp1 509 bp overlap
ChIP DE_D2 S53-DE-d2-SMAD2_3-exp2 535 bp overlap
SMAD4 1 dataset
ChIP endoderm GSE29422.SMAD4.endoderm 140 bp overlap
SMARCA4 3 datasets
ChIP 501-mel_SHSOX10 GSE61965.SMARCA4.501-mel_SHSOX10 118 bp overlap
ChIP A-549_AG15677 GSE132290.SMARCA4.A-549_AG15677 96 bp overlap
ChIP A-549_AG15685 GSE132290.SMARCA4.A-549_AG15685 106 bp overlap
SNAI2 1 dataset
ChIP RD_shSNAI2 GSE137168.SNAI2.RD_shSNAI2 229 bp overlap
SOX10 1 dataset
Motif DE_60h DE_60h-SOX10_MA0442.3 6 bp overlap
SOX12 1 dataset
Motif DE_60h DE_60h-SOX12_MA1561.2 10 bp overlap
SOX4 1 dataset
Motif DE_60h DE_60h-SOX4_MA0867.3 8 bp overlap
SP3 1 dataset
Motif DE_60h DE_60h-SP3_MA0746.3 11 bp overlap
SP8 1 dataset
Motif DE_60h DE_60h-SP8_MA0747.2 11 bp overlap
SP9 1 dataset
Motif DE_60h DE_60h-SP9_MA1564.2 10 bp overlap
Sox5 1 dataset
Motif DE_60h DE_60h-Sox5_MA0087.3 8 bp overlap
ZNF24 1 dataset
Motif DE_60h DE_60h-ZNF24_MA1124.1 13 bp overlap
ZNF418 1 dataset
Motif DE_60h DE_60h-ZNF418_MA1980.1 15 bp overlap
ZNF701 4 datasets
Motif DE_36h DE_36h-ZNF701_MA1987.2 17 bp overlap
Motif DE_48h DE_48h-ZNF701_MA1987.2 17 bp overlap
Motif DE_60h DE_60h-ZNF701_MA1987.2 17 bp overlap
Motif DE_72h DE_72h-ZNF701_MA1987.2 17 bp overlap
ZNF768 1 dataset
Motif DE_60h DE_60h-ZNF768_MA1731.2 9 bp overlap