chr4 : 144,647,743 144,648,344
601 bp 75 TFs 2 linked genes
This 601 bp open chromatin element is linked to HHIP and HHIP-AS1 and is bound by 75 transcription factors.
Linked Genes
2 genes
Gene Expression Dist. to TSS Distance Link type
HHIP 1.6 kb Proximal Proximity
HHIP-AS1 1.8 kb Proximal Proximity
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr4:144,642,743 – 144,653,344
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
75 transcription factors
Source
Cell type
AR 4 datasets
ChIP LNCaP_DHT GSE125245.AR.LNCaP_DHT 73 bp overlap
ChIP LNCaP_SHCTR_DHT GSE62492.AR.LNCaP_SHCTR_DHT 58 bp overlap
ChIP LNCaP_Talen_DHT GSE89938.AR.LNCaP_Talen_DHT 50 bp overlap
ChIP LNCaP_sgCHD1 GSE117430.AR.LNCaP_sgCHD1 50 bp overlap
ARID1A 1 dataset
ChIP HAP1 GSE108387.ARID1A.HAP1 238 bp overlap
ASXL3 1 dataset
ChIP NCI-H1963_Ab1 GSE145028.ASXL3.NCI-H1963_Ab1 284 bp overlap
ATF2 3 datasets
ChIP HEK293 ENCFF194VKZ 385 bp overlap
ChIP HEK293 ENCSR217HTK.ATF2.HEK293 385 bp overlap
ChIP WA01 ENCSR000BQU.ATF2.WA01 346 bp overlap
ATF3 2 datasets
ChIP HCT-116_DMSO GSE74355.ATF3.HCT-116_DMSO 108 bp overlap
ChIP K-562 ENCSR028UIU.ATF3.K-562 229 bp overlap
ATF4 9 datasets
ChIP HUDEP-2 GSE143961.ATF4.HUDEP-2 181 bp overlap
ChIP HUDEP-2_KO GSE143961.ATF4.HUDEP-2_KO 235 bp overlap
ChIP HepG2 ENCFF903ADR 400 bp overlap
ChIP Jurkat_ZBTB1-KO GSE145783.ATF4.Jurkat_ZBTB1-KO 333 bp overlap
ChIP Jurkat_ZBTB1-KO_Asp-deprivation GSE145783.ATF4.Jurkat_ZBTB1-KO_Asp-deprivation 488 bp overlap
ChIP Jurkat_ZBTB1-KO_Asp-deprivation_cDNA GSE145783.ATF4.Jurkat_ZBTB1-KO_Asp-deprivation_cDNA 453 bp overlap
ChIP Jurkat_ZBTB1-KO_cDNA GSE145783.ATF4.Jurkat_ZBTB1-KO_cDNA 348 bp overlap
ChIP K-562 ENCSR145TSJ.ATF4.K-562 397 bp overlap
ChIP K562 ENCFF674KTF 372 bp overlap
ATF7 3 datasets
ChIP Hep-G2 ENCSR545FXC.ATF7.Hep-G2 241 bp overlap
ChIP HepG2 ENCFF470FKK 381 bp overlap
ChIP MCF-7 ENCSR866QPZ.ATF7.MCF-7 224 bp overlap
BCOR 2 datasets
ChIP WA01_KDM2B_KO GSE104690.BCOR.WA01_KDM2B_KO 193 bp overlap
ChIP WA01_RNF2-R_Doxycyclin GSE104690.BCOR.WA01_RNF2-R_Doxycyclin 309 bp overlap
BRD2 2 datasets
ChIP SUM159PT_MGSK2801 GSE116879.BRD2.SUM159PT_MGSK2801 71 bp overlap
ChIP SUM159PT_MGSK2801 GSE116879.BRD2.SUM159PT_MGSK2801 350 bp overlap
BRD4 8 datasets
ChIP COLO-320 GSE73319.BRD4.COLO-320 301 bp overlap
ChIP HAP1 GSE108387.BRD4.HAP1 106 bp overlap
ChIP HUVEC-C_modETS1 GSE93030.BRD4.HUVEC-C_modETS1 236 bp overlap
ChIP RH4_shscr GSE140115.BRD4.RH4_shscr 167 bp overlap
ChIP SUM159PT_DMSO_24h GSE87418.BRD4.SUM159PT_DMSO_24h 301 bp overlap
ChIP SUM159PT_Dox_48h GSE87418.BRD4.SUM159PT_Dox_48h 254 bp overlap
ChIP SUM159PT_KO GSE131097.BRD4.SUM159PT_KO 319 bp overlap
ChIP SUM159PT_shMYC_DMSO_48h GSE87418.BRD4.SUM159PT_shMYC_DMSO_48h 318 bp overlap
CDK7 1 dataset
ChIP Jurkat GSE50622.CDK7.Jurkat 574 bp overlap
CDK9 1 dataset
ChIP MOLT-4_DMSO GSE79288.CDK9.MOLT-4_DMSO 82 bp overlap
CEBPB 8 datasets
ChIP A-549 ENCSR000BUB.CEBPB.A-549 116 bp overlap
ChIP A549 ENCFF235AIY 257 bp overlap
ChIP H1 ENCFF871PTR 261 bp overlap
ChIP HepG2 ENCFF536NTI 221 bp overlap
ChIP IMR-90 ENCFF468UGY 251 bp overlap
ChIP Ishikawa ENCSR000BTT.CEBPB.Ishikawa 133 bp overlap
ChIP K-562 ENCSR000EHE.CEBPB.K-562 131 bp overlap
ChIP WA01 ENCSR000EBV.CEBPB.WA01 236 bp overlap
CEBPD 1 dataset
ChIP HAEC_IL1b_4h GSE89970.CEBPD.HAEC_IL1b_4h 236 bp overlap
CHD1 1 dataset
ChIP hMSC-TERT GSE89179.CHD1.hMSC-TERT 537 bp overlap
CHD4 1 dataset
ChIP SCMC GSE155861.CHD4.SCMC 205 bp overlap
CHD7 2 datasets
ChIP H1 ENCFF126NLU 269 bp overlap
ChIP hiPSC_derived_lt-NES GSE108506.CHD7.hiPSC_derived_lt-NES 349 bp overlap
CTCF 1 dataset
ChIP dorsolateral prefrontal cortex ENCFF277CZQ 174 bp overlap
Cebpa 2 datasets
ChIP BLaER1 ENCFF140EYR 367 bp overlap
ChIP BLaER1 ENCFF140EYR 165 bp overlap
EP300 1 dataset
ChIP hESC GSE17917.EP300.hESC 297 bp overlap
ETS1 2 datasets
ChIP THP-6_shCtrl GSE138516.ETS1.THP-6_shCtrl 110 bp overlap
ChIP THP-6_shEts1 GSE138516.ETS1.THP-6_shEts1 265 bp overlap
EZH2 1 dataset
ChIP Jurkat_KO GSE147198.EZH2.Jurkat_KO 57 bp overlap
FOXA1 3 datasets
ChIP 22Rv1_R1881 GSE80742.FOXA1.22Rv1_R1881 289 bp overlap
ChIP LAPC-4_TFS_p358fs-V5 GSE123618.FOXA1.LAPC-4_TFS_p358fs-V5 103 bp overlap
ChIP PDAC_KOKLF5 GSE64557.FOXA1.PDAC_KOKLF5 331 bp overlap
FOXA2 2 datasets
ChIP BJ1-hTERT_MimoRelease GSE92491.FOXA2.BJ1-hTERT_MimoRelease 92 bp overlap
ChIP DE DE-FOXA2-1 329 bp overlap
GABPB1 1 dataset
ChIP WTC11 ENCFF166QKI 401 bp overlap
GATA4 1 dataset
ChIP BJ1-hTERT GSE92491.GATA4.BJ1-hTERT 116 bp overlap
IKZF3 1 dataset
ChIP HEK293 ENCSR304AMN.IKZF3.HEK293 249 bp overlap
JUND 1 dataset
ChIP liver ENCSR196HGZ.JUND.liver 201 bp overlap
KLF16 1 dataset
ChIP HEK293 ENCSR397DQC.KLF16.HEK293 368 bp overlap
KLF4 1 dataset
ChIP HAP1 GSE130417.KLF4.HAP1 207 bp overlap
KLF9 1 dataset
ChIP GBM1A GSE62211.KLF9.GBM1A 86 bp overlap
KMT2A 7 datasets
ChIP CCRF-CEM GSE83671.KMT2A.CCRF-CEM 601 bp overlap
ChIP HEK293T_N-term GSE90762.KMT2A.HEK293T_N-term 392 bp overlap
ChIP HEK293T_N-term_C104 GSE90762.KMT2A.HEK293T_N-term_C104 294 bp overlap
ChIP HEK293T_N-term_shCtrl_TASP1-KO GSE90762.KMT2A.HEK293T_N-term_shCtrl_TASP1-KO 95 bp overlap
ChIP KOPN-8 GSE83671.KMT2A.KOPN-8 231 bp overlap
ChIP KOPN-8 GSE83671.KMT2A.KOPN-8 181 bp overlap
ChIP MOLM-13_CBS79-KO-VP-HOTTIP GSE114981.KMT2A.MOLM-13_CBS79-KO-VP-HOTTIP 601 bp overlap
MED1 2 datasets
ChIP HUVEC-C_siERG GSE124892.MED1.HUVEC-C_siERG 596 bp overlap
ChIP Jurkat GSE59657.MED1.Jurkat 496 bp overlap
MEN1 1 dataset
ChIP PC-3 GSE132827.MEN1.PC-3 528 bp overlap
MYB 2 datasets
ChIP MOLT-3 GSE59657.MYB.MOLT-3 88 bp overlap
ChIP MOLT-3 GSE59657.MYB.MOLT-3 173 bp overlap
MZF1 1 dataset
ChIP HEK293 ENCSR298QUH.MZF1.HEK293 388 bp overlap
NANOG 8 datasets
ChIP GM23338 ENCSR061DGF.NANOG.GM23338 267 bp overlap
ChIP HUES-8 GSE109524.NANOG.HUES-8 601 bp overlap
ChIP WA01 ERP004238.NANOG.WA01 310 bp overlap
ChIP WA01_3IL ERP004238.NANOG.WA01_3IL 363 bp overlap
ChIP WA09 GSE105028.NANOG.WA09 510 bp overlap
ChIP WA09_heat-shock GSE105028.NANOG.WA09_heat-shock 262 bp overlap
ChIP hESC GSE18292.NANOG.hESC 260 bp overlap
ChIP hESC GSE20650.NANOG.hESC 245 bp overlap
OLIG2 1 dataset
ChIP brain-prefrontal-cortex_2017011 GSE129039.OLIG2.brain-prefrontal-cortex_2017011 225 bp overlap
ONECUT1 5 datasets
ChIP H9 ERP004206.ONECUT1.H9 308 bp overlap
ChIP Hep-G2 ENCSR956OSX.ONECUT1.Hep-G2 227 bp overlap
ChIP HepG2 ENCFF243FIR 341 bp overlap
ChIP liver ERP002306.ONECUT1.liver 229 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.ONECUT1.pancreatic-progenitor_PP1 422 bp overlap
PATZ1 2 datasets
ChIP HEK293 ENCFF016MNJ 434 bp overlap
ChIP HEK293 ENCSR966ULI.PATZ1.HEK293 316 bp overlap
POLR2A 4 datasets
ChIP endothelial cell of umbilical vein ENCFF131DWO 201 bp overlap
ChIP endothelial cell of umbilical vein ENCFF303XUJ 429 bp overlap
ChIP endothelial cell of umbilical vein ENCFF467WJF 187 bp overlap
ChIP endothelial cell of umbilical vein ENCFF467WJF 322 bp overlap
POU5F1 7 datasets
ChIP GM23338 ENCFF333SNB 311 bp overlap
ChIP HUES-8 GSE109524.POU5F1.HUES-8 601 bp overlap
ChIP NCCIT GSE36134.POU5F1.NCCIT 533 bp overlap
ChIP hESC ENCSR264RJX.POU5F1.hESC 306 bp overlap
ChIP hESC_NAIVE GSE69646.POU5F1.hESC_NAIVE 300 bp overlap
ChIP hESC_PRIMED GSE69646.POU5F1.hESC_PRIMED 463 bp overlap
ChIP neuron_bipolar_doxy_4d ENCSR362VCG.POU5F1.neuron_bipolar_doxy_4d 130 bp overlap
PRDM10 1 dataset
ChIP HEK293 ENCFF145WQQ 223 bp overlap
PRDM6 1 dataset
ChIP HEK293 ENCSR892QHR.PRDM6.HEK293 248 bp overlap
RAD21 1 dataset
ChIP HAP1 GSE152721.RAD21.HAP1 51 bp overlap
RARA 1 dataset
ChIP hiPSC_D4 GSE109172.RARA.hiPSC_D4 334 bp overlap
REST 1 dataset
ChIP HEK293 ENCSR896UBV.REST.HEK293 195 bp overlap
RING1 1 dataset
ChIP SYO-1_shRING1A-B GSE139053.RING1.SYO-1_shRING1A-B 50 bp overlap
RNF2 4 datasets
ChIP HUES-64 GSE104059.RNF2.HUES-64 118 bp overlap
ChIP SYO-1_shRING1A-B GSE139053.RNF2.SYO-1_shRING1A-B 200 bp overlap
ChIP SYO-1_shRING1A-B GSE139053.RNF2.SYO-1_shRING1A-B 289 bp overlap
ChIP fibroblast GSE139053.RNF2.fibroblast 250 bp overlap
RUNX1 1 dataset
ChIP Jurkat GSE68976.RUNX1.Jurkat 236 bp overlap
SIN3A 1 dataset
ChIP HUVEC-C GSE103245.SIN3A.HUVEC-C 145 bp overlap
SMAD2-3 2 datasets
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 338 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.SMAD2-3.HUES-8_DE_ctrl 317 bp overlap
SMAD3 1 dataset
ChIP SUM159PT GSE130364.SMAD3.SUM159PT 429 bp overlap
SMARCA4 6 datasets
ChIP NSC GSE125033.SMARCA4.NSC 578 bp overlap
ChIP TTC-1240_SMARCB1-FL GSE124903.SMARCA4.TTC-1240_SMARCB1-FL 267 bp overlap
ChIP TTC-1240_delC GSE124903.SMARCA4.TTC-1240_delC 158 bp overlap
ChIP endometrial-epithelial-cells_KO GSE106660.SMARCA4.endometrial-epithelial-cells_KO 264 bp overlap
ChIP hiPSC GSE124903.SMARCA4.hiPSC 373 bp overlap
ChIP hiPSC_WT GSE124903.SMARCA4.hiPSC_WT 422 bp overlap
SMARCC1 4 datasets
ChIP SYO-1 GSE108025.SMARCC1.SYO-1 341 bp overlap
ChIP SYO-1_shRING1A-B GSE139053.SMARCC1.SYO-1_shRING1A-B 425 bp overlap
ChIP TTC-1240_empty GSE124903.SMARCC1.TTC-1240_empty 423 bp overlap
ChIP hiPSC_WT GSE124903.SMARCC1.hiPSC_WT 286 bp overlap
SOX2 2 datasets
ChIP HNSC GSE69479.SOX2.HNSC 440 bp overlap
ChIP hiPSC GSE56567.SOX2.hiPSC 355 bp overlap
SS18 2 datasets
ChIP SYO-1_shRING1A-B GSE139053.SS18.SYO-1_shRING1A-B 187 bp overlap
ChIP SYO-1_shRING1A-B GSE139053.SS18.SYO-1_shRING1A-B 266 bp overlap
SS18-SSX 1 dataset
ChIP fibroblast_78aa-SSX-Tail GSE139053.SS18-SSX.fibroblast_78aa-SSX-Tail 290 bp overlap
SUPT5H 2 datasets
ChIP MOLT-4_SNS GSE89384.SUPT5H.MOLT-4_SNS 97 bp overlap
ChIP MOLT-4_SNS GSE89384.SUPT5H.MOLT-4_SNS 131 bp overlap
SUZ12 2 datasets
ChIP HEK293T_PCGF1352fl GSE119618.SUZ12.HEK293T_PCGF1352fl 418 bp overlap
ChIP HEK293T_PCGF2fl GSE119618.SUZ12.HEK293T_PCGF2fl 308 bp overlap
TAL1 1 dataset
ChIP MOLT-3 GSE59657.TAL1.MOLT-3 220 bp overlap
TCF3 1 dataset
ChIP NPC GSE154479.TCF3.NPC 253 bp overlap
YY1 1 dataset
ChIP ALL GSE145549.YY1.ALL 222 bp overlap
ZEB2 2 datasets
ChIP HEK293 ENCFF847JIE 456 bp overlap
ChIP HEK293 ENCSR417VWF.ZEB2.HEK293 588 bp overlap
ZFHX2 1 dataset
ChIP HEK293 ENCFF167TUA 299 bp overlap
ZMIZ1 1 dataset
ChIP THP-6_shCtrl GSE138516.ZMIZ1.THP-6_shCtrl 269 bp overlap
ZNF2 1 dataset
ChIP HEK293 ENCSR011CKE.ZNF2.HEK293 262 bp overlap
ZNF35 1 dataset
ChIP HEK293 GSE76494.ZNF35.HEK293 276 bp overlap
ZNF585A 1 dataset
ChIP HEK293T GSE78099.ZNF585A.HEK293T 225 bp overlap
ZNF90 1 dataset
ChIP HEK293T GSE78099.ZNF90.HEK293T 107 bp overlap