chr13 : 39,201,023 39,201,763
740 bp 125 TFs 2 linked genes
This 740 bp open chromatin element is linked to NHLRC3 and PROSER1 and is bound by 125 transcription factors.
Linked Genes
2 genes
Gene Expression Dist. to TSS Distance Link type
NHLRC3 163.2 kb Distal Multiome
PROSER1 163.4 kb Distal Multiome
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr13:39,196,023 – 39,206,763
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
125 transcription factors
Source
Cell type
ATF2 2 datasets
Motif DE_36h DE_36h-ATF2_MA1632.2 10 bp overlap
Motif DE_48h DE_48h-ATF2_MA1632.2 10 bp overlap
Ahr::Arnt 2 datasets
Motif DE_36h DE_36h-AhrArnt_MA0006.2 5 bp overlap
Motif DE_48h DE_48h-AhrArnt_MA0006.2 5 bp overlap
BCL6B 2 datasets
Motif DE_36h DE_36h-BCL6B_MA0731.1 17 bp overlap
Motif DE_48h DE_48h-BCL6B_MA0731.1 17 bp overlap
BRD1 1 dataset
ChIP RKO GSE47190.BRD1.RKO 119 bp overlap
CREB1 2 datasets
Motif DE_36h DE_36h-CREB1_MA0018.5 8 bp overlap
Motif DE_48h DE_48h-CREB1_MA0018.5 8 bp overlap
CTCF 2 datasets
Motif DE_36h DE_36h-CTCF_MA0139.2 15 bp overlap
Motif DE_36h DE_36h-CTCF_MA1929.2 31 bp overlap
EHF 1 dataset
Motif DE_36h DE_36h-EHF_MA0598.4 9 bp overlap
ELF1 1 dataset
Motif DE_36h DE_36h-ELF1_MA0473.4 9 bp overlap
ELF3 1 dataset
Motif DE_36h DE_36h-ELF3_MA0640.3 9 bp overlap
EOMES 3 datasets
Motif DE_36h DE_36h-EOMES_MA0800.2 9 bp overlap
Motif DE_48h DE_48h-EOMES_MA0800.2 9 bp overlap
ChIP hESC GSE26097.EOMES.hESC 527 bp overlap
ERF::HOXB13 2 datasets
Motif DE_36h DE_36h-ERFHOXB13_MA1937.2 13 bp overlap
Motif DE_48h DE_48h-ERFHOXB13_MA1937.2 13 bp overlap
ESRRG 1 dataset
ChIP cardiomyocyte GSE113760.ESRRG.cardiomyocyte 273 bp overlap
ETV1 1 dataset
Motif DE_36h DE_36h-ETV1_MA0761.3 9 bp overlap
Elf5 2 datasets
Motif DE_36h DE_36h-Elf5_MA0136.4 8 bp overlap
Motif DE_48h DE_48h-Elf5_MA0136.4 8 bp overlap
Erg 1 dataset
Motif DE_36h DE_36h-Erg_MA0474.4 10 bp overlap
FLI1 1 dataset
ChIP A-673_D11 GSE129155.FLI1.A-673_D11 220 bp overlap
FOXA2 3 datasets
ChIP BJ1-hTERT_FOXA2_GATA4_Coexp GSE92491.FOXA2.BJ1-hTERT_FOXA2_GATA4_Coexp 240 bp overlap
ChIP DE DE-FOXA2-1 601 bp overlap
ChIP DE DE-FOXA2-2 358 bp overlap
GABPA 1 dataset
Motif DE_36h DE_36h-GABPA_MA0062.4 10 bp overlap
GATA1 2 datasets
Motif DE_36h DE_36h-GATA1_MA0035.5 7 bp overlap
Motif DE_48h DE_48h-GATA1_MA0035.5 7 bp overlap
GATA2 2 datasets
Motif DE_36h DE_36h-GATA2_MA0036.4 7 bp overlap
Motif DE_48h DE_48h-GATA2_MA0036.4 7 bp overlap
GATA3 2 datasets
ChIP MCF-7_E2 GSE81510.GATA3.MCF-7_E2 179 bp overlap
ChIP MCF-7_E2_Dex GSE81510.GATA3.MCF-7_E2_Dex 186 bp overlap
GATA4 7 datasets
ChIP DE DE-GATA4-1 667 bp overlap
ChIP DE DE-GATA4-2 740 bp overlap
Motif DE_36h DE_36h-GATA4_MA0482.3 8 bp overlap
Motif DE_48h DE_48h-GATA4_MA0482.3 8 bp overlap
ChIP Hep-G2 GSE135714.GATA4.Hep-G2 240 bp overlap
ChIP foregut GSE117136.GATA4.foregut 526 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA4.pancreatic-progenitor 404 bp overlap
GATA5 2 datasets
Motif DE_36h DE_36h-GATA5_MA0766.3 8 bp overlap
Motif DE_48h DE_48h-GATA5_MA0766.3 8 bp overlap
GATA6 20 datasets
ChIP AGS GSE51705.GATA6.AGS 191 bp overlap
ChIP Caco-2_PROLIF GSE23436.GATA6.Caco-2_PROLIF 132 bp overlap
ChIP DE DE-GATA6-1 740 bp overlap
ChIP DE DE-GATA6-2 740 bp overlap
Motif DE_36h DE_36h-GATA6_MA1104.3 8 bp overlap
Motif DE_36h DE_36h-GATA6_MA1104.3 8 bp overlap
Motif DE_48h DE_48h-GATA6_MA1104.3 8 bp overlap
Motif DE_48h DE_48h-GATA6_MA1104.3 8 bp overlap
ChIP DE_D1 S09-DE-d1-GATA6-exp1 678 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 675 bp overlap
ChIP DE_D1 S41-DE-d1-GATA6-exp2 740 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 703 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 740 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 740 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 740 bp overlap
ChIP foregut GSE117136.GATA6.foregut 567 bp overlap
ChIP foregut_KO GSE117136.GATA6.foregut_KO 362 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA6.pancreatic-progenitor 405 bp overlap
ChIP pancreatic-progenitor_KO GSE117136.GATA6.pancreatic-progenitor_KO 434 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA6.pancreatic-progenitor_PP1 225 bp overlap
GLI3 1 dataset
Motif DE_36h DE_36h-GLI3_MA1491.3 15 bp overlap
Gata3 4 datasets
Motif DE_36h DE_36h-Gata3_MA0037.5 8 bp overlap
Motif DE_36h DE_36h-Gata3_MA0037.5 8 bp overlap
Motif DE_48h DE_48h-Gata3_MA0037.5 8 bp overlap
Motif DE_48h DE_48h-Gata3_MA0037.5 8 bp overlap
Gli1 1 dataset
Motif DE_36h DE_36h-Gli1_MA1990.2 10 bp overlap
HIC2 2 datasets
Motif DE_36h DE_36h-HIC2_MA0738.2 6 bp overlap
Motif DE_48h DE_48h-HIC2_MA0738.2 6 bp overlap
Hand1 2 datasets
Motif DE_36h DE_36h-Hand1_MA2123.1 9 bp overlap
Motif DE_48h DE_48h-Hand1_MA2123.1 9 bp overlap
Hnf1A 2 datasets
Motif DE_36h DE_36h-Hnf1A_MA1991.2 10 bp overlap
Motif DE_48h DE_48h-Hnf1A_MA1991.2 10 bp overlap
IKZF1 1 dataset
Motif DE_36h DE_36h-IKZF1_MA1508.2 8 bp overlap
IKZF2 2 datasets
Motif DE_36h DE_36h-IKZF2_MA2326.1 6 bp overlap
Motif DE_48h DE_48h-IKZF2_MA2326.1 6 bp overlap
IRF4 3 datasets
Motif DE_24h DE_24h-IRF4_MA1419.2 14 bp overlap
Motif DE_36h DE_36h-IRF4_MA1419.2 14 bp overlap
Motif DE_48h DE_48h-IRF4_MA1419.2 14 bp overlap
IRF8 3 datasets
Motif DE_24h DE_24h-IRF8_MA0652.2 13 bp overlap
Motif DE_36h DE_36h-IRF8_MA0652.2 13 bp overlap
Motif DE_48h DE_48h-IRF8_MA0652.2 13 bp overlap
Irf1 5 datasets
Motif DE_24h DE_24h-Irf1_MA0050.4 11 bp overlap
Motif DE_36h DE_36h-Irf1_MA0050.4 11 bp overlap
Motif DE_36h DE_36h-Irf1_MA0050.4 11 bp overlap
Motif DE_48h DE_48h-Irf1_MA0050.4 11 bp overlap
Motif DE_48h DE_48h-Irf1_MA0050.4 11 bp overlap
JUN 5 datasets
ChIP DE_D1 S08-DE-d1-JUN-exp1 514 bp overlap
ChIP DE_D1 S13-DE-d1-JUN-exp1 341 bp overlap
ChIP DE_D1 S40-DE-d1-JUN-exp2 526 bp overlap
ChIP DE_D2 S02-DE-d2-JUN-exp1 536 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.JUN.HUES-8_DE_ctrl 567 bp overlap
MED1 1 dataset
ChIP RH4 GSE83726.MED1.RH4 367 bp overlap
MEIS1 4 datasets
Motif DE_36h DE_36h-MEIS1_MA0498.3 5 bp overlap
Motif DE_36h DE_36h-MEIS1_MA0498.3 5 bp overlap
Motif DE_48h DE_48h-MEIS1_MA0498.3 5 bp overlap
Motif DE_48h DE_48h-MEIS1_MA0498.3 5 bp overlap
MXI1 2 datasets
Motif DE_36h DE_36h-MXI1_MA1108.3 6 bp overlap
Motif DE_48h DE_48h-MXI1_MA1108.3 6 bp overlap
MYB 3 datasets
Motif DE_36h DE_36h-MYB_MA0100.4 6 bp overlap
Motif DE_48h DE_48h-MYB_MA0100.4 6 bp overlap
ChIP GM12878 ENCSR819ATC.MYB.GM12878 142 bp overlap
MYOG 1 dataset
ChIP RH30_DMSO GSE85169.MYOG.RH30_DMSO 152 bp overlap
Mecom 2 datasets
Motif DE_36h DE_36h-Mecom_MA0029.2 11 bp overlap
Motif DE_48h DE_48h-Mecom_MA0029.2 11 bp overlap
NFATC3 2 datasets
Motif DE_36h DE_36h-NFATC3_MA0625.3 6 bp overlap
Motif DE_48h DE_48h-NFATC3_MA0625.3 6 bp overlap
NKX2-2 1 dataset
Motif DE_36h DE_36h-NKX2-2_MA1645.2 8 bp overlap
NKX2-4 2 datasets
Motif DE_36h DE_36h-NKX2-4_MA2003.2 8 bp overlap
Motif DE_48h DE_48h-NKX2-4_MA2003.2 8 bp overlap
NKX2-5 3 datasets
Motif DE_36h DE_36h-NKX2-5_MA0063.3 7 bp overlap
ChIP hESC_ab3584 GSE89457.NKX2-5.hESC_ab3584 271 bp overlap
ChIP hESC_sc-14033 GSE89457.NKX2-5.hESC_sc-14033 177 bp overlap
NKX2-8 2 datasets
Motif DE_36h DE_36h-NKX2-8_MA0673.2 8 bp overlap
Motif DE_48h DE_48h-NKX2-8_MA0673.2 8 bp overlap
NR2C1 2 datasets
Motif DE_36h DE_36h-NR2C1_MA1535.2 6 bp overlap
Motif DE_48h DE_48h-NR2C1_MA1535.2 6 bp overlap
NR2C2 2 datasets
Motif DE_36h DE_36h-NR2C2_MA1536.2 6 bp overlap
Motif DE_48h DE_48h-NR2C2_MA1536.2 6 bp overlap
NR2F1 2 datasets
Motif DE_36h DE_36h-NR2F1_MA0017.3 12 bp overlap
Motif DE_48h DE_48h-NR2F1_MA0017.3 12 bp overlap
NR2F2 1 dataset
ChIP liver ENCSR168SMX.NR2F2.liver 243 bp overlap
NR3C1 1 dataset
ChIP SUP-B15_DEX GSE107584.NR3C1.SUP-B15_DEX 302 bp overlap
Nfat5 2 datasets
Motif DE_36h DE_36h-Nfat5_MA0606.3 8 bp overlap
Motif DE_48h DE_48h-Nfat5_MA0606.3 8 bp overlap
Nfatc1 2 datasets
Motif DE_36h DE_36h-Nfatc1_MA0624.3 6 bp overlap
Motif DE_48h DE_48h-Nfatc1_MA0624.3 6 bp overlap
Nr1H2 2 datasets
Motif DE_36h DE_36h-Nr1H2_MA1996.2 6 bp overlap
Motif DE_48h DE_48h-Nr1H2_MA1996.2 6 bp overlap
Nr1H4 2 datasets
Motif DE_36h DE_36h-Nr1H4_MA1110.3 6 bp overlap
Motif DE_48h DE_48h-Nr1H4_MA1110.3 6 bp overlap
Nr1h3 2 datasets
Motif DE_36h DE_36h-Nr1h3_MA2337.1 6 bp overlap
Motif DE_48h DE_48h-Nr1h3_MA2337.1 6 bp overlap
Nr2f6 2 datasets
Motif DE_36h DE_36h-Nr2f6_MA0677.2 13 bp overlap
Motif DE_48h DE_48h-Nr2f6_MA0677.2 13 bp overlap
PAX7 1 dataset
ChIP H9_DOX GSE98976.PAX7.H9_DOX 212 bp overlap
PHOX2A 2 datasets
Motif DE_36h DE_36h-PHOX2A_MA0713.1 11 bp overlap
Motif DE_48h DE_48h-PHOX2A_MA0713.1 11 bp overlap
POU1F1 1 dataset
Motif DE_36h DE_36h-POU1F1_MA0784.3 14 bp overlap
POU2F2 1 dataset
Motif DE_36h DE_36h-POU2F2_MA0507.3 13 bp overlap
POU2F3 2 datasets
Motif DE_36h DE_36h-POU2F3_MA0627.3 9 bp overlap
Motif DE_48h DE_48h-POU2F3_MA0627.3 9 bp overlap
POU3F2 1 dataset
Motif DE_36h DE_36h-POU3F2_MA0787.1 12 bp overlap
POU5F1 3 datasets
Motif DE_36h DE_36h-POU5F1_MA1115.2 7 bp overlap
Motif DE_48h DE_48h-POU5F1_MA1115.2 7 bp overlap
ChIP DE_D1 DED1-OCT4_Batch_II 400 bp overlap
PRDM1 2 datasets
Motif DE_36h DE_36h-PRDM1_MA0508.4 7 bp overlap
Motif DE_48h DE_48h-PRDM1_MA0508.4 7 bp overlap
RBPJ 3 datasets
Motif DE_36h DE_36h-RBPJ_MA1116.2 6 bp overlap
Motif DE_36h DE_36h-RBPJ_MA1116.2 6 bp overlap
Motif DE_48h DE_48h-RBPJ_MA1116.2 6 bp overlap
RUNX1 1 dataset
ChIP MV4-11 GSE79899.RUNX1.MV4-11 194 bp overlap
SMAD2 2 datasets
Motif DE_36h DE_36h-SMAD2_MA1964.2 6 bp overlap
Motif DE_48h DE_48h-SMAD2_MA1964.2 6 bp overlap
SMAD2-3 3 datasets
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 659 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.SMAD2-3.HUES-8_DE_JNKi 740 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.SMAD2-3.HUES-8_DE_ctrl 648 bp overlap
SMAD2_3 5 datasets
ChIP DE_D1 S07-DE-d1-SMAD2_3-exp1 590 bp overlap
ChIP DE_D1 S12-DE-d1-SMAD2_3-exp1 740 bp overlap
ChIP DE_D1 S39-DE-d1-SMAD2_3-exp2 643 bp overlap
ChIP DE_D2 S01-DE-d2-SMAD2_3-exp1 532 bp overlap
ChIP DE_D2 S53-DE-d2-SMAD2_3-exp2 603 bp overlap
SMARCA4 1 dataset
ChIP 501-mel_SHSOX10 GSE61965.SMARCA4.501-mel_SHSOX10 236 bp overlap
SMARCC1 2 datasets
ChIP DE_D1 S10-DE-d1-BAF155-exp1 326 bp overlap
ChIP DE_D1 S15-DE-d1-BAF155-exp1 704 bp overlap
SNAI2 2 datasets
ChIP SMS-CTR_D48 GSE137168.SNAI2.SMS-CTR_D48 165 bp overlap
ChIP SMS-CTR_D48 GSE137168.SNAI2.SMS-CTR_D48 200 bp overlap
SOX17 1 dataset
ChIP DE_D2 DED2-SOX17_Batch_II 568 bp overlap
SOX17_M 1 dataset
ChIP DE_D2 DED2-SOX17-M_Batch_II 399 bp overlap
SPI1 1 dataset
ChIP primary-monocyte_donorA_ncMO GSE128834.SPI1.primary-monocyte_donorA_ncMO 141 bp overlap
SPIB 2 datasets
Motif DE_36h DE_36h-SPIB_MA0081.3 13 bp overlap
Motif DE_48h DE_48h-SPIB_MA0081.3 13 bp overlap
STAT1 1 dataset
Motif DE_36h DE_36h-STAT1_MA0137.4 9 bp overlap
STAT3 1 dataset
Motif DE_36h DE_36h-STAT3_MA0144.3 9 bp overlap
Spz1 1 dataset
Motif DE_36h DE_36h-Spz1_MA0111.1 11 bp overlap
Stat2 3 datasets
Motif DE_24h DE_24h-Stat2_MA1623.2 10 bp overlap
Motif DE_36h DE_36h-Stat2_MA1623.2 10 bp overlap
Motif DE_48h DE_48h-Stat2_MA1623.2 10 bp overlap
Stat4 1 dataset
Motif DE_36h DE_36h-Stat4_MA0518.2 10 bp overlap
Stat5b 2 datasets
Motif DE_36h DE_36h-Stat5b_MA1625.2 9 bp overlap
Motif DE_48h DE_48h-Stat5b_MA1625.2 9 bp overlap
T 2 datasets
ChIP H9_ENDODERM GSE60606.T.H9_ENDODERM 226 bp overlap
ChIP H9_MESODERM GSE60606.T.H9_MESODERM 183 bp overlap
TBR1 2 datasets
Motif DE_36h DE_36h-TBR1_MA0802.2 9 bp overlap
Motif DE_48h DE_48h-TBR1_MA0802.2 9 bp overlap
TBX18 2 datasets
Motif DE_36h DE_36h-TBX18_MA1565.2 9 bp overlap
Motif DE_48h DE_48h-TBX18_MA1565.2 9 bp overlap
TBX2 2 datasets
Motif DE_36h DE_36h-TBX2_MA0688.2 9 bp overlap
Motif DE_48h DE_48h-TBX2_MA0688.2 9 bp overlap
TBX21 2 datasets
Motif DE_36h DE_36h-TBX21_MA0690.3 10 bp overlap
Motif DE_48h DE_48h-TBX21_MA0690.3 10 bp overlap
TBX3 2 datasets
Motif DE_36h DE_36h-TBX3_MA1566.3 9 bp overlap
Motif DE_48h DE_48h-TBX3_MA1566.3 9 bp overlap
TBX5 3 datasets
ChIP G296S GSE85628.TBX5.G296S 211 bp overlap
ChIP G296S_2 GSE85628.TBX5.G296S_2 211 bp overlap
ChIP hiPSC GSE81585.TBX5.hiPSC 288 bp overlap
TCF7L1 4 datasets
Motif DE_36h DE_36h-TCF7L1_MA1421.1 12 bp overlap
Motif DE_36h DE_36h-TCF7L1_MA1421.1 12 bp overlap
Motif DE_48h DE_48h-TCF7L1_MA1421.1 12 bp overlap
Motif DE_48h DE_48h-TCF7L1_MA1421.1 12 bp overlap
TCF7L2 2 datasets
Motif DE_36h DE_36h-TCF7L2_MA0523.2 9 bp overlap
Motif DE_48h DE_48h-TCF7L2_MA0523.2 9 bp overlap
TFAP2A 2 datasets
Motif DE_36h DE_36h-TFAP2A_MA0872.1 13 bp overlap
Motif DE_48h DE_48h-TFAP2A_MA0872.1 13 bp overlap
TFAP2B 2 datasets
Motif DE_36h DE_36h-TFAP2B_MA0813.1 13 bp overlap
Motif DE_48h DE_48h-TFAP2B_MA0813.1 13 bp overlap
TFAP2C 2 datasets
Motif DE_36h DE_36h-TFAP2C_MA0815.1 13 bp overlap
Motif DE_48h DE_48h-TFAP2C_MA0815.1 13 bp overlap
TP63 1 dataset
ChIP foreskin GSE126390.TP63.foreskin 242 bp overlap
TRIM28 1 dataset
ChIP WIBR3_NAIVE GSE84382.TRIM28.WIBR3_NAIVE 100 bp overlap
TRPS1 4 datasets
Motif DE_36h DE_36h-TRPS1_MA1970.2 8 bp overlap
Motif DE_36h DE_36h-TRPS1_MA1970.2 8 bp overlap
Motif DE_48h DE_48h-TRPS1_MA1970.2 8 bp overlap
Motif DE_48h DE_48h-TRPS1_MA1970.2 8 bp overlap
Thap11 1 dataset
Motif DE_36h DE_36h-Thap11_MA1573.2 14 bp overlap
ZFP14 2 datasets
Motif DE_36h DE_36h-ZFP14_MA1972.1 15 bp overlap
Motif DE_48h DE_48h-ZFP14_MA1972.1 15 bp overlap
ZIM3 4 datasets
Motif DE_36h DE_36h-ZIM3_MA1709.2 11 bp overlap
Motif DE_48h DE_48h-ZIM3_MA1709.2 11 bp overlap
ChIP HEK293 GSE76494.ZIM3.HEK293 152 bp overlap
ChIP HEK293T GSE78099.ZIM3.HEK293T 100 bp overlap
ZKSCAN5 2 datasets
Motif DE_36h DE_36h-ZKSCAN5_MA1652.2 9 bp overlap
Motif DE_48h DE_48h-ZKSCAN5_MA1652.2 9 bp overlap
ZNF282 1 dataset
Motif DE_36h DE_36h-ZNF282_MA1154.2 15 bp overlap
ZNF317 2 datasets
Motif DE_36h DE_36h-ZNF317_MA1593.2 8 bp overlap
Motif DE_48h DE_48h-ZNF317_MA1593.2 8 bp overlap
ZNF382 2 datasets
Motif DE_36h DE_36h-ZNF382_MA1594.1 24 bp overlap
Motif DE_48h DE_48h-ZNF382_MA1594.1 24 bp overlap
ZNF410 1 dataset
Motif DE_36h DE_36h-ZNF410_MA0752.2 16 bp overlap
ZNF416 2 datasets
Motif DE_36h DE_36h-ZNF416_MA1979.2 10 bp overlap
Motif DE_48h DE_48h-ZNF416_MA1979.2 10 bp overlap
ZNF418 2 datasets
Motif DE_36h DE_36h-ZNF418_MA1980.1 15 bp overlap
Motif DE_48h DE_48h-ZNF418_MA1980.1 15 bp overlap
ZNF90 1 dataset
ChIP HEK293T GSE78099.ZNF90.HEK293T 389 bp overlap
Zic2 1 dataset
Motif DE_36h DE_36h-Zic2_MA1629.2 9 bp overlap
Zic3 1 dataset
Motif DE_36h DE_36h-Zic3_MA0697.3 7 bp overlap