chr3 : 190,325,950 190,326,598
648 bp 76 TFs 2 linked genes
This 648 bp open chromatin element is linked to CLDN16 and CLDN1 and is bound by 76 transcription factors.
Linked Genes
2 genes
Gene Expression Dist. to TSS Distance Link type
CLDN16 3.4 kb Proximal Proximity
CLDN1 3.5 kb Proximal Proximity
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr3:190,320,950 – 190,331,598
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
76 transcription factors
Source
Cell type
ATF2 1 dataset
ChIP GM12878 ENCSR000BQK.ATF2.GM12878 148 bp overlap
BATF 1 dataset
ChIP GM12878 ENCFF954REE 231 bp overlap
BRD3 1 dataset
ChIP HEK293T GSE39579.BRD3.HEK293T 137 bp overlap
BRD4 6 datasets
ChIP CD4_JG1 GSE33281.BRD4.CD4_JG1 272 bp overlap
ChIP HAP1 GSE108387.BRD4.HAP1 648 bp overlap
ChIP RH4_shscr GSE140115.BRD4.RH4_shscr 201 bp overlap
ChIP SUM229PE_pos_30nMtrametinib_24h GSE87418.BRD4.SUM229PE_pos_30nMtrametinib_24h 392 bp overlap
ChIP SUM229PE_pos_DMSO_24h GSE87418.BRD4.SUM229PE_pos_DMSO_24h 551 bp overlap
ChIP keratinocyte GSE140992.BRD4.keratinocyte 648 bp overlap
CHD4 1 dataset
ChIP HaCaT GSE139685.CHD4.HaCaT 169 bp overlap
CREBBP 2 datasets
ChIP keratinocyte GSE140991.CREBBP.keratinocyte 278 bp overlap
ChIP keratinocyte_KLF3i GSE140991.CREBBP.keratinocyte_KLF3i 215 bp overlap
CREM 1 dataset
ChIP GM12878 ENCSR839XZU.CREM.GM12878 175 bp overlap
FEZF1 2 datasets
ChIP HEK293 GSE76494.FEZF1.HEK293 200 bp overlap
ChIP HEK293 ENCSR827NWO.FEZF1.HEK293 56 bp overlap
FLI1 1 dataset
ChIP A-673_1_KRAB_GFP GSE106914.FLI1.A-673_1_KRAB_GFP 201 bp overlap
FOS 2 datasets
ChIP MCF-10A ENCSR000DON.FOS.MCF-10A 236 bp overlap
ChIP MCF-7 ENCSR569XNP.FOS.MCF-7 224 bp overlap
FOSL2 2 datasets
ChIP NPC_R1159Q GSE122631.FOSL2.NPC_R1159Q 251 bp overlap
ChIP hESC GSE69539.FOSL2.hESC 187 bp overlap
FOXA2 1 dataset
ChIP KerCT GSE90454.FOXA2.KerCT 377 bp overlap
FOXM1 3 datasets
ChIP GM12878 ENCFF264DJE 486 bp overlap
ChIP GM12878 ENCFF264DJE 286 bp overlap
ChIP GM12878 ENCSR000BRU.FOXM1.GM12878 138 bp overlap
GATA2 1 dataset
ChIP ESF GSE108408.GATA2.ESF 86 bp overlap
IKZF3 1 dataset
ChIP HEK293 ENCSR304AMN.IKZF3.HEK293 172 bp overlap
JUN 2 datasets
ChIP MCF10A-Er-Src_EtOH GSE115597.JUN.MCF10A-Er-Src_EtOH 200 bp overlap
ChIP keratinocyte_CHD4-KD GSE139685.JUN.keratinocyte_CHD4-KD 151 bp overlap
JUNB 3 datasets
ChIP GM12878 ENCFF667EJQ 227 bp overlap
ChIP GM12878 ENCSR897MMC.JUNB.GM12878 367 bp overlap
ChIP MCF10A-Er-Src_EtOH GSE115597.JUNB.MCF10A-Er-Src_EtOH 164 bp overlap
KDM1A 1 dataset
ChIP NCI-H526_DMSO GSE66297.KDM1A.NCI-H526_DMSO 356 bp overlap
KLF5 2 datasets
ChIP HEK293_E419K GSE88976.KLF5.HEK293_E419K 68 bp overlap
ChIP HEK293_E419Q GSE88976.KLF5.HEK293_E419Q 132 bp overlap
MAML1 1 dataset
ChIP SCC_4h GSE156486.MAML1.SCC_4h 286 bp overlap
MAZ 2 datasets
ChIP HEK293 GSE76494.MAZ.HEK293 193 bp overlap
ChIP HEK293 ENCSR290SSQ.MAZ.HEK293 126 bp overlap
MED1 1 dataset
ChIP hMSC-TERT4_4h GSE104537.MED1.hMSC-TERT4_4h 155 bp overlap
MED12 2 datasets
ChIP leiomyoma_PT886 GSE128230.MED12.leiomyoma_PT886 124 bp overlap
ChIP leiomyoma_PT967 GSE128230.MED12.leiomyoma_PT967 75 bp overlap
MYC 1 dataset
ChIP MCF-10A ENCSR000DOS.MYC.MCF-10A 187 bp overlap
NANOG 3 datasets
ChIP HUES-8 GSE109524.NANOG.HUES-8 89 bp overlap
ChIP WA01_3IL ERP004238.NANOG.WA01_3IL 91 bp overlap
ChIP WA09 GSE105028.NANOG.WA09 271 bp overlap
NFIC 1 dataset
ChIP GM12878 ENCSR000BRN.NFIC.GM12878 217 bp overlap
NFKB1 1 dataset
ChIP MCF10A-Er-Src_TAM GSE115597.NFKB1.MCF10A-Er-Src_TAM 180 bp overlap
NKX2-1 1 dataset
ChIP NCI-H3122 GSE39998.NKX2-1.NCI-H3122 290 bp overlap
NR3C1 3 datasets
ChIP BEAS-2B_TNF_IA1 GSE125623.NR3C1.BEAS-2B_TNF_IA1 491 bp overlap
ChIP HCC1937 GSE152203.NR3C1.HCC1937 150 bp overlap
ChIP SUM159PT GSE152203.NR3C1.SUM159PT 156 bp overlap
PAX7 1 dataset
ChIP H9_DOX GSE98976.PAX7.H9_DOX 213 bp overlap
PGR 1 dataset
ChIP T-47D_R5020-MTVL GSE126859.PGR.T-47D_R5020-MTVL 168 bp overlap
PML 1 dataset
ChIP GM12878 ENCSR000BQM.PML.GM12878 142 bp overlap
POLR2A 3 datasets
ChIP esophagus squamous epithelium ENCFF708IOX 284 bp overlap
ChIP vagina ENCFF305NWS 439 bp overlap
ChIP vagina ENCFF305NWS 477 bp overlap
POU2F2 1 dataset
ChIP GM12878 ENCFF207RKY 263 bp overlap
POU2F3 1 dataset
ChIP NCI-H1048 GSE115123.POU2F3.NCI-H1048 441 bp overlap
POU5F1 2 datasets
ChIP HUES-8 GSE109524.POU5F1.HUES-8 107 bp overlap
ChIP hESC_NAIVE GSE69646.POU5F1.hESC_NAIVE 91 bp overlap
PRDM10 1 dataset
ChIP HEK293 ENCFF145WQQ 327 bp overlap
RBPJ 1 dataset
ChIP SCC_4h GSE156486.RBPJ.SCC_4h 266 bp overlap
RELA 8 datasets
ChIP Detroit-562_LPS GSE91018.RELA.Detroit-562_LPS 367 bp overlap
ChIP Detroit-562_Pam2CSK4 GSE91018.RELA.Detroit-562_Pam2CSK4 296 bp overlap
ChIP Detroit-562_Poly-I GSE91018.RELA.Detroit-562_Poly-I 354 bp overlap
ChIP Detroit-562_TNFa GSE91018.RELA.Detroit-562_TNFa 377 bp overlap
ChIP Detroit-562_tri-DAP GSE91018.RELA.Detroit-562_tri-DAP 355 bp overlap
ChIP FaDu_LPS GSE132018.RELA.FaDu_LPS 648 bp overlap
ChIP HDF_NUTLIN GSE77225.RELA.HDF_NUTLIN 289 bp overlap
ChIP aortic-endothelial-cell_IL1B_D2 GSE139377.RELA.aortic-endothelial-cell_IL1B_D2 201 bp overlap
RUNX3 1 dataset
ChIP GM12878 ENCFF395WHA 326 bp overlap
SIX2 3 datasets
ChIP HEK GSE73865.SIX2.HEK 75 bp overlap
ChIP kidney_fetal_16w GSE75948.SIX2.kidney_fetal_16w 57 bp overlap
ChIP kidney_fetal_17w GSE75948.SIX2.kidney_fetal_17w 73 bp overlap
SMAD3 2 datasets
ChIP HCC1954 GSE104760.SMAD3.HCC1954 519 bp overlap
ChIP HCC1954_TGFb GSE104760.SMAD3.HCC1954_TGFb 238 bp overlap
SMARCA4 5 datasets
ChIP A-549_AG15686 GSE132290.SMARCA4.A-549_AG15686 237 bp overlap
ChIP A-549_AG15688 GSE132290.SMARCA4.A-549_AG15688 283 bp overlap
ChIP A-549_AG15725 GSE132290.SMARCA4.A-549_AG15725 57 bp overlap
ChIP endometrial-epithelial-cells_KO GSE106660.SMARCA4.endometrial-epithelial-cells_KO 534 bp overlap
ChIP hiPSC GSE124903.SMARCA4.hiPSC 159 bp overlap
SOX2 1 dataset
ChIP KYSE-70 GSE46837.SOX2.KYSE-70 287 bp overlap
SP4 1 dataset
ChIP HEK293 GSE76494.SP4.HEK293 204 bp overlap
TBP 1 dataset
ChIP GM12878 ENCFF571OXR 385 bp overlap
TBX21 1 dataset
ChIP GM12878 ENCSR739IHN.TBX21.GM12878 260 bp overlap
TEAD1 1 dataset
ChIP keratinocyte GSE138727.TEAD1.keratinocyte 293 bp overlap
TFAP2C 2 datasets
ChIP HCC1954 GSE126897.TFAP2C.HCC1954 186 bp overlap
ChIP UCLA1-hESCs_TFAP2C--_2d_TFAP2C_Induc GSE101074.TFAP2C.UCLA1-hESCs_TFAP2C--_2d_TFAP2C_Induc 266 bp overlap
TP53 44 datasets
ChIP A-498_2h_4GY GSE100292.TP53.A-498_2h_4GY 203 bp overlap
ChIP GM00011 GSE55727.TP53.GM00011 648 bp overlap
ChIP GM06170 GSE55727.TP53.GM06170 648 bp overlap
ChIP H9 GSE142050.TP53.H9 492 bp overlap
ChIP H9_ectoderm GSE142050.TP53.H9_ectoderm 383 bp overlap
ChIP H9_mesoderm GSE142050.TP53.H9_mesoderm 483 bp overlap
ChIP HCT-116_5FU GSE58506.TP53.HCT-116_5FU 278 bp overlap
ChIP HCT-116_Nutlin3a-SC GSE125927.TP53.HCT-116_Nutlin3a-SC 200 bp overlap
ChIP HCT-116_nutlin GSE86164.TP53.HCT-116_nutlin 563 bp overlap
ChIP IMR-90 GSE115940.TP53.IMR-90 648 bp overlap
ChIP IMR-90_APO GSE53491.TP53.IMR-90_APO 188 bp overlap
ChIP IMR-90_DMSO GSE58740.TP53.IMR-90_DMSO 155 bp overlap
ChIP IMR-90_NUT3A GSE58740.TP53.IMR-90_NUT3A 648 bp overlap
ChIP IMR-90_SENES_SHLUC GSE42728.TP53.IMR-90_SENES_SHLUC 146 bp overlap
ChIP MCF-10A_Nutlin3A GSE111009.TP53.MCF-10A_Nutlin3A 590 bp overlap
ChIP MCF-7_1h_IR_10Gy GSE100099.TP53.MCF-7_1h_IR_10Gy 321 bp overlap
ChIP MCF-7_2-5h_IR_10Gy GSE100099.TP53.MCF-7_2-5h_IR_10Gy 347 bp overlap
ChIP MCF-7_7-5h_IR_10Gy GSE100099.TP53.MCF-7_7-5h_IR_10Gy 487 bp overlap
ChIP MCF-7_7-5h_IR_10Gy_Nutlin GSE100099.TP53.MCF-7_7-5h_IR_10Gy_Nutlin 540 bp overlap
ChIP MCF-7_NCS-treated GSE101737.TP53.MCF-7_NCS-treated 237 bp overlap
ChIP MCF-7_nutlin GSE86164.TP53.MCF-7_nutlin 648 bp overlap
ChIP MOLM-13_R282W_DMSO GSE131484.TP53.MOLM-13_R282W_DMSO 432 bp overlap
ChIP MOLM-13_R282W_Daunorubicin GSE131484.TP53.MOLM-13_R282W_Daunorubicin 377 bp overlap
ChIP SJSA-1 GSE86164.TP53.SJSA-1 482 bp overlap
ChIP SJSA-1_nutlin GSE86164.TP53.SJSA-1_nutlin 648 bp overlap
ChIP SaOS-2 GSE15780.TP53.SaOS-2 592 bp overlap
ChIP U2OS_DMSO GSE46641.TP53.U2OS_DMSO 172 bp overlap
ChIP U2OS_DXR GSE46641.TP53.U2OS_DXR 238 bp overlap
ChIP U2OS_NUT GSE46641.TP53.U2OS_NUT 475 bp overlap
ChIP U2OS_UV_16H ERP004176.TP53.U2OS_UV_16H 308 bp overlap
ChIP U2OS_UV_8H ERP004176.TP53.U2OS_UV_8H 212 bp overlap
ChIP UACC-257_nutlin_2h GSE100292.TP53.UACC-257_nutlin_2h 289 bp overlap
ChIP UO-31_2h_4GY GSE100292.TP53.UO-31_2h_4GY 595 bp overlap
ChIP keratinocyte GSE56674.TP53.keratinocyte 254 bp overlap
ChIP keratinocyte_ADRIA GSE56674.TP53.keratinocyte_ADRIA 352 bp overlap
ChIP keratinocyte_CISP GSE56674.TP53.keratinocyte_CISP 428 bp overlap
ChIP lymphocyte_104_Nutlin GSE110368.TP53.lymphocyte_104_Nutlin 390 bp overlap
ChIP lymphocyte_116 GSE110368.TP53.lymphocyte_116 260 bp overlap
ChIP lymphocyte_116_DXR GSE110368.TP53.lymphocyte_116_DXR 480 bp overlap
ChIP lymphocyte_116_Nutlin GSE110368.TP53.lymphocyte_116_Nutlin 627 bp overlap
ChIP lymphocyte_45_DXR GSE110368.TP53.lymphocyte_45_DXR 420 bp overlap
ChIP lymphocyte_45_Nutlin GSE110368.TP53.lymphocyte_45_Nutlin 246 bp overlap
ChIP lymphocyte_90_DXR GSE110368.TP53.lymphocyte_90_DXR 588 bp overlap
ChIP lymphocyte_90_Nutlin GSE110368.TP53.lymphocyte_90_Nutlin 506 bp overlap
TP53_PS15 2 datasets
ChIP U2OS_ACTD GSE21939.TP53_PS15.U2OS_ACTD 295 bp overlap
ChIP U2OS_ETO GSE21939.TP53_PS15.U2OS_ETO 329 bp overlap
TP63 22 datasets
ChIP BxPC-3 GSE115461.TP63.BxPC-3 576 bp overlap
ChIP EP156T GSE43111.TP63.EP156T 239 bp overlap
ChIP HCC95 GSE46837.TP63.HCC95 244 bp overlap
ChIP HaCaT_LacZ GSE60814.TP63.HaCaT_LacZ 195 bp overlap
ChIP HaCaT_LacZ_TGFB GSE60814.TP63.HaCaT_LacZ_TGFB 486 bp overlap
ChIP HaCaT_caRAS_TGFB GSE60814.TP63.HaCaT_caRAS_TGFB 484 bp overlap
ChIP HaCaT_caRAS_TGFB GSE60814.TP63.HaCaT_caRAS_TGFB 122 bp overlap
ChIP HaCaT_dnRAS_TGFB GSE60814.TP63.HaCaT_dnRAS_TGFB 492 bp overlap
ChIP JHU-029 GSE88859.TP63.JHU-029 365 bp overlap
ChIP KYSE-70 GSE46837.TP63.KYSE-70 368 bp overlap
ChIP MCF-10A_DCIS GSE72009.TP63.MCF-10A_DCIS 383 bp overlap
ChIP MCF-10A_Nutlin3A GSE111009.TP63.MCF-10A_Nutlin3A 648 bp overlap
ChIP SUIT-2 GSE115461.TP63.SUIT-2 648 bp overlap
ChIP keratinocyte GSE56674.TP63.keratinocyte 389 bp overlap
ChIP keratinocyte_ADRIA GSE56674.TP63.keratinocyte_ADRIA 263 bp overlap
ChIP keratinocyte_CISP GSE56674.TP63.keratinocyte_CISP 369 bp overlap
ChIP keratinocyte_D0 GSE59824.TP63.keratinocyte_D0 441 bp overlap
ChIP keratinocyte_D2 GSE59824.TP63.keratinocyte_D2 415 bp overlap
ChIP keratinocyte_D4 GSE59824.TP63.keratinocyte_D4 358 bp overlap
ChIP keratinocyte_D7 GSE59824.TP63.keratinocyte_D7 370 bp overlap
ChIP keratinocyte_epidermal GSE67382.TP63.keratinocyte_epidermal 523 bp overlap
ChIP keratinocyte_epidermal_KDPAF GSE67382.TP63.keratinocyte_epidermal_KDPAF 304 bp overlap
TP73 1 dataset
ChIP GM12878 GSE97661.TP73.GM12878 293 bp overlap
TP73_TA 1 dataset
ChIP SaOS-2 GSE15780.TP73_TA.SaOS-2 315 bp overlap
TRIM28 1 dataset
ChIP HEK293 ENCFF582MWI 568 bp overlap
WT1 2 datasets
ChIP HEK293 ENCFF906HIR 65 bp overlap
ChIP HEK293 ENCSR966PJJ.WT1.HEK293 161 bp overlap
YY1 3 datasets
ChIP HEK293 ENCSR859RAO.YY1.HEK293 276 bp overlap
ChIP Hep-G2_NC GSE120104.YY1.Hep-G2_NC 142 bp overlap
ChIP Hep-G2_RBM25-KD GSE120104.YY1.Hep-G2_RBM25-KD 498 bp overlap
ZBED1 2 datasets
ChIP GM12878 ENCFF007OSW 440 bp overlap
ChIP GM12878 ENCFF007OSW 505 bp overlap
ZBTB17 1 dataset
ChIP HEK293 ENCFF865LIO 441 bp overlap
ZFP37 3 datasets
ChIP HEK293 ENCFF968PWB 491 bp overlap
ChIP HEK293 ENCFF968PWB 366 bp overlap
ChIP HEK293 ENCSR365GRX.ZFP37.HEK293 314 bp overlap
ZNF335 2 datasets
ChIP HEK293 ENCFF784SLD 577 bp overlap
ChIP HEK293 ENCSR328SUD.ZNF335.HEK293 448 bp overlap
ZNF341 1 dataset
ChIP HEK293 GSE76494.ZNF341.HEK293 81 bp overlap
ZNF823 1 dataset
ChIP HEK293T GSE78099.ZNF823.HEK293T 290 bp overlap